Mackerel Explained
Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. Mackerel species typically have deeply forked tails and vertical "tiger-like" stripes on their backs with an iridescent green-blue quality.[1] [2] Many are restricted in their distribution ranges and live in separate populations or fish stocks based on geography. Some stocks migrate in large schools along the coast to suitable spawning grounds, where they spawn in fairly shallow waters. After spawning they return the way they came in smaller schools to suitable feeding grounds, often near an area of upwelling. From there they may move offshore into deeper waters and spend the winter in relative inactivity. Other stocks migrate across oceans.
Smaller mackerel are forage fish for larger predators, including larger mackerel and Atlantic cod.[3] Flocks of seabirds, whales, dolphins, sharks, and schools of larger fish such as tuna and marlin follow mackerel schools and attack them in sophisticated and cooperative ways. Mackerel flesh is high in omega-3 oils and is intensively harvested by humans. In 2009, over 5 million tons were landed by commercial fishermen. Sport fishermen value the fighting abilities of the king mackerel.[4]
Species
Over 30 different species, principally belonging to the family Scombridae, are commonly referred to as mackerel. The term "mackerel" is derived from Old French and may have originally meant either "marked, spotted" or "pimp, procurer". The latter connection is not altogether clear, but mackerel spawn enthusiastically in shoals near the coast, and medieval ideas on animal procreation were creative.[5]
Scombroid mackerels
About 21 species in the family Scombridae are commonly called mackerel. The type species for the scombroid mackerel is the Atlantic mackerel, Scomber scombrus. Until recently, Atlantic chub mackerel and Indo-Pacific chub mackerel were thought to be subspecies of the same species. In 1999, Collette established, on molecular and morphological considerations, that these are separate species. Mackerel are smaller with shorter lifecycles than their close relatives, the tuna, which are also members of the same family.[6] [7]
Scombrini, the true mackerels
The true mackerels belong to the tribe Scombrini. The tribe consists of seven species, each belonging to one of two genera: Scomber or Rastrelliger.
Scomberomorini, the Spanish mackerels
The Spanish mackerels belong to the tribe Scomberomorini, which is the "cousin tribe" of the true mackerels. This tribe consists of 21 species in all—18 of those are classified into the genus Scomberomorus, two into Grammatorcynus, and a single species into the monotypic genus Acanthocybium.
Spanish Mackerels (tribe Scomberomorini) |
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Common name | Scientific name | Maximum length | Common length | Maximum weight | Maximum age | Trophic level | FishBase | FAO | IUCN status |
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Wahoo | Acanthocybium solandri (Cuvier in Cuvier and Valenciennes, 1832) | 250 cm | 170 cm | 83 kg | years | 4.4 | | | Least concern[19] |
| Grammatorcynus bicarinatus (Quoy & Gaimard, 1825) | 112 cm | cm | 13.5 kg | years | 4.5 | | | Least concern[20] |
Double-lined mackerel | G. bilineatus (Rüppell, 1836) | 100 cm | 50 cm | 3.5 kg | years | 4.18 | | | Least concern[21] |
Serra Spanish mackerel | Scomberomorus brasiliensis (Collette, Russo & Zavala-Camin, 1978) | cm | cm | kg | years | 3.31 | | | Least concern[22] |
King mackerel | S. cavalla (Cuvier, 1829) | 184 cm | 70 cm | 45 kg | 14 years | 4.5 | | [23] | Least concern[24] |
Narrow-barred Spanish mackerel | S. commerson (Lacepède, 1800) | 240 cm | 120 cm | 70 kg | years | 4.5 | | [25] | Near threatened[26] |
Monterey Spanish mackerel | S. concolor (Lockington, 1879) | cm | cm | kg | years | 4.24 | | | Vulnerable[27] |
Indo-Pacific king mackerel | S. guttatus (Bloch & Schneider, 1801) | 76 cm | 55 cm | kg | years | 4.28 | | [28] | Data deficient[29] |
Korean mackerel | S. koreanus (Kishinouye, 1915) | 150 cm | 60 cm | 15 kg | years | 4.2 | | | Least concern[30] |
Streaked Spanish mackerel | S. lineolatus (Cuvier, 1829) | 80 cm | 70 cm | kg | years | 4.5 | | | Least concern[31] |
Atlantic Spanish mackerel | S. maculatus (Mitchill, 1815) | 91 cm | cm | 5.89 kg | 5 years | 4.5 | | [32] | Least concern[33] |
Papuan Spanish mackerel | S. multiradiatus Munro, 1964 | 35 cm | cm | 0.5 kg | years | 4.0 | | | Least concern[34] |
Australian spotted mackerel | S. munroi (Collette & Russo, 1980) | 104 cm | cm | 10.2 kg | years | 4.3 | | | Near threatened[35] |
Japanese Spanish mackerel | S. niphonius (Cuvier, 1832) | 100 cm | cm | 7.1 kg | years | 4.5 | | [36] | Data deficient[37] |
