Largest and heaviest animals explained

The largest animal currently alive is the blue whale. The maximum recorded weight was 190 tonnes for a specimen measuring 27.6m (90.6feet), whereas longer ones, up to 33m (108feet), have been recorded but not weighed.[1] [2] [3] It is estimated that this individual could have a mass of 250 tonnes or more.[4] [5] [6] The longest non-colonial animal is the lion's mane jellyfish .[7]

In 2023, paleontologists estimated that the extinct whale Perucetus, discovered in Peru, may have outweighed the blue whale, with a mass of .[8] However, more recent studies suggest this whale was much smaller than previous estimates putting its weight at 60 to 113 tonnes.[9] While controversial, estimates for the weight of the sauropod Bruhathkayosaurus suggest it was around 110–170 tons, with the highest estimate being 240 tons, if scaled with Patagotitan, although actual fossil remains no longer exist, and that estimation is based on described dimensions in 1987. In April 2024, Ichthyotitan severnensis was established as a valid shastasaurid taxon and is considered both the largest marine reptile ever discovered and the largest macropredator ever discovered. The Lilstock specimen was estimated to be around 26m (85feet) whilst the Aust specimen was an even more impressive 30mto35mm (100feetto115feetm) in length. Whilst no weight estimates was made as of yet, Ichthyotitan would have easily rivalled or surpassed the blue whale.[10] [11] The upper estimates of weight for these prehistoric animals would have easily rivaled or exceeded the largest rorquals and sauropods.[12]

The African bush elephant (Loxodonta africana) is the largest living land animal. A native of various open habitats in sub-Saharan Africa, males weigh about 6tonne on average.[13] The largest elephant ever recorded was shot in Angola in 1974. It was a male measuring 10.67m (35.01feet) from trunk to tail and 4.17m (13.68feet) lying on its side in a projected line from the highest point of the shoulder, to the base of the forefoot, indicating a standing shoulder height of 3.96m (12.99feet). This male had a computed weight of 10.4 to 12.25 tonnes.[1]

Heaviest living animals

The heaviest living animals are all whales. Since no scale can accommodate the whole body of a large whale, most have been weighed by parts.

RankAnimalAverage mass
[tonnes]		Maximum mass
[tonnes]		Average total length
[m (ft)]
1 Blue whale[14] 110[15] 19024 (79)[16]
2 60[17] 120 15.5 (51)
3 58 110[18] 15.25 (50)
4 57 120 19.5 (64)
5 54.5[19] 120 15 (49)
6 54[20] 110[21] 15 (49)
7 31.25[22] 57[23] 13.25 (43.5)
8 29[24] 48[25] 13.5 (44)
9 22.5 45[26] 14.8 (49)
10 19.5 45[27] 13.5 (44)

Heaviest terrestrial animals

The heaviest land animals are all mammals. The African elephant is now listed as two species, the African bush elephant and the African forest elephant, as they are now generally considered to be two separate species.[28]

RankAnimalAverage mass
[tonnes]		Maximum mass
[tonnes]		Average total length
[m (ft)]
1 6[29] [30] 10.4[31] 7 (23)[32]
2 4.5[33] 8.15 6.8 (22.3)
3 2.7[34] 6.0 6.2 (20.3)[35]
4 2[36] [37] 4.5[38] 4.4 (14.4)[39]
5 1.9[40] [41] 4.0[42] 4.2 (13.8)[43]
6 1.8[44] [45] 4.5[46] 5.05 (16.5)[47]
7 1.75[48] [49] 2.3[50] 3.8 (12.5)[51]
8 1.1[52] 2.9[53] 4 (13.1)[54]
9 1.0 2[55] 5.15 (16.9)[56]
10 0.95[57] 1.5 3.8 (12.5)[58]

Vertebrates

Mammals (Mammalia)

See main article: List of largest mammals. The blue whale is the largest mammal of all time, with the longest known specimen being 331NaN1 long and the heaviest weighted specimen being 190 tonnes.[59] [60] The extinct whale species Perucetus colossus was shorter than the blue whale, at 17.0–20.1 meters (55.8–65.9 ft) but it is estimated to have rivaled or surpassed it in weight, at 85–340 tonnes. At the highest estimates, this would make Perucetus the heaviest known animal in history.

The largest land mammal extant today is the African bush elephant. The largest extinct land mammal known was long considered to be Paraceratherium orgosensis, a rhinoceros relative thought to have stood up to 4.81NaN1 tall, measured over 7.41NaN1 long and may have weighed about 17 tonnes.[61] In 2015, a study suggested that one example of the proboscidean Palaeoloxodon namadicus may have been the largest land mammal ever, based on extensive research of fragmentary leg bone fossils from one individual, with a maximum estimated size of 22 tonnes.[62] [63]

Stem-mammals (Synapsida)

The Late Triassic Lisowicia bojani, from what is now southern Poland, probably was the largest of all non-mammalian synapsids (most of which became extinct 250 million years ago), at 4.5m (14.8feet) in length, 2.6m (08.5feet) in height and in weight. However, one study suggested a more conservative weight of 4.87 tonnes to 7.02 tonnes for the adult taxon, with an average body mass of 5.88 tonnes.[64] The largest carnivorous synapsid was Anteosaurus at 5– and 500–.[65]

Caseasaurs (Caseasauria)
  • The herbivorous Alierasaurus was the largest caseid and the largest amniote to have lived at the time, with an estimated length around .[66] Another huge caseasaur is Cotylorhynchus hancocki, with an estimated length and weight of at least [67] and more than .[68]
    Sphenacodontids (Sphenacodontidae)
  • The biggest carnivorous synapsid of Early Permian was Dimetrodon, which could reach and .[69] The largest members of the genus Dimetrodon were also the world's first fully terrestrial apex predators.[70]
    Tappenosaurids (Tappenosauridae)
  • The Middle Permian Tappenosaurus was estimated at in length which is comparable in size with the largest dinocephalians.[71]
    Therapsids (Therapsida)
  • The plant-eating dicynodont Lisowicia bojani is the largest-known of all non-mammalian synapsids, at and .[72] [73] [74] The largest carnivorous therapsid was the aforementioned Anteosaurus from what is now South Africa during Middle Permian epoch. It reached 5– long, and about 500– in weight.[65]

    Reptiles (Reptilia)

    See main article: List of largest reptiles. The largest living reptile, a representative of the order Crocodilia, is the saltwater crocodile (Crocodylus porosus) of Southern Asia and Australia, with adult males being typically 3.9– long. The largest confirmed saltwater crocodile on record was 6.321NaN1 long, and weighed about 1360kg (3,000lb).[1] Unconfirmed reports of much larger crocodiles exist, but examinations of incomplete remains have never suggested a length greater than 7m (23feet).[75] Also, a living specimen estimated at 7m (23feet) and 2000kg (4,000lb) has been accepted by the Guinness Book of World Records.[76] However, due to the difficulty of trapping and measuring a very large living crocodile, the accuracy of these dimensions has yet to be verified. A specimen named Lolong caught alive in the Philippines in 2011 (died February 2013) was found to have measured 6.17m (20.24feet) in length.[77] [78] [79] [80] [81]

