Septum (cell biology) explained

A septum in cell biology is the new cell wall that forms between two daughter cells as a result of cell division.[1]

In yeast, septins form a ring structure, to which other proteins are recruited.[2] In particular, chitin synthase 2 is required, an enzyme that synthesises chitin thereby building up the primary septum. A secondary septum of β-glucans and mannoproteins is then assembled using the enzyme 1,3-Beta-glucan synthase, and the primary septum degraded during cell separation. After degradation of the primary septum, a chitinous bud scar remains on both the mother and daughter cell. [2] [3]

Composition

In Schizosaccharomyces pombe, the primary septum is composed of linear β(1,3)-D-glucan, β(1,6) branches, and α(1,3)-D-glucan.[4] The secondary septum in Schizosaccharomyces pombe is composed of β(1,6)-D-glucan, β(1,6) branches, and α(1,3)-D-glucan. The synthesis of linear β(1,3)-D-glucan for the primary septum is done by the enzyme β(1,3)-D-glucan synthase and regulated by a Rho GTPase. Ags1/Mok1 enzyme is responsible for the synthesis of α(1,3)-D-glucan in the primary septum and secondary septum.

Notes and References

  1. O'Connor C . 2008 . Cell Division: Stages of Mitosis. . Nature Education . 1 . 1 . 188 .
  2. Cabib E, Roh DH, Schmidt M, Crotti LB, Varma A . The yeast cell wall and septum as paradigms of cell growth and morphogenesis . The Journal of Biological Chemistry . 276 . 23 . 19679–82 . June 2001 . 11309404 . 10.1074/jbc.R000031200 . free .
  3. Lesage G, Bussey H . Cell wall assembly in Saccharomyces cerevisiae . Microbiology and Molecular Biology Reviews . 70 . 2 . 317–43 . June 2006 . 16760306 . 1489534 . 10.1128/MMBR.00038-05 .
  4. García Cortés JC, Ramos M, Osumi M, Pérez P, Ribas JC . The Cell Biology of Fission Yeast Septation . Microbiology and Molecular Biology Reviews . 80 . 3 . 779–91 . September 2016 . 27466282 . 4981666 . 10.1128/MMBR.00013-16 .