Queen mackerel | S. plurilineatus Fourmanoir, 1966 | 120 cm | cm | 12.5 kg | years | 4.2 | | | Data deficient[38] |
Queensland school mackerel | S. queenslandicus (Munro, 1943) | 100 cm | 80 cm | 12.2 kg | years | 4.5 | | | Least concern[39] |
Cero mackerel | S. regalis (Bloch, 1793) | 183 cm | cm | 7.8 kg | years | 4.5 | | | Least concern[40] |
Broadbarred king mackerel | S. semifasciatus (Macleay, 1883) | 120 cm | cm | kg | 10 years | 4.5 | | | Least concern[41] |
Pacific sierra | S. sierra (Cuvier, 1832) | 99 cm | 60 cm | 8.2 kg | years | 4.5 | | | Least concern[42] |
Chinese mackerel | S. sinensis (Cuvier, 1832) | 247 cm | 100 cm | kg | years | 4.5 | | | Data deficient |
West African Spanish mackerel | S. tritor (Cuvier, 1832) | cm | cm | kg | years | 4.26 | | | Least concern | |
Other mackerel
In addition, a number of species with mackerel-like characteristics in the families Carangidae, Hexagrammidae and Gempylidae are commonly referred to as mackerel. Some confusion had occurred between the Pacific jack mackerel (Trachurus symmetricus) and the heavily harvested Chilean jack mackerel (T. murphyi). These have been thought at times to be the same species, but are now recognised as separate species.[43]
Other mackerel species |
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Family | Common name | Scientific name | Maximum length | Common length | Maximum weight | Maximum age | Trophic level | FishBase | FAO | IUCN status |
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Scombridae Gasterochisma | Butterfly mackerel | Gasterochisma melampus Richardson, 1845 | 175 cm | 153 cm | kg | years | 4.4 | | | Least concern[44] |
Carangidae Jack mackerel | Atlantic horse mackerel | Trachurus trachurus (Linnaeus, 1758) | 70 cm | 22 cm | 2.0 kg | years | 3.64 | | [45] | Not assessed |
Blue jack mackerel | T. picturatus (Bowdich, 1825) | 60 cm | 25 cm | kg | years | 3.32 | | | Data deficient[46] |
Cape horse mackerel | T. capensis (Castelnau, 1861) | 60 cm | 30 cm | kg | years | 3.47 | | [47] | Not assessed[48] |
Chilean jack mackerel | T. murphyi (Nichols, 1920) | 70 cm | 45 cm | kg | 16 years | 3.49 | | [49] | Data deficient |
| T. trecae (Cadenat, 1950) | 35 cm | cm | 2.0 kg | years | 3.49 | | [50] | Not assessed |
Greenback horse mackerel | T. declivis (Jenyns, 1841) | 64 cm | 42 cm | kg | 25 years | 3.93 | | [51] | Not assessed[52] |
Japanese horse mackerel | T. japonicus (Temminck & Schlegel, 1844) | 50 cm | 35 cm | 0.66 kg | 12 years | 3.4 | | [53] | Not assessed |
Mediterranean horse mackerel | T. mediterraneus (Steindachner, 1868) | 60 cm | 30 cm | kg | years | 3.59 | | [54] | Not assessed |
Pacific jack mackerel | T. symmetricus (Ayres, 1855) | 81 cm | 55 cm | kg | 30 years | 3.56 | | | Least concern[55] |
Yellowtail horse mackerel | T. novaezelandiae (Richardson, 1843) | 50 cm | 35 cm | kg | 25 years | 4.5 | | | Not assessed |
Gempylidae Snake mackerel | Black snake mackerel | Nealotus tripes (Johnson, 1865) | 25 cm | 15 cm | kg | years | 4.2 | | | Not assessed |
Blacksail snake mackerel | Thyrsitoides marleyi (Fowler, 1929) | 200 cm | 100 cm | kg | years | 4.19 | | | Not assessed |
| Gempylus serpens (Cuvier, 1829) | 100 cm | 60 cm | kg | years | 4.35 | | | Not assessed |
Violet snake mackerel | Nesiarchus nasutus (Johnson, 1862) | 130 cm | 80 cm | kg | years | 4.33 | | | Not assessed |
| Thyrsitops lepidopoides (Cuvier, 1832) | 40 cm | 25 cm | kg | years | 3.86 | | | Not assessed |
Hexagrammidae | Okhotsk atka mackerel | Pleurogrammus azonus (Jordan & Metz, 1913) | 62 cm | cm | 1.6 kg | 12 years | 3.58 | | [56] | Not assessed |
Atka mackerel | P. monopterygius (Pallas, 1810) | 56.5 cm | cm | 2.0 kg | 14 years | 3.33 | | | Not assessed | |
The term "mackerel" is also used as a modifier in the common names of other fish, sometimes indicating the fish has vertical stripes similar to a scombroid mackerel:
By extension, the term is applied also to other species such as the mackerel tabby cat,[57] and to inanimate objects such as the altocumulus mackerel sky cloud formation.[58] [59]
Characteristics
Most mackerel belong to the family Scombridae, which also includes tuna and bonito. Generally, mackerel are much smaller and slimmer than tuna, though in other respects, they share many common characteristics. Their scales, if present at all, are extremely small. Like tuna and bonito, mackerel are voracious feeders, and are swift and manoeuvrable swimmers, able to streamline themselves by retracting their fins into grooves on their bodies. Like other scombroids, their bodies are cylindrical with numerous finlets on the dorsal and ventral sides behind the dorsal and anal fins, but unlike the deep-bodied tuna, they are slim.