    The Komodo dragon (Varanus komodoensis), also known as the "Komodo monitor", is a large species of lizard found in the Indonesian islands of Komodo, Rinca, Flores, Gili Motang, Nusa kode and Padar. A member of the monitor lizard family (Varanidae), it is the largest living species of lizard, growing to a maximum length of more than 3m (10feet) in rare cases and weighing up to approximately 166kg (366lb).[82]

    Nevertheless, current extant reptiles are still dwarfed by their prehistoric ancestors. The largest non-dinosaurian terrestrial reptile to have ever lived were crocodilomorphs such as Deinosuchus, Sarcosuchus and Purussaurus, each have weight varying between 5-10 metric tons.[83] [84] [85] The largest snake to have ever lived was Titanoboa which could grow up to 12.8– and weigh up to 730-.[86] Prehistoric marine reptiles from the Mesozoic were even larger. Archelon was the largest testudines, being around long from head to tail and in weight.[87] Mosasaurus hoffmanni was the largest squamate lizard to have ever lived, with a length of 11m (36feet) and a body mass of 10MT[88] However, the largest reptiles, period, were the shastasaurids, specifically Ichthyotitan, which approached lengths rivalling to exceeding those of a blue whale, at around 26– in length.[10] [11]

    Largest living reptiles

    The following is a list of the largest living reptile species ranked by average weight, which is dominated by the crocodilians. Unlike mammals, birds, or fish, the mass of large reptiles is frequently poorly documented and many are subject to conjecture and estimation.[1]

    RankAnimalAverage mass
    [kg (lb)]    		Maximum mass
    [kg (lb)]    		Average total length
    [m (ft)]
    1 450 (1,000)[89] [90] 2,000 (4,409 lbs)[91] [92] 4.5 (14.8)[93]
    2 410 (900)[94] 1,090 (2,400) 4.2 (13.8)
    3 380 (840) 1,100 (2,400) 4.1 (13.5)[95] [96]
    4 364 (800)[97] [98] 932 (2,050) 2.0 (6.6)
    5 336 (740)[99] 1,000 (2,200)[100] 4.0 (13.1)[101] [102]
    6 300 (661) 1,000 (2,200)3.9 (12.8)[103] [104] [105] [106]
    7 250 (550)[107] 1,000 (2,200)[108] 4.5 (14.8)
    8 240 (530)[109] [110] 1,000 (2,200) 3.4 (11.2)
    9 225 (495) 700 (1,500)[111] 3.3 (10.8)
    10 210 (460)[112] 590 (1,300)[113] 4.0 (13.1)[114]
    11 205 (450)[115] 360 (790) 1.4 (4.6)[116]
    12 200 (441) 545 (1,202)0.95 (3.2)
    13 190 (418.9)[117] 395 (870.8)1.12 (3.67)
    14 180 (400)[118] 325 (720) 3.3 (10.8)
    15 175 (390)[119] 417 (919)[120] 1.5 (4.9)[121]

    Dinosaurs (Dinosauria)

    See main article: Dinosaur size.

    See also: Largest prehistoric animals.

    Dinosaurs are now extinct, except for birds, which are theropods.

    Sauropods (Sauropoda)
  • The largest dinosaurs, and the largest animals to ever live on land, were the plant-eating, long-necked Sauropoda. The tallest and heaviest sauropod known from a complete skeleton is a specimen of an immature Giraffatitan discovered in Tanzania between 1907 and 1912, now mounted in the Museum für Naturkunde of Berlin. It is NaNm (-2,147,483,648feet) tall and weighed 23.3–39.5 tonnes.[122] [123] [124] [125] [126] [127] The longest is a 25m (82feet) long specimen of Diplodocus discovered in Wyoming, and mounted in Pittsburgh's Carnegie Natural History Museum in 1907. A Patagotitan specimen found in Argentina in 2014 is estimated to have been NaNm (-2,147,483,648feet) long and 20m (70feet) tall, with a weight of 69–77 tonnes.[128] [129]
  • There were larger sauropods, but they are known only from a few bones. The current record-holders include Argentinosaurus, which may have weighed 100 tonnes; Supersaurus which might have reached 34m (112feet) in length and Sauroposeidon which might have been 18m (59feet) tall. Some abnormal specimens such as specimen BYU 9024 of the Barosaurus/Supersaurus genus could reach an astounding 45-50 meters long,[130] with mass varying from the 'modest' 60-66 tons to the more immense 92-120 tons. Two other such sauropods include Bruhathkayosaurus and Maraapunisaurus. Both are known only from fragments that no longer exist. Bruhathkayosaurus might have been between 40– in length and 175–220 tonnes in weight according to some estimates, with recent estimates being place between 110-170 tons.[130] [131] [132] Maraapunisaurus might have been approximately 35–40 m long and 80–120 tonnes or more.[133] Each of these two 'super-sauropods' would have easily rivalled the largest blue whale in size.[132]
    RankAnimalAverage mass
    [tonnes]    		Maximum mass
    [tonnes]    		Average total length
    [m (ft)]
    1 Bruhathkayosaurus matleyi110-170 240 44.1-45 (144.6-148)[134] [135]
    2 Maraapunisaurus fragillimus 80-120[136] 150[137] 35-40 (115–131)
    3 Barosaurus lentus/Supersaurus vivianae BYU 9024 60-66[138] [139] [140] 92-120[141] 45-50 (148–160)
    4 Argentinosaurus huinculensis 75-80 100 35–39.7 (115–130)[142]
    5 Mamenchisaurus 50-80 80 26–35 (85–115)[143]
    6 Notocolossus gonzalezparejasi 44.9–75.9 75.9 28 (92)[144]
    7 Patagotitan mayorum 55-69 77 33–37 (108–121)[145]
    8 Puertasaurus reuili 50-60[146] 60 27-30 (89–98)[147] [148] [149]
    9 Sauroposeidon proteles 40-60 60 27–34 (89–112)[150] [151]
    10 Dreadnoughtus schrani 22.1–59.3 59.3 26 (85)[152]
    Theropods (Theropoda)
  • The largest theropod known from a nearly complete skeleton is the most complete Tyrannosaurus rex specimen, nicknamed "Sue", which was discovered in South Dakota in 1990 and now mounted in the Field Museum of Chicago at a total length of 12.3m (40.4feet). Body mass estimates have reached over 9,500 kg,[153] though other figures, such as Hartman's 2013 estimate of 8,400 kg,[154] have been lower.
  • Another giant theropod is Spinosaurus aegyptiacus from the mid-Cretaceous of North Africa. Size estimates have been fluctuating far more over the years, with length estimates ranging from 12.6 to 18 m and mass estimates from 7 to 20.9 t.[155] [156] Recent findings favor a length exceeding 15 m [157] and a body mass of 7.5 tons.[158]
  • Other contenders known from partial skeletons include Giganotosaurus carolinii (est. 12.2–13.2 m and 6-13.8 tonnes) and Carcharodontosaurus saharicus (est. 12-13.3 m and 6.2-15.1 tonnes).[156] [159] [160] [161] [162] [163] Saurophaganax maximus is another contender for largest theropod since the bones of the largest specimens (tibia, centrum and atlas) are reportedly as large or even larger than the same bones in Giganotosaurus, though these remains are yet to be properly described.[164]
  • The largest extant theropod is the common ostrich (see birds, below).
    Armored dinosaurs (Thyreophora)
  • The largest thyreophorans were Ankylosaurus and Stegosaurus, from the Late Cretaceous and Late Jurassic periods (respectively) of what is now North America, both measuring up to 9m (30feet) in length and estimated to weigh up to 6 tonnes.[165] [166]
    Ornithopods (Ornithopoda)
  • The largest ornithopods were the hadrosaurids Shantungosaurus, a late Cretaceous dinosaur found in the Shandong Peninsula of China, and Magnapaulia from the late Cretaceous of North America. Both species are known from fragmentary remains but are estimated to have reached over 15m (49feet) in length[167] [168] and were likely the heaviest non-sauropod dinosaurs, estimated at over 23 tonnes.[168]
    Ceratopsians (Ceratopsia)
  • The largest ceratopsians were Triceratops and its ancestor Eotriceratops from the late Cretaceous of North America. Both estimated to have reached about 9m (30feet) in length[169] and weighed 12 tonnes.[170] [171]

