Thynnidae Explained

The Thynnidae (also known as thynnid wasps, flower wasps, or thynnid flower wasps) are a family of large, solitary wasps whose larvae are almost universally parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea. Until recently, the constituents of this family were classified in the family Tiphiidae, but multiple studies have independently confirmed that thynnids are a separate lineage.

Description

Most species are small, but they can be up to 30 mm long. The females of some subfamilies (Diamminae, Methochinae, and most Thynninae) are wingless and hunt ground-dwelling (fossorial) beetle larvae, or (in one species) mole crickets. The prey is paralysed with the female's sting, and an egg is laid on it so the wasp larva has a ready supply of food. In species where both sexes are winged, males are similar in size to the females, but are much more slender. The males of species with wingless females, however, are often much larger than the females and have wings; the adults mate in the air, with the female carried by the male's genitalia. Adults feed on nectar and are minor pollinators. As some of the ground-dwelling scarab species attacked by thynnids are pests, some of these wasps are considered beneficial as biological control agents.

Taxonomy

The family has five extant subfamilies, which were previously placed in Tiphiidae before it was found to be non-monophyletic. Thynnidae genera are classified as follows:

Extant genera

Subfamily: Anthoboscinae Turner, 1912

Subfamily: Diamminae Turner, 1907

Subfamily: Methochinae Rohwer, 1916

Subfamily: Myzininae Borner, 1919

Tribe: Mesini Argaman, 1994
Tribe: Meriini Costa, 1858

Subfamily: Thynninae Ashmead, 1903

Tribe: Elaphropterini Kimsey, 1992
Tribe: Rhagigasterini Ashmead, 1903
Tribe: Scotaenini Kimsey,1992
Tribe: Thynnini

Extinct genera

The following 4 genera are extinct members of Thynnidae:

Subfamily: Anthoboscinae Turner, 1912

Subfamily: Methochinae Rohwer, 1916

Subfamily: Myzininae Borner, 1919

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