The type species for scombroid mackerels is the Atlantic mackerel, Scomber scombrus. These fish are iridescent blue-green above with a silvery underbelly and near-vertical wavy black stripes running along their upper bodies.[60]
The prominent stripes on the back of mackerels seemingly are there to provide camouflage against broken backgrounds. That is not the case, though, because mackerel live in midwater pelagic environments which have no background.[61] However, fish have an optokinetic reflex in their visual systems that can be sensitive to moving stripes.[62] For fish to school efficiently, they need feedback mechanisms that help them align themselves with adjacent fish, and match their speed. The stripes on neighbouring fish provide "schooling marks", which signal changes in relative position.[63]
A layer of thin, reflecting platelets is seen on some of the mackerel stripes. In 1998, E J Denton and D M Rowe argued that these platelets transmit additional information to other fish about how a given fish moves. As the orientation of the fish changes relative to another fish, the amount of light reflected to the second fish by this layer also changes. This sensitivity to orientation gives the mackerel "considerable advantages in being able to react quickly while schooling and feeding."[64]
Mackerel range in size from small forage fish to larger game fish. Coastal mackerel tend to be small.[65] The king mackerel is an example of a larger mackerel. Most fish are cold-blooded, but exceptions exist. Certain species of fish maintain elevated body temperatures. Endothermic bony fishes are all in the suborder Scombroidei and include the butterfly mackerel, a species of primitive mackerel.[66]
Mackerel are strong swimmers. Known in the latin family as "punctualis piscis" which translates to "punctual fish." This is due to its punctuality of migration during mating season as it moves from warm to cold waters. Atlantic mackerel can swim at a sustained speed of 0.98 m/sec with a burst speed of 5.5 m/sec,[67] [68] while chub mackerel can swim at a sustained speed of 0.92 m/sec with a burst speed of 2.25 m/sec.[69]
Distribution
Most mackerel species have restricted distribution ranges.
Some mackerel species migrate vertically. Adult snake mackerel conduct a diel vertical migration, staying in deeper water during the day and rising to the surface at night to feed. The young and juveniles also migrate vertically, but in the opposite direction, staying near the surface during the day and moving deeper at night.[70]
Lifecycle
Mackerel are prolific broadcast spawners, and must breed near the surface of the water because the eggs of the females float. Individual females lay between 300,000 and 1,500,000 eggs. Their eggs and larvae are pelagic, that is, they float free in the open sea. The larvae and juvenile mackerel feed on zooplankton. As adults, they have sharp teeth, and hunt small crustaceans such as copepods, forage fish, shrimp, and squid. In turn, they are hunted by larger pelagic animals such as tuna, billfish, sea lions, sharks, and pelicans.[71]
Off Madagascar, spinner sharks follow migrating schools of mackerel.[72] Bryde's whales feed on mackerel when they can find them. They use several feeding methods, including skimming the surface, lunging, and bubble nets.[73]
Fisheries
Chub mackerel, Scomber japonicus, are the most intensively fished scombroid mackerel. They account for about half the total capture production of scombroid mackerels. As a species, they are easily confused with Atlantic mackerel. Chub mackerel migrate long distances in oceans and across the Mediterranean. They can be caught with drift nets and suitable trawls, but are most usually caught with surround nets at night by attracting them with lampara lamps.[74]
The remaining catch of scombroid mackerels is divided equally between the Atlantic mackerel and all other scombroid mackerels. Just these two species (Chub mackerel and Atlantic mackerel) account for about 75% of the total catch of scombroid mackerels.
Chilean jack mackerel are the most commonly fished nonscombroid mackerel, fished as heavily as chub mackerel. The species has been overfished, and its fishery may now be in danger of collapsing.[75] [76]
Smaller mackerel behave like herrings, and are captured in similar ways. Fish species like these, which school near the surface, can be caught efficiently by purse seining. Huge purse-seine vessels use spotter planes to locate the schooling fish. Then they close in using sophisticated sonar to track the shape of the school, which is then encircled with fast auxiliary boats that deploy purse seines as they speed around the school.[77] [78]
Suitably designed trollers can also catch mackerels effectively when they swim near the surface. Trollers typically have several long booms which they lift and drop with "topping lifts". They haul their lines with electric or hydraulic reels.[79] Fish aggregating devices are also used to target mackerel.[80]
Management
The North Sea has been overfished to the point where the ecological balance has become disrupted and many jobs in the fishing industry have been lost.[81]
The Southeast US region spans the Gulf of Mexico, the Caribbean Sea, and the US Southeast Atlantic. Overfishing of king and Spanish mackerel occurred in the 1980s. Regulations were introduced to restrict the size, fishing locations, and bag limits for recreational fishers and commercial fishers. Gillnets were banned in waters off Florida. By 2001, the mackerel stocks had bounced back.[82]
As food
See main article: Mackerel as food. Mackerel is an important food fish that is consumed worldwide.[83] As an oily fish, it is a rich source of omega-3 fatty acids. The flesh of mackerel spoils quickly, especially in the tropics, and can cause scombroid food poisoning. Accordingly, it should be eaten on the day of capture, unless properly refrigerated or cured.[84]
Mackerel preservation is not simple. Before the 19th-century development of canning and the widespread availability of refrigeration, salting and smoking were the principal preservation methods available. Historically in England, this fish was not preserved, but was consumed only in its fresh form. However, spoilage was common, leading the authors of The Cambridge Economic History of Europe to remark: "There are more references to stinking mackerel in English literature than to any other fish!" In France, mackerel was traditionally pickled with large amounts of salt, which allowed it to be sold widely across the country.[85]
For many years mackerel was regarded as 'unclean' in the UK and other places due to folklore which suggested that the fish fed on the corpses of dead sailors.[86] A 1976 survey of housewives in Britain undertaken by the White Fish Authority indicated a reluctance to departing from buying the traditional staples of cod, haddock or salmon. Less than 10% of the survey's 1,931 respondents had ever bought mackerel, and only 3% did so regularly. As a result of this trend, many UK fishmongers during the 1970s did not display or even stock mackerel.[86]
Further reading
External links
Notes and References
- Web site: Mackerel. Scientific name: Scomber scombrus. The Wildlife Trusts. 2022-08-02.