    Birds (Aves)

    See main article: List of largest birds. The largest living bird, a member of the Struthioniformes, is the common ostrich (Struthio camelus), from the plains of Africa. A large male ostrich can reach a height of 2.8m (09.2feet) and weigh over 156kg (344lb).[172] A mass of 200kg (400lb) has been cited for the common ostrich but no wild ostriches of this weight have been verified.[173] Eggs laid by the ostrich can weigh 1.4kg (03.1lb) and are the largest eggs in the world today.

    The largest bird in the fossil record may be the extinct elephant birds (Aepyornithidae) of Madagascar, which were related to the kiwis. Aepyornis exceeded 3m (10feet) in height and 500kg (1,100lb), while Vorombe could reach a similar height and a mass of 732kg (1,614lb).[174] The last of the elephant birds became extinct about 300 years ago. Of almost exactly the same upper proportions as the largest elephant birds was Dromornis stirtoni of Australia, part of a 26,000-year-old group called mihirungs of the family Dromornithidae.[175] The largest carnivorous bird was Brontornis, an extinct flightless bird from South America which reached a weight of 350to and a height of about 2.8m (09.2feet).[176] The tallest carnivorous bird was Kelenken, which could reach 3 to 3.2 meters in height and 220 to 250 kilograms. The tallest bird ever was the giant moa (Dinornis maximus), part of the moa family of New Zealand that went extinct around 1500 AD. This particular species of moa stood up to 3.7m (12.1feet) tall,[172] but weighed about half as much as a large elephant bird or mihirung due to its comparatively slender frame.[1]

    The heaviest bird ever capable of flight was Argentavis magnificens, the largest member of the now extinct family Teratornithidae, found in Miocene-aged fossil beds of Argentina, with a wingspan up to 5.5m (18feet), a length of up to 1.25m (04.1feet), a height on the ground of up to 1.75m (05.74feet) and a body weight of at least 71kg (157lb).[1] [177] [178] Pelagornis sandersi is thought to have had an even larger wingspan of about 6.1-, but is only about 22-, half the mass of the former.[179] [178]

    Heaviest living bird species

    The following is a list of the heaviest living bird species based on maximum reported or reliable mass, but average weight is also given for comparison. These species are almost all flightless, which allows for these particular birds to have denser bones and heavier bodies. Flightless birds comprise less than 2% of all living bird species.

    RankAnimalBinomial nameAverage mass
    [kg (lb)]    		Maximum mass
    [kg (lb)]    		Average total length
    [cm (ft)]    		Flighted
    1 Struthio camelus104 (230)[180] 156.8 (346) 210 (6.9)[181] No
    2 Struthio molybdophanes90 (200) 130 (287) 200 (6.6) No
    3 Casuarius casuarius45 (99) 85 (190)[182] 155 (5.1) No
    4 Casuarius unappendiculatus44 (97) 75 (170) 149 (4.9) No
    5 Dromaius novaehollandiae33 (73)[183] 70 (150) 153 (5) No
    6 Aptenodytes forsteri31.5 (69)[184] 46 (100) 114 (3.7) No
    7 Rhea americana23 (51) 40 (88) 134 (4.4) No
    8 Meleagris gallopavo13.5 (29.8) [185] 39 (86)[186] 100 - 124.9 (3.3 – 4.1) Yes
    9 Casuarius bennetti19.7 (43) 34 (75) 105 (3.4) No
    10 Rhea pennata19.6 (43) 28.6 (63) 96 (3.2) No
    11 Cygnus olor11.87 (26.2) 23 (51) 100-130 (3.3 - 4.3) Yes
    12 Otis tarda10.6 (23.4) 21 (46) 115 (3.8) Yes
    13 Aptenodytes patagonicus13.6 (30) 20 (44)[187] 92 (3) No
    14 Ardeotis kori11.4 (25.1) 20 (44.1) 150 (5) Yes
    15 Cygnus buccinator11.6 (25.1) 17.2 (38) 138 - 165 (4.5 - 5.4) Yes
    16 Diomedea exulans11.9 (24) 16.1 (38)[188] 107 - 135 (3.5 - 4.4) Yes
    17 Cygnus cygnus11.4 (25) 15.5 (32) 140 - 165 (4.5 - 5.4) Yes
    18 Pelecanus crispus11.5 (25) 15 (33.1) 183 (6) Yes
    19 Vultur gryphus11.3 (25)[189] 14.9 (33) 100 - 130 (3.3 - 4.3) Yes

    Amphibians (Amphibia)

    The largest living amphibian is the South China giant salamander (Andrias sligoi). Formerly considered conspecific with the Chinese giant salamander (A. davidianus), the maximum size of this nearly human-sized river-dweller is 64kg (141lb) and almost 1.83m (06feet).[1] Before amniotes became the dominant tetrapods, several giant amphibian proto-tetrapods existed and were certainly the dominant animals in their ecosystems. The largest known was the crocodile-like Prionosuchus, which reached a length of 9m (30feet).[190]

