Glossary of leaf morphology explained

The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular or irregular, may be smooth or bearing hair, bristles or spines. For more terms describing other aspects of leaves besides their overall morphology see the leaf article.

The terms listed here all are supported by technical and professional usage, but they cannot be represented as mandatory or undebatable; readers must use their judgement. Authors often use terms arbitrarily, or coin them to taste, possibly in ignorance of established terms, and it is not always clear whether because of ignorance, or personal preference, or because usages change with time or context, or because of variation between specimens, even specimens from the same plant. For example, whether to call leaves on the same tree "acuminate", "lanceolate", or "linear" could depend on individual judgement, or which part of the tree one collected them from. The same cautions might apply to "caudate", "cuspidate", and "mucronate", or to "crenate", "dentate", and "serrate".

Another problem is to establish definitions that meet all cases or satisfy all authorities and readers. For example, it seems altogether reasonable to define a mucro as "a small sharp point as a continuation of the midrib", but it may not be clear how small is small enough, how sharp is sharp enough, how hard the point must be, and what to call the point when one cannot tell whether the leaf has a midrib at all. Various authors or field workers might come to incompatible conclusions, or might try to compromise by qualifying terms so vaguely that a description of a particular plant practically loses its value.

Use of these terms is not restricted to leaves, but may be applied to morphology of other parts of plants, e.g. bracts, bracteoles, stipules, sepals, petals, carpels or scales. Some of these terms are also used for similar-looking anatomical features on animals.

Leaf structure

Leaves of most plants include a flat structure called the blade or lamina, but not all leaves are flat, some are cylindrical. Leaves may be simple, with a single leaf blade, or compound, with several leaflets. In flowering plants, as well as the blade of the leaf, there may be a petiole and stipules; compound leaves may have a rachis supporting the leaflets. Leaf structure is described by several terms that include:

ImageTermLatinDescription
Having two leaflets[1]
Having two leaflets, each leaflet being bifoliolate
Latin: bipinnatus The leaflets are themselves pinnately-compound; twice pinnate
With three components, each with three leaflets
With an odd number of leaflets, pinnate with a terminal leaflet (the opposite of paripinnate)
Pinnate with an even number of leaflets, lacking a terminal leaflet (the opposite of imparipinnate)
even-pinnate
palmatusConsisting of leaflets all radiating from one point
pinnatusHaving two rows of leaflets on opposite sides of a central axis, see imparipinnate and paripinnate
Leaf blade in one continuous section, without leaflets (not compound)
Latin: ternatus With three leaflets
Latin: trifoliatus
Latin: trifoliolatus
Latin: tripinnatus Pinnately compound in which each leaflet is itself bipinnate

Leaf and leaflet shapes

Being one of the more visible features, leaf shape is commonly used for plant identification. Similar terms are used for other plant parts, such as petals, tepals, and bracts.