- Web site: Mackerel: to fish or not to fish?. Marine Stewardship Council. 2022-08-02.
- Daan. N.. A quantitative analysis of the food intake of North Sea cod, Gadus Morhua. Netherlands Journal of Sea Research. December 1973. 6. 4. 479–517. 10.1016/0077-7579(73)90002-1. 1973NJSR....6..479D.
- Book: 2008 . King mackerel . 11th . Merriam-Webster's Collegiate Dictionary . Merriam Webster . 9780877798095 . 688 .
- Mackerel . Online Etymology Dictionary . 2012-03-03 . https://web.archive.org/web/20181222125413/https://www.etymonline.com/word/mackerel . 2018-12-22 . live.
- Juan-Jorda . MJ . Mosqueira . I . Cooper . AB . Freire . J . Dulvy . NK . 2011 . Global population trajectories of tunas and their relatives . Proceedings of the National Academy of Sciences . 108 . 51. 20650–20655 . 10.1073/pnas.1107743108 . 22143785 . 3251139 . 2011PNAS..10820650J . free .
- News: Tuna and mackerel populations have reduced by 60% in the last century . 2012-02-08 . ScienceDaily . https://web.archive.org/web/20170707065234/https://www.sciencedaily.com/releases/2012/02/120208103226.htm . 2017-07-07 . live.
- Web site: Species Fact Sheet: Rastrelliger brachysoma (Bleeker, 1851) . . 2 March 2012 . https://web.archive.org/web/20181018194549/http://www.fao.org/fishery/species/2477/en . 2018-10-18 . live.
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Rastrelliger brachysoma . 2011 . e.T170318A6745895 . 10.2305/IUCN.UK.2011-2.RLTS.T170318A6745895.en . 11 November 2021.
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Rastrelliger faughni . 2011 . e.T170324A6748697 . 10.2305/IUCN.UK.2011-2.RLTS.T170324A6748697.en . 11 November 2021.
- Web site: Species Fact Sheet: Rastrelliger kanagurta (Cuvier, 1817) . . 2012-03-02 . https://web.archive.org/web/20180806051043/http://www.fao.org/fishery/species/2478/en . 2018-08-06 . live.
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Rastrelliger kanagurta . 2011 . e.T170328A6750032 . 10.2305/IUCN.UK.2011-2.RLTS.T170328A6750032.en . 11 November 2021.
- Collette, B. . Acero, A. . Canales Ramirez, C. . Cardenas, G. . Carpenter, K.E. . Chang, S.-K. . Chiang, W. . Di Natale, A. . Fox, W. . Guzman-Mora, A. . Juan Jorda, M. . Miyabe, N. . Montano Cruz, R. . Nelson, R. . Salas, E. . Schaefer, K. . Serra, R. . Sun, C. . Uozumi, Y. . Wang, S. . Wu, J. . Yanez, E. . Yeh, S. . 2011 . Scomber australasicus . 2011 . e.T170329A6750490 . 10.2305/IUCN.UK.2011-2.RLTS.T170329A6750490.en . 11 November 2021.
- Collette, B. . Amorim, A.F. . Boustany, A. . Carpenter, K.E. . de Oliveira Leite Jr. . N. . Di Natale, A. . Fox, W. . Fredou, F.L. . Graves, J. . Viera Hazin, F.H. . Juan Jorda, M. . Kada, O. . Minte Vera, C. . Miyabe, N. . Nelson, R. . Oxenford, H. . Teixeira Lessa, R.P. . Pires Ferreira Travassos, P.E. . 2011 . Scomber colias . 2011 . e.T170357A6767497 . 10.2305/IUCN.UK.2011-2.RLTS.T170357A6767497.en . 11 November 2021.
- Web site: Species Fact Sheet: Scomber japonicus (Houttuyn, 1782) . . 2012-03-02 . https://web.archive.org/web/20181221172602/http://www.fao.org/fishery/species/3277/en . 2018-12-21 . live.