    Frogs (Anura)
  • The largest member of the largest order of amphibians is the African goliath frog (Conraua goliath). The maximum size this species is verified to attain is a weight of 3.8kg (08.4lb) and a snout-to-vent length of 39cm (15inches).[1] The largest of the toads, the cane toad (Rhinella marina), is also the second largest member of the frog order. This infamous, often invasive species can grow to maximum mass of 2.65kg (05.84lb) and measure a maximum of 33cm (13inches) from snout-to-vent.[1] Rivaling the previous two species, the African bullfrog (Pyxicephalus adspersus) can range up to a weight of 2kg (04lb) and 25.5cm (10inches) from snout to vent.[191] Another large frog is the largest frog in North America, the American bullfrog, which can reach weights of up to 0.8kg (01.8lb) and snout-to-vent-length (SVL) of 20cm (10inches). However, the toad Beelzebufo ampinga, found in fossil from the Cretaceous era in what is now Madagascar, was estimated to grow to 41cm (16inches) long and weigh up to 4.5kg (09.9lb), making it the largest frog ever known.[192] But in more recent studies, animals of this species have been estimated to have grown to at least 23.21NaN1 (snout-vent length), which is around the size a modern African bullfrog can reach.[193] The largest tree frog is the Australasian white-lipped tree frog (Litoria infrafrenata), the females of which can reach a length of 14cm (06inches) from snout to vent and can weigh up to 115g.[194] The family Leptodactylidae, one of the most diverse anuran families, also has some very large members. The largest is the Surinam horned frog (Ceratophrys cornuta), which can reach 20cm (10inches) in length from snout to vent and weigh up to 0.48kg (01.06lb).[195] While not quite as large as Ceratophrys cornuta, Leptodactylus pentadactylus is often heavier; it can reach 18.5cm (07.3inches) long and weigh 0.6kg (01.3lb). The largest dendrobatid is the Colombian golden poison frog (Phyllobates terribilis), which can attain a length of 6cm (02inches) and nearly 28.3g.[196] Most frogs are classified under the suborder Neobatrachia, although nearly 200 species are part of the suborder Mesobatrachia, or ancient frogs. The largest of these are the little-known Brachytarsophrys or Karin Hills frogs, of South Asia, which can grow to a maximum snout-to-vent length of 17cm (07inches) and a maximum weight of 0.54kg (01.19lb).[197]
    Caecilians (Gymnophiona)
  • The largest of the worm-like caecilians is the Colombian Thompson's caecilian (Caecilia thompsoni), which reaches a length of 1.5m (04.9feet), a width of about 4.6cm (01.8inches) and can weigh up to about 1kg (02lb).[1]
    Salamanders (Urodela)
  • Besides the previously mentioned Chinese and South China giant salamanders, the closely related Japanese giant salamander (Andrias japonicus) is also sometimes cited as the largest living amphibian, but salamanders of a greater size than 1.53m (05.02feet) and 36kg (79lb) have never been verified for this species. Another giant of the amphibian world is the North American hellbender (Cryptobranchus alleganiensis), which can measure up to 0.76m (02.49feet). The recently described reticulated siren of the southeastern United States rivals the hellbender in size, although it is more lean in build.[198] The largest of the newts is the Iberian ribbed newt (Pleurodeles waltl), which can grow up to 30cm (10inches) in length.[199]

    Fish

    See main article: List of largest fish.

    Invertebrate chordates

    Tunicates (Tunicata)

    The largest tunicate is Synoicum pulmonaria, found at depths of 20and, and are up to 14 centimetres (6 in) in diameter. It is also present in the northwestern Atlantic Ocean, around the coasts of Greenland and Newfoundland, but is less common here than in the east, and occurs only at depths between 10and.[200]

    Entergonas (Enterogona)
  • The largest entergona is Synoicum pulmonaria it is usually found at depths between about 20and and can grow to over a metre (yard) in length. It is also present in the northwestern Atlantic Ocean, around the coasts of Greenland and Newfoundland, but is less common here than in the east, and occurs only at depths between 10and.
    Pleurogonas (Pleurogona)
  • The largest pleurogona is Pyura pachydermatina .[201] In colour it is off-white or a garish shade of reddish-purple. The stalk is two thirds to three quarters the length of the whole animal which helps distinguish it from certain invasive tunicates not native to New Zealand such as Styela clava and Pyura stolonifera.[202] It is one of the largest species of tunicates and can grow to over a metre (yard) in length.[203]
    Aspiraculates (Aspiraculata)
  • The largest aspiraculate is Oligotrema large and surrounded by six large lobes; the cloacal syphon is small. They live exclusively in deep water and range in size from less than one inch (2 cm) to 2.4 inches (6 cm).

    Thaliacea

    The largest thaliacean, Pyrosoma atlanticum, is cylindrical and can grow up to 60 cm (2 ft) long and 4–6 cm wide. The constituent zooids form a rigid tube, which may be pale pink, yellowish, or bluish. One end of the tube is narrower and is closed, while the other is open and has a strong diaphragm. The outer surface or test is gelatinised and dimpled with backward-pointing, blunt processes. The individual zooids are up to long and have a broad, rounded branchial sac with gill slits. Along the side of the branchial sac runs the endostyle, which produces mucus filters. Water is moved through the gill slits into the centre of the cylinder by cilia pulsating rhythmically. Plankton and other food particles are caught in mucus filters in the processes as the colony is propelled through the water. P. atlanticum is bioluminescent and can generate a brilliant blue-green light when stimulated.[204]

    Doliolida (Doliolida)
  • The largest doliolida is Doliolida [205] The doliolid body is small, typically 1–2 cm long, and barrel-shaped; it features two wide siphons, one at the front and the other at the back end, and eight or nine circular muscle strands reminiscent of barrel bands. Like all tunicates, they are filter feeders. They are free-floating; the same forced flow of water through their bodies with which they gather plankton is used for propulsion - not unlike a tiny ramjet engine. Doliolids are capable of quick movement. They have a complicated lifecycle consisting of sexual and asexual generations. They are nearly exclusively tropical animals, although a few species are found as far north as northern California.
    Salps (Salpida)
  • The largest salp is Cyclosalpa bakeri 15cm (6ins) long. There are openings at the anterior and posterior ends of the cylinder which can be opened or closed as needed. The bodies have seven transverse bands of muscle interspersed by white, translucent patches. A stolon grows from near the endostyle (an elongated glandular structure producing mucus for trapping food particles). The stolon is a ribbon-like organ on which a batch of aggregate forms of the animal are produced by budding. The aggregate is the second, colonial form of the salp and is also gelatinous, transparent and flabby. It takes the shape of a radial whorl of individuals up to about 20cm (4in) in diameter. It is formed of approximately 12 zooids linked side by side in a shape that resembles a crown.[206] [207] are largest thetyses: Thetys vagina Individuals can reach up to 30cm (10inches) long.
    Larvaceans (Larvacea)
  • The largest larvacean is Appendicularia 1cm (00inches) in body length (excluding the tail).

    Cephalochordates (Leptocardii)

    The largest lancelet is the European lancelet (Branchiostoma lanceolatum) "primitive fish". It can grow up to 6 cm (2.5 in) long.[208]

    Invertebrate non-chordates

    Echinoderms (Echinodermata)

    The largest species of echinoderm in terms of bulk is probably the starfish species Thromidia gigas, of the class Asteroidea, which reaches a weight of over 6kg (13lb),[209] but it might be beaten by some giant sea cucumbers such as Thelenota anax. However, at a maximum span of 63cm (25inches), Thromidia gigas is quite a bit shorter than some other echinoderms.[1] The longest echinoderm known is the conspicuous sea cucumber Synapta maculata, with a slender body that can extend up to 3m (10feet). In comparison, the biggest sea star is the brisingid sea star Midgardia xandaros, reaching a span of 1.4m (04.6feet), despite being quite slender.[1] Evasterias echinosoma is another giant echinoderm and can measure up to 1m (03feet) across and weigh 5.1kg (11.2lb).[1]

