Labeobarbus Explained
Labeobarbus is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular for the southern species, is yellowfish. The scientific name refers to the fact that these large barbs recall the fairly closely related "carps" in the genus Labeo in size and shape. As far as can be told, all Labeobarbus species are hexaploid.[1] One species, L. microbarbis from Rwanda, is known to have gone extinct in recent times.
Systematics
Like many other "barbs", it was long included in Barbus. It appears to be a fairly close relative of the typical barbels and relatives - the genus Barbus proper - but closer still to the large Near Eastern species nowadays separated in Carasobarbus. Barbus has been split to account for the improved phylogenetic knowledge which indicated it was highly paraphyletic in its wide circumscription; it may be that Carasobarbus and some other closely related "barbs" (e.g. Labeobarbus reinii) are now included in Labeobarbus to avoid a profusion of very small genera.[2]
The taxonomy of many species in the "wastebin genus" Barbus has recently been re-evaluated. Though hybrid introgression may confound studies based in mtDNA data alone, a number of these species appear to be so closely related to Labeobarbus as to warrant inclusion in the present genus outright, irrespective of whether Carasobarbus is considered distinct or not. These include L. ethiopicus.
Species
Labeobarbus at present contains the following species:
- Labeobarbus acuticeps (Matthes, 1959)
- Labeobarbus acutirostris (Bini, 1940)
- Labeobarbus aeneus (Burchell, 1822) (Smallmouth yellowfish)
- Labeobarbus alluaudi Pellegrin, 1909
- Labeobarbus altianalis Boulenger, 1900 (Ripon barbel)
- Labeobarbus altipinnis Banister & Poll, 1973
- Labeobarbus ansorgii Boulenger, 1906
- Labeobarbus aspius (Boulenger, 1912)
- Labeobarbus axelrodi (Getahun, Stiassny & Teugels, 2004)
- Labeobarbus batesii (Boulenger, 1903)
- Labeobarbus beso (Rüppell, 1835)
- Labeobarbus boulengeri (Vreven, Musschoot, Snoeks & Schliewen, 2016)
- Labeobarbus brauni (Pellegrin, 1935)
- Labeobarbus brevicephalus (Nagelkerke & Sibbing, 1997)
- Labeobarbus brevispinis (Holly, 1927)
- Labeobarbus bynni (Forsskål, 1775)
- Labeobarbus cardozoi (Boulenger, 1912)
- Labeobarbus caudovittatus (Boulenger, 1902)
- Labeobarbus clarkeae (Banister, 1984)
- Labeobarbus claudinae De Vos & Thys van den Audenaerde, 1990
- Labeobarbus codringtonii (Boulenger, 1908) (Upper Zambezi yellowfish)
- Labeobarbus compiniei (Sauvage, 1879)
- Labeobarbus crassibarbis (Nagelkerke & Sibbing, 1997)
- Labeobarbus dainellii (Bini, 1940)
- Labeobarbus dartevellei Poll, 1945
- Labeobarbus dimidiatus Tweddle & P. H. Skelton, 1998
- Labeobarbus ensifer Boulenger, 1910
- Labeobarbus ethiopicus Zolezzi, 1939
- Labeobarbus fasolt Pappenheim, 1914
- Labeobarbus fimbriatus Holly, 1926
- Labeobarbus gananensis Vinciguerra, 1895
- Labeobarbus gestetneri Banister & R. G. Bailey, 1979
- Labeobarbus girardi Boulenger, 1910
- Labeobarbus gorgorensis (Bini, 1940)
- Labeobarbus gorguari (Rüppell, 1835)
- Labeobarbus gruveli Pellegrin, 1911
- Labeobarbus gulielmi Boulenger, 1910
- Labeobarbus habereri (Steindachner, 1912)
- Labeobarbus huloti Banister, 1976
- Labeobarbus humphri Banister, 1976
- Labeobarbus intermedius (Rüppell, 1835)
- Labeobarbus iphthimostoma Banister & Poll, 1973
- Labeobarbus iturii Holly, 1929
- Labeobarbus jaegeri Holly, 1930
- Labeobarbus johnstonii (Boulenger, 1907)
- Labeobarbus jubae Banister, 1984
- Labeobarbus jubbi Poll, 1967
- Labeobarbus kimberleyensis (Gilchrist & W. W. Thompson, 1913) (Largemouth yellowfish)
- Labeobarbus lagensis (Günther, 1868)
- Labeobarbus latirostris (Keilhack, 1908)
- Labeobarbus leleupanus Matthes, 1959\