- Collette, B. . Acero, A. . Canales Ramirez, C. . Cardenas, G. . Carpenter, K.E. . Chang, S.-K. . Di Natale, A. . Fox, W. . Guzman-Mora, A. . Juan Jorda, M. . Miyabe, N. . Montano Cruz, R. . Nelson, R. . Salas, E. . Schaefer, K. . Serra, R. . Sun, C. . Uozumi, Y. . Wang, S. . Wu, J. . Yeh, S. . 2011 . Scomber japonicus . 2011 . e.T170306A6737373 . 10.2305/IUCN.UK.2011-2.RLTS.T170306A6737373.en . 11 November 2021.
- Web site: Species Fact Sheet: Scomber scombrus (Linnaeus, 1758) . . 2012-03-02 . https://web.archive.org/web/20190502001326/http://www.fao.org/fishery/species/2473/en . 2019-05-02 . live .
- Collette, B. . Boustany, A. . Carpenter, K.E. . Di Natale, A. . Fox, W. . Graves, J. . Juan Jorda, M. . Kada, O. . Nelson, R. . Oxenford, H. . 2011 . Scomber scombrus . 2011 . e.T170354A6764313 . 10.2305/IUCN.UK.2011-2.RLTS.T170354A6764313.en . 11 November 2021.
- Collette, B. . Acero, A. . Amorim, A.F. . Boustany, A. . Canales Ramirez, C. . Cardenas, G. . Carpenter, K.E. . de Oliveira Leite Jr. . N. . Di Natale, A. . Die, D. . Fox, W. . Fredou, F.L. . Graves, J. . Guzman-Mora, A. . Viera Hazin, F.H. . Hinton, M. . Juan Jorda, M. . Kada, O. . Minte Vera, C. . Miyabe, N. . Montano Cruz, R. . Nelson, R. . Oxenford, H. . Restrepo, V. . Salas, E. . Schaefer, K. . Schratwieser, J. . Serra, R. . Sun, C. . Teixeira Lessa, R.P. . Pires Ferreira Travassos, P.E. . Uozumi, Y. . Yanez, E. . 2011 . Acanthocybium solandri . 2011 . e.T170331A6750961 . 10.2305/IUCN.UK.2011-2.RLTS.T170331A6750961.en . 11 November 2021.
- Collette, B. . Fox, W. . Nelson, R. . 2011 . Grammatorcynus bicarinatus . 2011 . e.T170308A6738658 . 10.2305/IUCN.UK.2011-2.RLTS.T170308A6738658.en . 11 November 2021.
- Collette, B. . Chiang, W. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Grammatorcynus bilineatus . 2011 . e.T170358A6768577 . 10.2305/IUCN.UK.2011-2.RLTS.T170358A6768577.en . 11 November 2021.
- Collette, B. . Amorim, A.F. . Boustany, A. . Carpenter, K.E. . de Oliveira Leite Jr. . N. . Di Natale, A. . Fox, W. . Fredou, F.L. . Graves, J. . Viera Hazin, F.H. . Juan Jorda, M. . Minte Vera, C. . Miyabe, N. . Nelson, R. . Oxenford, H. . Teixeira Lessa, R.P. . Pires Ferreira Travassos, P.E. . 2011 . Scomberomorus brasiliensis . 2011 . e.T170335A6753567 . 10.2305/IUCN.UK.2011-2.RLTS.T170335A6753567.en . 11 November 2021.
- Web site: Species Fact Sheet: Scomberomorus cavalla (Cuvier, 1829) . . 2012-03-02 . https://web.archive.org/web/20181015151018/http://www.fao.org/fishery/species/2483/en . 2018-10-15 . live.
- Collette, B. . Amorim, A.F. . Boustany, A. . Carpenter, K.E. . de Oliveira Leite Jr. . N. . Di Natale, A. . Fox, W. . Fredou, F.L. . Graves, J. . Viera Hazin, F.H. . Juan Jorda, M. . Minte Vera, C. . Miyabe, N. . Nelson, R. . Oxenford, H. . Teixeira Lessa, R.P. . Pires Ferreira Travassos, P.E. . 2011 . Scomberomorus cavalla . 2011 . e.T170339A6755835 . 10.2305/IUCN.UK.2011-2.RLTS.T170339A6755835.en . 11 November 2021.
- Web site: Species Fact Sheet: Scomberomorus commerson (Lacepède, 1800) . . 2012-03-02 . https://web.archive.org/web/20181016014535/http://www.fao.org/fishery/species/3280/en . 2018-10-16 . live .
- Collette, B. . Chang, S.-K. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Miyabe, N. . Nelson, R. . 2011 . Scomberomorus commerson . 2011 . e.T170316A6745396 . 10.2305/IUCN.UK.2011-2.RLTS.T170316A6745396.en . 11 November 2021.
- Collette, B. . Acero, A. . Boustany, A. . Canales Ramirez, C. . Cardenas, G. . Carpenter, K.E. . Di Natale, A. . Die, D. . Fox, W. . Graves, J. . Hinton, M. . Juan Jorda, M. . Minte Vera, C. . Miyabe, N. . Montano Cruz, R. . Nelson, R. . Restrepo, V. . Schaefer, K. . Schratwieser, J. . Serra, R. . Yanez, E. . 2011 . Scomberomorus concolor . 2011 . e.T20047A9138383 . 10.2305/IUCN.UK.2011-2.RLTS.T20047A9138383.en . 11 November 2021.