    Crinoids (Crinoidea)
  • The largest species of crinoid is the unstalked feather-star Heliometra glacialis, reaching a total width of 78cm (31inches) and an individual arm length of 35cm (14inches). A width of 91.4cm (36inches) was claimed for one unstalked feather-star but is not confirmed.[1] The genus Metacrinus has a stalk span of 61cm (24inches) but, due to its bulk and multiple arms, it is heavier than Heliometra.[1] In the past, crinoids grew much larger, and stalk lengths up to 40m (130feet) have been found in the fossil record.[210]
    Sea urchins and allies (Echinoidea)
  • The largest sea urchin is the species Sperosoma giganteum from the deep northwest Pacific Ocean, which can reach a shell width of about 30cm (10inches).[211] Another deep sea species Hygrosoma hoplacantha is only slightly smaller.[211] The largest species found along the North America coast is the Pacific red sea urchin (Mesocentrotus franciscanus) where the shell can reach 19cm (07inches).[212] If the spines enter into count, the biggest species may be a Diadematidae like Diadema setosum, with a test up to 10cm (00inches) only, but its spines can reach up to 30cm (10inches) in length.[213]
    Sea cucumbers (Holothuroidea)
  • The bulkiest species of sea cucumber are Stichopus variegatus and Thelenota anax, weighing several pounds, being about 21cm (08inches) in diameter, and reaching a length of 1m (03feet) when fully extended. Synapta maculata can reach an extended length of 3m (10feet), but is extremely slender (3-5cm) and weigh much less than Stichopodids.[1]
    Brittle stars (Ophiuroidea)
  • The largest known specimen of brittle star is the basket star Astrotoma agassizii. This species can grow to have a span of 1m (03feet).[1] Sometimes, Gorgonocephalus stimpsoni is considered the largest but the maximum this species is can measure 70cm (30inches) and a disk diameter of about 14.3cm (05.6inches). Outside from euryalids, the biggest ophiurid brittle star may be Ophiopsammus maculata (6–7 inches).[214]
    Sea stars (Asteroidea)
  • The heaviest sea star is Thromidia gigas from the Indo-Pacific, which can surpass 6kg (13lb) in weight, but only has a diameter of about 65cm (26inches).[209] [211] Despite its relatively small disk and weight, the long slender arms of Midgardia xandaros from the Gulf of California makes it the sea star with the largest diameter at about 4.5abbr=onNaNabbr=on.[211] Mithrodia clavigera may also become wider than 1m (03feet) in some cases, with stout arms.

    Flatworms (Platyhelminthes)

    Monogenean flatworms (Monogenea)
  • The largest known members of this group of very small parasites are among the genus of capsalids, Listrocephalos, reaching a length of 2cm (01inches).[215]
    Flukes (Trematoda)
  • The largest known species of fluke is Fasciolopsis buski, which most often attacks humans and livestock. One of these flukes can be up to 7.5cm (03inches) long and 2cm (01inches) thick.[216]
    Tapeworms (Cestoda)
  • The largest known species of tapeworm is the whale tapeworm, Polygonoporus giganticus, which can grow to over 30m (100feet).[217] [218]

    Segmented worms (Annelida)

    The largest of the segmented worms (including earthworms, leeches, and polychaetes) is the African giant earthworm (Microchaetus rappi). Although it averages about 1.36m (04.46feet) in length, this huge worm can reach a length of as much as 6.7m (22feet) and can weigh over 1.5kg (03.3lb).[219] Only the giant Gippsland earthworm, Megascolides australis, and a few giant polychaetes, including the notorious Eunice aphroditois, reach nearly comparable sizes, reaching 4and, respectively.[1]

    Ribbon worms (Nemertea)

    The largest nemertean is the bootlace worm, Lineus longissimus. A specimen found washed ashore on a beach in St. Andrews, Scotland in 1864 was recorded at a length of 55m (180feet).[220]

    Mollusks (Mollusca)

    Both the largest mollusks and the largest of all invertebrates (in terms of mass) are the largest squids. The colossal squid (Mesonychoteuthis hamiltoni) is projected to be the largest invertebrate.[221] Current estimates put its maximum size at 12to long and 750kg (1,650lb),[222] based on analysis of smaller specimens. In 2007, authorities in New Zealand announced the capture of the largest known colossal squid specimen. It was initially thought to be 10m (30feet) and 450kg (990lb). It was later measured at 4.2m (13.8feet) long and 495kg (1,091lb) in weight. The mantle was 2.5m (08.2feet) long when measured.[223] [224]

    The giant squid (Architeuthis dux) was previously thought to be the largest squid, and while it is less massive and has a smaller mantle than the colossal squid, it may exceed the colossal squid in overall length including tentacles. One giant squid specimen that washed ashore in 1878 in Newfoundland reportedly measured 16.8m (55.1feet) in total length (from the tip of the mantle to the end of the long tentacles), head and body length 6.1m (20feet), 4.6m (15.1feet) in circumference at the thickest part of mantle, and weighed about 900kg (2,000lb). This specimen is still often cited as the largest invertebrate that has ever been examined.[1] [225] [226] However, no animals approaching this size have been scientifically documented and, according to giant squid expert Steve O'Shea, such lengths were likely achieved by greatly stretching the two tentacles like elastic bands.[227]

    Aplacophorans (Aplacophora)
  • The largest known of these worm-like, shell-less mollusks are represented in the genus Epimenia, which can reach 30cm (10inches) long. Most aplacophorans are less than 5cm (02inches) long.[228]
    Chitons (Polyplacophora)
  • The largest of the chitons is the gumboot chiton, Cryptochiton stelleri, which can reach a length of 33cm (13inches) and weigh over 2kg (04lb).[229]
    Bivalves (Bivalvia)
  • The largest of the bivalve mollusks is the giant clam, Tridacna gigas. Although even larger sizes have been reported for this passive animal, the top verified size was for a specimen from the Great Barrier Reef. This creature weighed 270kg (600lb), had an axial length of 1.14m (03.74feet) and depth of 0.75m (02.46feet).[1] The largest bivalve ever was Platyceramus platinus, a Cretaceous giant that reached an axial length of up to 3 m (nearly 10 ft).[230]
    Gastropods (Gastropoda)
  • The "largest" of this most diverse and successful mollusk class of slugs and snails can be defined in various ways.
  • The living gastropod species that has the largest (longest) shell is Syrinx aruanus with a maximum shell length of 0.91m (02.99feet), a weight of 18kg (40lb) and a width of 96cm (38inches).[231] [232] Another giant species is Melo amphora, which in a 1974 specimen from Western Australia, measured 0.71m (02.33feet) long, had a maximum girth of 0.97m (03.18feet) and weighed 16kg (35lb).[1]
  • The largest shell-less gastropod is the giant black sea hare (Aplysia vaccaria) at 0.99m (03.25feet) in length and almost 14kg (31lb) in weight.
  • The largest of the land snails is the giant African snail (Achatina achatina) at up to 1kg (02lb) and 35cm (14inches) long.
    Cephalopods (Cephalopoda)
  • (See Cephalopod size.) While generally much smaller than the giant Architeuthis and Mesonychoteuthis, the largest of the octopuses, the giant Pacific octopus (Enteroctopus dofleini), can grow to be very large. The largest confirmed weight of a giant octopus is 74kg (163lb),[233] with a 7m (23feet) arm span (with the tentacles fully extended) and a head-to-tentacle-tip length of 3.9m (12.8feet).[234] Specimens have been reported up to 125kg (276lb) but are unverified. A weight of 10 - 50kg is a much more common size.[1]