- Labeobarbus lobogenysoides Pellegrin, 1935
- Labeobarbus longidorsalis Pellegrin, 1935
- Labeobarbus longifilis Pellegrin, 1935
- Labeobarbus longissimus (Nagelkerke & Sibbing, 1997)
- Labeobarbus lucius (Boulenger, 1910)
- Labeobarbus lufupensis Banister & R. G. Bailey, 1979
- Labeobarbus macroceps Fowler, 1936
- Labeobarbus macrolepidotus Pellegrin, 1928
- Labeobarbus macrolepis Pfeffer, 1889
- Labeobarbus macrophtalmus (Bini, 1940)
- Labeobarbus malacanthus (Pappenheim, 1911)
- Labeobarbus marequensis (A. Smith, 1841) (Largescale yellowfish)
- Labeobarbus mariae Holly, 1926
- Labeobarbus matris Holly, 1928
- Labeobarbus mawambiensis Steindachner, 1911
- Labeobarbus mbami (Holly, 1927)
- Labeobarbus megastoma (Nagelkerke & Sibbing, 1997)
- †Labeobarbus microbarbis L. R. David & Poll, 1937 (declared extinct in 2006)
- Labeobarbus micronema (Boulenger, 1904)
- Labeobarbus microterolepis Boulenger, 1902
- Labeobarbus mirabilis Pappenheim, 1914
- Labeobarbus mungoensis (Trewavas, 1974)
- Labeobarbus nanningsi (de Beaufort, 1933)
- Labeobarbus natalensis (Castelnau, 1861) (Scaly yellowfish)
- Labeobarbus nedgia Rüppell, 1835
- Labeobarbus nelspruitensis Gilchrist & W. W. Thompson, 1911 (Incomati chiselmouth)
- Labeobarbus nthuwa Tweddle & P. H. Skelton, 2008
- Labeobarbus osseensis (Nagelkerke & Sibbing, 2000)
- Labeobarbus oxyrhynchus Pfeffer, 1889 (Pangani barb)
- Labeobarbus pagenstecheri J. G. Fischer, 1884
- Labeobarbus parawaldroni Lévêque, Thys van den Audenaerde & Traoré, 1987
- Labeobarbus pellegrini Bertin & Estève, 1948
- Labeobarbus petitjeani Daget, 1962
- Labeobarbus platydorsus (Nagelkerke & Sibbing, 1997)
- Labeobarbus platyrhinus Boulenger, 1900
- Labeobarbus platystomus Pappenheim, 1914 (Rwandese carp)
- Labeobarbus pojeri (Poll, 1944)
- Labeobarbus polylepis (Boulenger, 1907) (Smallscale yellowfish)
- Labeobarbus progenys (Boulenger, 1903)
- Labeobarbus pungweensis R. A. Jubb, 1959 (Pungwe chiselmouth)
- †Labeobarbus reinii Günther, 1874 (Giant Atlas barbel) (declared extinct in 2022)
- Labeobarbus rhinoceros (Copley, 1938) (Rhinofish)
- Labeobarbus rhinophorus Boulenger, 1910
- Labeobarbus robertsi Banister, 1984
- Labeobarbus rocadasi (Boulenger, 1910)
- Labeobarbus rosae (Boulenger, 1910)
- Labeobarbus roylii (Boulenger, 1912)
- Labeobarbus ruandae Pappenheim, 1914
- Labeobarbus ruasae Pappenheim, 1914
- Labeobarbus ruwenzorii (Pellegrin, 1909)
- Labeobarbus sacratus Daget, 1963
- Labeobarbus sandersi Boulenger, 1912
- Labeobarbus seeberi (Gilchrist & Thompson, 1913) (Clanwilliam yellowfish)
- Labeobarbus semireticulatus Pellegrin, 1924
- Labeobarbus somereni Boulenger, 1911
- Labeobarbus stappersii Boulenger, 1915
- Labeobarbus steindachneri Boulenger, 1910
- Labeobarbus stenostoma Boulenger, 1910
- Labeobarbus surkis (Rüppell, 1835)
- Labeobarbus tornieri Steindachner, 1906
- Labeobarbus trachypterus Boulenger, 1915
- Labeobarbus tropidolepis Boulenger, 1900
- Labeobarbus truttiformis (Nagelkerke & Sibbing, 1997)
- Labeobarbus tsanensis (Nagelkerke & Sibbing, 1997)
- Labeobarbus upembensis Banister & R. G. Bailey, 1979
- Labeobarbus urotaenia Boulenger, 1913
- Labeobarbus varicostoma Boulenger, 1910
- Labeobarbus versluysii (Holly, 1929)
- Labeobarbus werneri Holly, 1929
- Labeobarbus wittei Banister & Poll, 1973
- Labeobarbus wurtzi Pellegrin, 1908
- Labeobarbus xyrocheilus Tweddle & P. H. Skelton, 1998
References
- (2007): Evolutionary origin of Lake Tana's (Ethiopia) small Barbus species: indications of rapid ecological divergence and speciation. Anim. Biol. 57(1): 39–48. (HTML abstract)
- (2009): 2009 IUCN Red List of Threatened Species. Version 2009.1. Retrieved 2009-SEP-20.
Notes and References
- de Graaf et al. (2007), IUCN (2009)
- de Graaf et al. (2007)