- Web site: Species Fact Sheet: Scomberomorus guttatus (Bloch & Schneider, 1801) . . 2012-03-02 . https://web.archive.org/web/20181009102312/http://www.fao.org/fishery/species/3281/en . 2018-10-09 . live .
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Scomberomorus guttatus . 2011 . e.T170311A6742170 . 10.2305/IUCN.UK.2011-2.RLTS.T170311A6742170.en . 11 November 2021.
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Scomberomorus koreanus . 2011 . e.T170343A6757043 . 10.2305/IUCN.UK.2011-2.RLTS.T170343A6757043.en . 11 November 2021.
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Scomberomorus lineolatus . 2011 . e.T170353A6764121 . 10.2305/IUCN.UK.2011-2.RLTS.T170353A6764121.en . 11 November 2021.
- Web site: Species Fact Sheet: Scomberomorus maculatus (Mitchill, 1815) . . 2012-03-02 . https://web.archive.org/web/20180724053321/http://www.fao.org/fishery/species/3282/en . 2018-07-24 . live.
- Collette, B. . Boustany, A. . Carpenter, K.E. . Fox, W. . Graves, J. . Juan Jorda, M. . Nelson, R. . Oxenford, H. . 2011 . Scomberomorus maculatus . 2011 . e.T170323A6748550 . 10.2305/IUCN.UK.2011-2.RLTS.T170323A6748550.en . 11 November 2021.
- Collette, B. . Carpenter, K.E. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Scomberomorus multiradiatus . 2011 . e.T170347A6760831 . 10.2305/IUCN.UK.2011-2.RLTS.T170347A6760831.en . 11 November 2021.
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Scomberomorus munroi . 2011 . e.T170330A6750789 . 10.2305/IUCN.UK.2011-2.RLTS.T170330A6750789.en . 11 November 2021.
- Web site: Species Fact Sheet: Scomberomorus niphonius (Cuvier, 1831) . . 2012-03-02 . https://web.archive.org/web/20180112140508/http://www.fao.org/fishery/species/3285/en . 2018-01-12 . live.
- Collette, B. . Chang, S.-K. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . Uozumi, Y. . 2011 . Scomberomorus niphonius . 2011 . e.T170356A6767322 . 10.2305/IUCN.UK.2011-2.RLTS.T170356A6767322.en . 11 November 2021.
- Collette, B. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Scomberomorus plurilineatus . 2011 . e.T170348A6760962 . 10.2305/IUCN.UK.2011-2.RLTS.T170348A6760962.en . 11 November 2021.
- Collette, B. . Di Natale, A. . Dooley, J. . Fox, W. . Fritzsche, R. . Juan Jorda, M. . Matsuura, K. . Nelson, J. . Nelson, R. . 2011 . Scomberomorus queenslandicus . 2011 . e.T154946A4674361 . 10.2305/IUCN.UK.2011-2.RLTS.T154946A4674361.en . 11 November 2021.
- Collette, B. . Amorim, A.F. . Boustany, A. . Carpenter, K.E. . de Oliveira Leite Jr. . N. . Di Natale, A. . Fox, W. . Fredou, F.L. . Graves, J. . Viera Hazin, F.H. . Juan Jorda, M. . Minte Vera, C. . Miyabe, N. . Nelson, R. . Oxenford, H. . Teixeira Lessa, R.P. . Pires Ferreira Travassos, P.E. . 2011 . Scomberomorus regalis . 2011 . e.T170327A6749725 . 10.2305/IUCN.UK.2011-2.RLTS.T170327A6749725.en . 11 November 2021.
- Collette, B. . Carpenter, K.E. . Di Natale, A. . Fox, W. . Juan Jorda, M. . Nelson, R. . 2011 . Scomberomorus semifasciatus . 2011 . e.T170337A6754351 . 10.2305/IUCN.UK.2011-2.RLTS.T170337A6754351.en . 11 November 2021.
- Collette, B. . Amorim, A.F. . Boustany, A. . Carpenter, K.E. . de Oliveira Leite Jr. . N. . Di Natale, A. . Fox, W. . Fredou, F.L. . Viera Hazin, F.H. . Juan Jorda, M. . Kada, O. . Minte Vera, C. . Miyabe, N. . Nelson, R. . Oxenford, H. . Teixeira Lessa, R.P. . Pires Ferreira Travassos, P.E. . 2011 . Scomberomorus tritor . 2011 . e.T170326A6749128 . 10.2305/IUCN.UK.2011-2.RLTS.T170326A6749128.en . 11 November 2021.
- Poulin . E . Cárdenas . L . Hernández . CE . Kornfield . I . Ojeda . FP . 2004 . Resolution of the taxonomic status of Chilean and Californian jack mackerels using mitochondrial DNA sequence . Journal of Fish Biology . 65 . 4. 1160–1164 . 10.1111/j.0022-1112.2004.00514.x . 2004JFBio..65.1160P .