    Roundworms (Nematoda)

    The largest roundworm, Placentonema gigantissima,[235] is a parasite found in the placentas of sperm whales which can reach up to 9m (30feet) in length.[236]

    Velvet worms (Onychophora)

    The largest velvet worm known is Solórzano's velvet worm (Peripatus solorzanoi). An adult female was recorded to have a body length of 22 cm (approximately 8.7 in).[237]

    Arthropods (Arthropoda)

    The largest arthropod known to have existed is the eurypterid (sea scorpion) Jaekelopterus, reaching up to 2.5m (08.2feet) in body length, followed by the millipede relative Arthropleura at around 2.1m (06.9feet) in length.[238] Among living arthropods, the Japanese spider crab (Macrocheira kaempferi) is the largest in overall size, the record specimen, caught in 1921, had an extended arm span of 3.8m (12.5feet) and weighed about 19kg (42lb).[1] The heaviest is the American lobster (Homarus americanus), the largest verified specimen, caught in 1977 off of Nova Scotia weighed 20kg (40lb) and its body length was 1.1m (03.6feet).[1] The largest land arthropod and the largest land invertebrate is the coconut crab (Birgus latro), up to 40cm (20inches) long and weighing up to 4kg (09lb) on average. Its legs may span 1m (03feet).[1]

    Arachnids (Arachnida)

    Both spiders and scorpions include contenders for the largest arachnids.

    Spiders (Araneae)
  • The largest species of arachnid by length is probably the giant huntsman spider (Heteropoda maxima) of Laos, which in 2008 replaced the Goliath birdeater (Theraphosa blondi) of northern South America as the largest known spider by leg-span.[239] However the most massive arachnids, of comparable dimensions and possibly even greater mass, are the Chaco golden knee (Grammostola pulchripes), and the Brazilian salmon pink (Lasiodora parahybana). The huntsman spider may span up to 29cm (11inches) across the legs, while in the New World tarantulas like Theraphosa can range up to 26cm (10inches).[1] In Grammostola, Theraphosa and Lasiodora, the weight is projected to be up to at least 150g and body length is up to 10cm (00inches).[240]
    Scorpions (Scorpiones)
  • The largest of the scorpions is the species Heterometrus swammerdami of the Indian subcontinent, which have a maximum length of 29.2cm (11.5inches) and weigh around 60g. Another extremely large scorpion is the African emperor scorpion (Pandinus imperator), which can weigh 57g but is not known to exceed a length of 23cm (09inches).[1] However, they were dwarfed by Pulmonoscorpius kirktonensis, a giant extinct species of scorpion from Scotland, at an estimated length of 0.7m (02.3feet),[241] and the aquatic Brontoscorpio, at up to 94cm (37inches) which is only known from a free finger.[242]
    Pseudoscorpions (Pseudoscorpiones)
  • The largest pseudoscorpion is Garypus titanius, from Ascension island, which can be 12mm long.[243]

    Crustaceans (Crustacea)

    The largest crustacean is the Tasmanian giant crab (Pseudocarcinus gigas), with a weight of and a carapace width of up to . It is the only species in the genus Pseudocarcinus.[244] Males reach more than twice the size of females.[245] At a length of up to, Lysiosquillina maculata is the largest mantis shrimp in the world.[246] Tasmanian giant freshwater crayfish (Astacopsis gouldi) in weight and over long have been known in the past, but now, even individuals over are rare.[247] The species is only found in Tasmanian rivers flowing north into the Bass Strait below above sea level, and is listed as an endangered species on the IUCN Red List.[248]

    Branchiopods (Branchiopoda)
  • The largest of these primarily freshwater crustaceans is probably Branchinecta gigas, which can reach a length 10cm (00inches).[249]
    Barnacles and allies (Maxillopoda)
  • The largest species is Pennella balaenopterae, a copepod and ectoparasite specialising in parasitising marine mammals. The maximum size attained is 32 cm (about 13 in).[250] The largest of the barnacles is the giant acorn barnacle, Balanus nubilis, reaching 7cm (03inches) in diameter and 12.7cm (05inches) high.[251]
    Ostracods (Ostracoda)
  • The largest living representative of these small and little-known but numerous crustaceans is the species Gigantocypris australis females of which reaching a maximum length of 3cm (01inches).
    Amphipods, isopods, and allies (Peracarida)
  • The largest species is the giant isopod (Bathynomus pergiganteus), which can reach a length of 45 cm (18 inches) and a weight of 1.7 kg (3.7 lb).[252]
    Remipedes (Remipedia)
  • The largest of these cave-dwelling crustaceans is the species Godzillius robustus, at up to 4.5cm (01.8inches).[253]

    Horseshoe crabs (Xiphosura)

    The four modern horseshoe crabs are of roughly the same sizes, with females measuring up to 60cm (20inches) in length and 5kg (11lb) in weight.[254]

    Sea spiders (Pycnogonida)

    The largest of the sea spiders is the deep-sea species Colossendeis colossea, attaining a leg span of nearly 60cm (20inches).[255]

    Trilobites (Trilobita)

    Some of these extinct marine arthropods exceeded 60cm (20inches) in length. A nearly complete specimen of Isotelus rex from Manitoba attained a length over 70cm (30inches), and an Ogyginus forteyi from Portugal was almost as long. Fragments of trilobites suggest even larger record sizes. An isolated pygidium of Hungioides bohemicus implies that the full animal was 90cm (40inches) long.[256] [257]

    Myriapods (Myriapoda)

    Centipedes (Chilopoda)
  • The biggest of the centipedes is Scolopendra gigantea of the neotropics, reaching a length of 33cm (13inches).[258]
    Millipedes (Diplopoda)
  • Two species of millipede both reach a very large size: Archispirostreptus gigas of East Africa and Scaphistostreptus seychellarum, endemic to the Seychelles islands. Both of these species can slightly exceed a length of 28cm (11inches) and measure over 2cm (01inches) in diameter.[1] The largest ever known was the Arthropleura, a gigantic prehistoric specimen that reached nearly 189cm (74inches).
    Symphylans (Symphyla)
  • The largest known symphylan is Hanseniella magna, originating in Tasmanian caves, which can reach lengths from 25mm up to 30mm.[259]

    Insects (Insecta)

    Insects, a class of Arthropoda, are easily the most numerous class of organisms, with over one million identified species, and probably many undescribed species. The heaviest insect is almost certainly a species of beetle, which incidentally is the most species-rich order of organisms. Although heavyweight giant wetas (Deinacrida heteracantha) are known, the elephant beetles of Central and South America, (Megasoma elephas) and (M. actaeon), the Titan beetle (Titanus giganteus) of the neotropical rainforest or the Goliath beetles, (Goliathus goliatus) and (G. regius), of Africa's rainforest are thought to reach a higher weight.[260] The most frequently crowned are the Goliath beetles, the top known size of which is at least 100g and 11.5cm (04.5inches).[1] The elephant beetles and titan beetle can reach greater lengths than the Goliath, at up to 13.1and, respectively, but this is in part thanks to their rather large horns. The Goliath beetle's wingspan can range up to 25cm (10inches).[1]

    Some moths and butterflies have much larger areas than the heaviest beetles, but weigh a fraction as much.