- Collette, B. . Boustany, A. . Carpenter, K.E. . Di Natale, A. . Fox, W. . Graves, J. . Juan Jorda, M. . Miyabe, N. . Nelson, R. . Oxenford, H. . Uozumi, Y. . 2011 . Gasterochisma melampus . 2011 . e.T170340A6756181 . 10.2305/IUCN.UK.2011-2.RLTS.T170340A6756181.en . 11 November 2021.
- Web site: Species Fact Sheet: Trachurus trachurus (Linnaeus, 1758) . . 2012-03-02 . https://web.archive.org/web/20180716221543/http://www.fao.org/fishery/species/2306/en . 2018-07-16 . live.
- Smith-Vaniz, B. . Robertson, R. . Dominici-Arosemena, A. . 2010 . Trachurus murphyi . 2010 . e.T183965A8207652 . 10.2305/IUCN.UK.2010-3.RLTS.T183965A8207652.en . 11 November 2021.
- Web site: Species Fact Sheet: Trachurus capensis (Castelnau, 1861) . . 2012-03-02 . https://web.archive.org/web/20171126111348/http://www.fao.org/fishery/species/3105/en . 2017-11-26 . live.
- Web site: Species Phallomedusa solida (Martens, 1878) . dead . https://web.archive.org/web/20120712095240/http://www.environment.gov.au/biodiversity/abrs/online-resources/fauna/afd/taxa/Phallomedusa_solida . 2012-07-12 . Department of the Environment, Water, Heritage and the Arts, Australian Biological Resources Study . 2012-03-02 .
- Web site: Species Fact Sheet: Trachurus murphyi (Nichols, 1920) . . 2012-03-02 . https://web.archive.org/web/20181015012246/http://www.fao.org/fishery/species/2309/en . 2018-10-15 . live.
- Web site: Species Fact Sheet: Trachurus trecae (Cadenat, 1949) . . 2012-03-02 . https://web.archive.org/web/20180724042410/http://www.fao.org/fishery/species/3106/en . 2018-07-24 . live.
- Web site: Species Fact Sheet: Trachurus declivis (Jenyns, 1841) . . 2012-03-02 . https://web.archive.org/web/20180330125236/http://www.fao.org/fishery/species/2313/en . 2018-03-30 . live.
- Web site: Phallomedusa solida (Martens, 1878) . dead . https://web.archive.org/web/20120712095240/http://www.environment.gov.au/biodiversity/abrs/online-resources/fauna/afd/taxa/Phallomedusa_solida . 2012-07-12 . Department of the Environment, Water, Heritage and the Arts, Australian Biological Resources Study . 2012-03-02 .
- Web site: Species Fact Sheet: Trachurus japonicus (Temminck & Schlegel, 1844) . . 2012-03-02 . https://web.archive.org/web/20170730140855/http://www.fao.org/fishery/species/2308/en . 2017-07-30 . live.
- Web site: Species Fact Sheet: Trachurus mediterraneus (Steindachner, 1868) . . 2012-03-02 . https://web.archive.org/web/20190328231000/http://www.fao.org/fishery/species/2311/en . 2019-03-28 . live.
- Smith-Vaniz, B. . Robertson, R. . Dominici-Arosemena, A. . 2010 . Trachurus symmetricus . 2010 . e.T183729A8166054 . 10.2305/IUCN.UK.2010-3.RLTS.T183729A8166054.en . 11 November 2021.
- Web site: Species Fact Sheet: Pleurogrammus azonus (Jordan & Metz, 1913) . . 2012-03-02 . https://web.archive.org/web/20180214082907/http://www.fao.org/fishery/species/2534/en . 2018-02-14 . live.
- Web site: Glossary of definitions of cat terms for the breeder . Cats online . 2012-04-07 . https://web.archive.org/web/20190109122235/http://www.showcatsonline.com/x/glossary.shtml#M . 2019-01-09 . live.
- Book: Metereology of Clouds. 154. Downing, L. L.. 2013. AuthorHouse . 9781491804339.
- Book: Metereology Today. C. Donald . Ahrens. Robert. Henson. 2015. Cengage Learning. 153. 9781305480629.
- Web site: Atlantic mackerel . National Oceanic and Atmospheric Administration . FishWatch . 2012-03-12 . https://web.archive.org/web/20120320184708/http://www.fishwatch.gov/seafood_profiles/species/mackerel/species_pages/atlantic_mackerel.htm . 2012-03-20 . dead.
- Denton . EJ . Rowe . DM . 1998 . Bands against stripes on the backs of mackerel, Scomber scombrus L. . Proc. R. Soc. Lond. B . 265 . 1401 . 1051–1058 . 1689176 . 10.1098/rspb.1998.0398 .
- Shaw . E . Tucker . A . 1965 . The optomotor reaction of schooling carangid fishes . Animal Behaviour . 13 . 2–3. 330–336 . 10.1016/0003-3472(65)90052-7. 5835850 .
- Book: Bone . Q . Moore . RH . 2008 . Biology of Fishes . Taylor & Francis Group . 418–422 . 978-0-415-37562-7.
- Rowe . DM . Denton . EJ . Eric James Denton. 1997 . The physical basis of reflective communication between fish, with special reference to the horse mackerel, Trachurus trachurus . Phil. Trans. R. Soc. Lond. B . 352 . 1353. 531–549 . 1691948 . 10.1098/rstb.1997.0037. 1997RSPTB.352..531R .