    The longest insects are the stick insects, see below.

    Representatives of the extinct dragonfly-like order Meganisoptera, such as the Carboniferous Meganeura monyi of what is now France and the Permian Meganeuropsis permiana of what is now North America, are the largest insect species known to have existed. These creatures had a wingspan of some 75cm (30inches) and a mass of over 1lb, making them about the size of a crow.[1]

    Cockroaches and termites (Blattodea)
  • The largest cockroach by body mass is the Australian giant burrowing cockroach (Macropanesthia rhinoceros), also known as the rhinoceros cockroach. This species can attain a length of 8.3cm (03.3inches) and a weight of 36g. It does not have wings.[261] The Brazilian giant cockroach (Blaberus giganteus) of the neotropics reaches greater sizes of up to 10 cm in length and 15 cm in wingspan, although it is not as massive and heavy as the burrowing species. The termites, traditionally classified in their own order (Isoptera), have recently been re-considered to belong in Blattodea. The largest of the termites is the African species Macrotermes bellicosus. The queen of this species can attain a length of 14cm (06inches) and breadth of 5.5cm (02.2inches) across the abdomen; other adults, on the other hand, are about a third of the size.[1]
    Beetles (Coleoptera)
  • The beetles are the largest order of organisms on earth, with about 400,000 species so far identified. The most massive species are the Goliathus, Megasoma and Titanus beetles already mentioned. Another fairly large species is the Hercules beetle (Dynastes hercules) of the neotropic rainforest with a maximum overall length of at least 19cm (07inches) including the extremely long pronotal horn. The weight in this species does not exceed 16.5g.[1] The longest overall beetle is a species of longhorn beetle, Batocera wallacei, from New Guinea, which can attain a length of 26.6cm (10.5inches), about 19cm (07inches) of which is comprised by the long antennae.[1]
    Earwigs (Dermaptera)
  • Since 1798, the largest of the earwigs has been the Saint Helena giant earwig (Labidura herculeana), endemic to the island of its name, measuring up to 8cm (03inches) in length.[262] As of 2014, with the declaring of the organism extinct by the IUCN,[263] this may no longer be the case, although some believe a small number individuals are still extant.[264]
    True flies (Diptera)
  • The largest species of this order, which includes the common housefly, is the neotropical species Gauromydas heros, which can reach a length of 6frac=8NaNfrac=8 and a wingspan of 10cm (00inches).[1] Species of crane fly, the largest of which is Holorusia brobdignagius, can attain a length of 23cm (09inches) but are extremely slender and much lighter in weight than Gauromydas.
    Mayflies (Ephemeroptera)
  • The largest mayflies are members of the genus Proboscidoplocia from Madagascar. These insects can reach a length of 7cm (03inches).[265]
    True bugs (Hemiptera)
  • The largest species of this diverse order is usually listed as the giant water bug in the genus Lethocerus, with L. maximus from the Neotropics being the absolutely largest.[266] They can surpass 12cm (05inches) in length,[267] with some suggesting that the maximum size is 15cm (06inches).[268] It is more slender and less heavy than most other insects of this size (principally the huge beetles). The largest cicada is Megapomponia imperatoria, which has a head-body length of about 7cm (03inches) and a wingspan of 18-.[269] [270] The cicadas of the genus Tacua can also grow to comparably large sizes. The largest type of aphid is the giant oak aphid (Stomaphis quercus), which can reach an overall length of 2cm (01inches).[271] The biggest species of leafhopper is Ledromorpha planirostris, which can reach a length of 2.8cm (01.1inches).[272]
    Ants and allies (Hymenoptera)
  • The largest of the ants, and the heaviest species of the order, are the females of the African Dorylus helvolus, reaching a length of 5.1cm (02inches) and a weight of 8.5g.[1] The ant that averages the largest for the mean size within the whole colony is a ponerine ant, Dinoponera gigantea, from South America, averaging up to 3.3cm (01.3inches) from the mandibles to the end of abdomen.[1] Workers of the bulldog ant (Myrmecia brevinoda) of Australia are up to 3.7cm (01.5inches) in total length, although much of this is from their extremely large mandibles.[1] The largest of the bee species, also in the order Hymenoptera, is Megachile pluto of Indonesia, the females of which can be 3.8cm (01.5inches) long, with a 6.3cm (02.5inches) wingspan. Nearly as large, the carpenter bees can range up to 2.53cm (01inches).[1] The largest wasp is probably the so-called tarantula hawk species Pepsis pulszkyi of South America, at up to 6.8cm (02.7inches) long and 11.6cm (04.6inches) wingspan, although many other Pepsis approach a similar size. The giant scarab-hunting wasp Megascolia procer may rival the largest tarantula hawks in weight and wingspan, though its body is not as long.[1]
    Moths and allies (Lepidoptera)
  • The Hercules moth (Coscinocera hercules), in the family Saturniidae, is endemic to New Guinea and northern Australia, and its wings have the largest documented surface area (300 square centimeters) of any living insect,[273] [274] and a maximum wingspan which is confirmed to 28cm (11inches) while unconfirmed specimens have spanned up to 35.5cm (14inches). The largest species overall is often claimed to be either the Queen Alexandra's birdwing (Ornithoptera alexandrae), a butterfly from Papua New Guinea, or the Atlas moth (Attacus atlas), a moth from Southeast Asia. Both of these species can reach a length of 8cm (03inches), a wingspan of 28cm (11inches) and a weight of 12g. One Atlas moth allegedly had a wingspan of 30cm (10inches) but this measurement was not verified.[1] The larvae in the previous species can weigh up to 58and, respectively. The white witch (Thysania agrippina) of Central and South America, has the largest recorded wingspan of the order, and indeed of any living insect. The verified record-sized Thysania spanned 30.8cm (12.1inches) across the wings, although specimens have been reported to 36cm (14inches).[1] The heaviest mature moths have been cited in the giant carpenter moth (Xyleutes boisduvali) of Australia, which has weighed up to 20g although the species does not surpass 25.5cm (10inches) in wingspan.[1]
    Mantises (Mantodea)
  • The largest species of this order is Toxodera denticulata from Java, which has been measured up to 20cm (10inches) in overall length.[275] However, an undescribed species from the Cameroon jungle is allegedly much larger than any other mantis and may rival the larger stick insects for the longest living insect.[276] Among widespread mantis species, the largest is the Chinese mantis (Tenodera aridifolia). The females of this species can attain a length of up to 10.6cm (04.2inches).
    Scorpionflies (Mecoptera)
  • The largest scorpionfly, the common scorpionfly (Panorpa communis), can reach a body length of about .[277]
    Alderflies and allies (Megaloptera)
  • This relatively small insect order includes some rather large species, many of which are noticeable for their elongated, imposing mandibles. The dobsonflies reach the greatest sizes of the order and can range up to 12.5cm (04.9inches) in length.[278]
    Net-winged insects (Neuroptera)
  • These flying insects reach their largest size in Palparellus voeltzkowi, which can have a wingspan over 16cm (06inches).[279] The largest lacewing is the "blue eyes lacewing" (Nymphes myrmeleonides) of Australia, which can measure up to 4cm (02inches) in length and span 11cm (04inches) across the wings.[280] Some forms of this ancient order could grow extremely large during the Jurassic period and may have ranked among the largest insects ever.[281] Found in the Early Cretaceous sedimentary rocks, Makarkinia adamsi had wings nearly 140- in length.[282]
    Dragonflies and damselflies (Odonata)
  • The largest species of Odonata is the damselfly Megaloprepus caerulatus of the neotropics, attaining a size of as much as 19cm (07inches) across the wings and a body length of over 12cm (05inches).[1] The largest species of dragonfly is Tetracanthagyna plagiata of Southeast Asia, spanning up to 17.6cm (06.9inches) and measuring up to 11.8cm (04.6inches) long; while bulkier and heavier than Megaloprepus (at up to 7g), it is smaller in its linear dimensions.[1]
    Grasshoppers and allies (Orthoptera)
  • The largest of this widespread, varied complex of insects are the giant wetas of New Zealand, which is now split among 12 species. The largest of these is the Little Barrier Island giant weta (Deinacrida heteracantha), the largest specimen was weighed at 71.3g, one of the largest insects weights ever known. These heavyweight insects can be over 9cm (04inches) long.[1] The largest grasshopper species is often considered to be the Australian giant grasshopper (Valanga irregularis), which ranges up to 9cm (04inches) in length.[283] The American eastern lubber grasshopper (Romalea guttata) can allegedly range up to 10cm (00inches) in length.[284] However, the greatest grasshopper sizes known, to 12cm (05inches), have been cited in the South American giant grasshopper (Tropidacris violaceus). The longest members of this order (although much lighter than the giant wetas) is the katydid Macrolyristes corporalis of Southeast Asia which can range up to 21.5cm (08.5inches) with its long legs extended and can have a wingspan of 20cm (10inches).[285]
    Stick insects (Phasmatodea)
  • The longest known stick insects are also the longest known insects, notably species in the tribe Pharnaciini, but they are generally relatively lightweight because of their slender shape. The longest is an unnamed species of Phryganistria discovered in China in 2016, where a specimen held at the Insect Museum of West China in Chengdu has a total length of .[286] The second-longest species is the Australian Ctenomorpha gargantua, females of which have been measured at over in total length.[287] Other very large species, formerly believed to be longest but now considered third longest is Sadyattes chani; a specimen held in the Natural History Museum in London has a total length of .[288] [289] These measurements are, however, with the front legs fully extended; it has a body length measuring .[290] Another very large species is Phobaeticus kirbyi where the total length (including extended legs) is up to and the body alone up to .[291] Another of the longest insect in terms of total length is Phobaeticus serratipes of Malaysia and Singapore, measuring up to 55.5cm (21.9inches).[292] Another extremely long stick insect is Pharnacia maxima, which measured 51cm (20inches) with its legs extended.[1] The spiny stick insect (Heteropteryx dilatata) of Malaysia does not reach the extreme lengths of its cousins, the body reaching up to 16cm (06inches) long, but it is much bulkier. The largest Heteropteryx weighed about 65g and was 3.5cm (01.4inches) wide across the thickest part of the body.[1]
    Lice (Phthiraptera)
  • These insects, which live parasitically on other animals, are as a rule quite small. The largest known species is the hog louse, Haematopinus suis, a sucking louse that lives on large livestock like pigs and cattle. It can range up to 6mm in length.[293]
    Stoneflies (Plecoptera)
  • The largest species of stonefly is Pteronarcys californica of western North America, a species favored by fishermen as lures. This species can attain a length of 5cm (02inches) and a wingspan of over 9.5cm (03.7inches).[294]
    Booklice (Psocoptera)
  • The largest of this order of very small insects are the barklice of the genus Psocus, the top size of which is about 1 cm.[295]
    Fleas (Siphonaptera)
  • The largest species of flea is Hystrichopsylla schefferi. This parasite is known exclusively from the fur of the mountain beaver (Aplodontia rufa) and can reach a length of 1.2cm (00.5inches).[1]
    Silverfishes and allies (Thysanura)
  • These ancient flightless insects, some of which feed on human household objects, can range up to 4.3cm (01.7inches) in length. A 350 million year old form was known to grow quite large, at up to 6cm (02inches).
    Thrips (Thysanoptera)
  • Members of the genus Phasmothrips are the largest kinds of thrips. The maximum size these species attain is approximately 1.3cm (00.5inches) in length.[296]
    Caddisflies (Trichoptera)
  • The largest of the small, moth-like caddisflies is Eubasilissa maclachlani. This species can range up to 7cm (03inches) across the wings.[297]
    Angel insects (Zoraptera)
  • The largest angel insect species, Hubbard's angel insect (Zorotypus hubbardi), grows up to 32NaN2 in length.[298] [299]