- Book: Lal . BV . Fortune . K . 2000 . The Pacific Islands: An encyclopedia . University of Hawaii Press . 8 . 9780824822651 .
- Block . BA . Finnerty . JR . 1993 . Endothermy in fishes: a phylogenetic analysis of constraints, predispositions, and selection pressures . Environmental Biology of Fishes . 40 . 3 . 283–302 . 10.1007/BF00002518 . 28644501 .
- Wardle . CS . He . P . 1988 . Burst swimming speeds of mackerel, Scomber scombrus . Journal of Fish Biology . 32 . 3. 471–478 . 10.1111/j.1095-8649.1988.tb05382.x. 1988JFBio..32..471W .
- Wardle . CS . He . P . 1988 . Endurance at intermediate swimming speeds of Atlantic mackerel, Scomber scombrus L., herring, Clupea harengus L., and saithe, Pollachius virens L . Journal of Fish Biology . 33 . 2. 255–266 . 10.1111/j.1095-8649.1988.tb05468.x. 1988JFBio..33..255H .
- Web site: FAO Fact Sheet: Biological characteristics of tuna . 2012-03-06 . https://web.archive.org/web/20190205223834/http://www.fao.org/fishery/topic/16082/en . 2019-02-05 . live.
- Book: International Wildlife Encyclopedia . Burton, R. . Marshall Cavendish . 2002 . 978-0-7614-7266-7.
- Web site: Forage species . . 2019-05-06 . https://web.archive.org/web/20181221041523/http://www.fao.org/fishery/topic/4290/en . 2018-12-21 . live .
- Book: Compagno, L.J.V. . 1984 . Sharks of the World: An Annotated and Illustrated Catalogue of Shark Species Known to Date . Rome . Food and Agricultural Organization . 978-92-5-101384-7 . 466–468.
- Web site: Bryde's Whale (Balaenoptera edeni) . Noaa Fisheries, Office of Protected Resources . 2009-12-31 . https://web.archive.org/web/20100619094748/http://www.nmfs.noaa.gov/pr/species/mammals/cetaceans/brydeswhale.htm . 2010-06-19 . dead .
- Web site: Chub mackerel . https://web.archive.org/web/20130313192422/http://www.sicilianfishontheroad.com/en/il-pescato-siciliano/il-pesce-azzurro/lanzardo . 2013-03-13 . dead . Sicilian Fish on the Road . 2012-04-06 .
- News: In mackerel's plunder, hints of epic fish collapse . 2012-01-25 . The New York Times . https://web.archive.org/web/20181221043313/https://www.nytimes.com/2012/01/25/science/earth/in-mackerels-plunder-hints-of-epic-fish-collapse.html?_r=1&pagewanted=all . 2018-12-21 . live.
- News: Lords of the fish . 2012-01-25 . dead . iWatch News . https://web.archive.org/web/20120320194143/http://www.iwatchnews.org/2012/01/25/7914/lords-fish . 2012-03-20 .
- Web site: Fishing vessel types: Purse seiners . . 2019-05-06 . https://web.archive.org/web/20190506095459/http://www.fao.org/fishery/vesseltype/140/en . 2019-05-06 . live.
- Book: Gabriel . O . von Brandt . A . Lange . K . Dahm . E . Wendt . T . 2005 . Seining in fresh and sea water . Fish catching methods of the world . Wiley-Blackwell . 9780852382806 . 431–448 .
- Web site: Fishing Vessel type: Trollers . . https://web.archive.org/web/20190403212959/http://www.fao.org/fishery/vesseltype/360/en . 2019-04-03 . live.
- Web site: The FAD FAQ . 2019-05-06 . https://web.archive.org/web/20181029004642/http://www.himb.hawaii.edu/FADS/ . 2018-10-29 . live.
- Book: Clover . Charles . 2004 . The End of the Line: How overfishing is changing the world and what we eat . London . Ebury Press . 0-09-189780-7 .
- Web site: [ftp://ftp.fao.org/FI/DOCUMENT/fcp/en/FI_CP_US.pdf FISHERY COUNTRY PROFILE: THE UNITED STATES OF AMERICA ]. https://web.archive.org/web/20170518220246/ftp://ftp.fao.org/FI/DOCUMENT/fcp/en/FI_CP_US.pdf . dead . 2017-05-18 . .
- Book: The California mackerel fishery . Croker . Richard Symonds . 1933 . Division of Fish and Game of California . Contribution No. 125 from the California State Fisheries Laboratory . 9–10.
- Web site: November 2007 . Scombrotoxin (Histamine) . dead . Food Safety Watch . https://web.archive.org/web/20120713020029/http://www.foodsafetywatch.com/public/484.cfm . 2012-07-13 .
- Book: Clapham . JH . Postan . MM . Rich . EE . 1941 . The Cambridge economic history of Europe . CUP Archive . 166–168 . 978-0-521-08710-0.
- Web site: McFarlane. Andrew. 2010-08-24. Why is Britain braced for a mackerel war?. 2022-08-02. BBC News. en.