    Cnidarians (Cnidaria)

    The lion's mane jellyfish (Cyanea capillata) is the largest cnidarian species, of the class Scyphozoa. The largest known specimen of this giant, found washed up on the shore of Massachusetts Bay in 1870,[300] [301] had a bell diameter of 2.5m (08.2feet), a weight of 150kg (330lb). The tentacles of this specimens were as long as 37m (121feet) and were projected to have a tentacular spread of about 75m (246feet) making it one of the longest extant animals.[1]

    Corals and sea anemones (Anthozoa)
  • The largest individual species are the sea-anemones of the genus Discoma, which can attain a mouth disc diameter of 60cm (20inches).[302] Longer, but much less massive overall, are the anemones of the genus Ceriantharia, at up to 2m (07feet) tall.[303] Communities of coral can be truly massive, a single colony of the genus Porites can be over 10m (30feet), but the actual individual organisms are quite small.
    Hydrozoans (Hydrozoa)
  • The colonial siphonophore Praya dubia can attain lengths of 40–.[304] The Portuguese man o' war's (Physalia physalis) tentacles can attain a length of up to 50m (160feet).[305] On 6 April 2020 the Schmidt Ocean Institute announced the discovery of a giant Apolemia siphonophore in submarine canyons near Ningaloo Coast, measuring 15 m (49 ft) diameter with a ring approximately 47 m (154 ft) long, claiming it was possibly the largest siphonophore ever recorded.[306] [307]

    Sponges (Porifera)

    The largest known species of sea sponge is the giant barrel sponge, Xestospongia muta. These massively built sponges can reach 8abbr=onNaNabbr=on in height and can be about the same thickness at the thickest part of the "body".[308] Some of these creatures have been estimated to be over 2,400 years of age.[309]

    Calcareous sponges (Calcarea)
  • The largest known of these small, inconspicuous sponges is probably the species Pericharax heteroraphis, attaining a height of 30cm (10inches). Most calcareous sponges do not exceed 10cm (00inches) tall.
    Hexactinellid sponges (Hexactinellida)
  • A relatively common species, Rhabdocalyptus dawsoni, can reach a height of 1m (03feet) once they are of a very old age.[310] This is the maximum size recorded for a hexactinellid sponge.

    See also

    Notes and References

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    304. http://www.lifesci.ucsb.edu/~biolum/organism/pictures/praya.html Praya picture
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