ImageTermLatinRefers principally toDescription
Latin: acicularis whole leaf Slender and pointed, needle-like
Latin: acuminatus leaf tip Tapering to a long point in a concave manner
leaf tip or base Pointed, having a short sharp apex angled less than 90°
leaf tip Tapering and ending in a short, slender point
Latin: aristatus leaf tip Ending in a stiff, bristle-like point
whole leaf With the blade shape different on each side of the midrib
Latin: attenuatusleaf base Having leaf tissue taper down the petiole to a narrow base and always having some leaf material on each side of the petiole
Latin: auriculatusleaf base Having ear-shaped appendages reaching beyond the attachment to the petiole or stem (in case of a seated leaf)
Latin: caudatus leaf tip Tailed at the apex
, leaf tip Having a rachis that extends beyond the leaf blade or leaflets into a long whip-like extension or cirrus (common in climbing palms); antonym: ecirrate
, Latin: cordatus whole leaf or baseHeart-shaped, with the petiole or stem attached to the notch
Latin: cuneatus leaf base Triangular, wedge-shaped, stem attaches to point
whole leaf Narrowly triangular, widest on the opposite end from the stem, with the corners at that end rounded
Latin: cuspidatus leaf tip With a sharp, elongated, rigid tip; tipped with a cusp
, Latin: deltoideus whole leaf Shaped like the Greek letter delta; triangular with stem attached to side
Latin: digitatus whole leaf A palmately compound leaf with leaflets, similar to palmate[2]
leaf tip Without a cirrus; antonym: cirrate
Latin: ellipticus whole leaf Shaped like an ellipse (widest at mid-blade and with similar convex tapering towards apex and base), with a short or no point
Latin: emarginatusleaf tip Slightly indented at the tip
Latin: ensiformis whole leaf Shaped like a sword; long and narrow with a sharp pointed tip
Latin: falcatus whole leaf Sickle-shaped
Latin: fenestratus leaf surface features Large openings through the leaf; see perforate; sometimes used to describe leaf epidermal windows
Latin: filiformis whole leaf Thread- or filament-shaped
Latin: flabellatus whole leaf Semi-circular or fan-like
Latin: hastatus whole leaf or base Spear-shaped: pointed, with barbs, shaped like a spear point, with flaring pointed lobes at the base
lacinatuswhole leaf Very deeply lobed with the lobes being very drawn out and often making the leaf look somewhat like a branch or a pitchfork
3-D shape Flat (like most leaves)
Latin: lanceolatus whole leaf Long, wider in the middle, shaped like a lance tip
Latin: linearis whole leaf Long and very narrow like a blade of grass
Latin: lobatus whole leaf Being divided by clefts; may be pinnately lobed or palmately lobed
Latin: loratus whole leaf Having the form of a thong or strap
Latin: lyratus whole leaf Shaped like a lyre, pinnately lobed leaf with an enlarged terminal lobe and smaller lateral lobes. See also List of lyrate plants.
mucronatusleaf tip Ending abruptly in a small sharp point as a continuation of the midrib[3]
Latin: multi + Latin: findere whole leaf Cleft into many parts or lobes
Latin: obcordatus whole leaf Heart-shaped, stem attaches at the tapering end
Latin: oblanceolatus whole leaf Much longer than wide and with the widest portion near the tip; reversed lanceolate
leaf base Asymmetrical leaf base, with one side lower than the other
Latin: oblongus whole leaf Having an elongated form with slightly parallel sides; roughly rectangular
Latin: obovatus whole leaf Teardrop-shaped, stem attaches to the tapering end; reversed ovate
whole leaf Reversed trullate; the longer sides meet at the base rather than the apex.
Latin: obtusus leaf tip or base Blunt, forming an angle > 90°
Latin: orbicularis whole leaf Circular
Latin: ovatus whole leaf Egg-shaped, with a tapering point and the widest portion near the petiole
Latin: palmatus whole leaf Palm-shaped, i.e. with lobes or leaflets stemming from the leaf base[4]
Latin: palmatus whole leaf Lobes spread radially from a point[5]
Latin: palma + Latin: findere whole leaf Palm-shaped, having lobes with incisions that extend less than halfway toward the petiole
Latin: palma + Latin: partiri whole leaf Having palmate lobes with incisions that extend over halfway toward the petiole
Latin: palma + Latin: secare whole leaf Having palmate lobes with incisions that extend almost up, but not quite to the petiole.
panduratuswhole leaf Fiddle-shaped; obovate with a constriction near the middle.
Latin: pedatus whole leaf Palmate, with cleft lobes[6]
Latin: peltatus stem attachment A round leaf where the petiole attaches near the center, e.g. a lotus leaf
Latin: perfoliatus stem attachment With the leaf blade surrounding the stem such that the stem appears to pass through the leaf
Latin: perforatus leaf surface featuresMany holes, or perforations, on leaf surface. Compare with fenestrate.
Latin: pinna + Latin: lobus whole leaf Having lobes pinnately arranged on the central axis
Latin: pinna + Latin: findere whole leaf Having lobes with incisions that extend less than halfway to the midrib
Latin: pinnatus + Latin: partiri whole leaf Having lobes with incisions that extend more than halfway to the midrib
Latin: pinnatus + Latin: sectus whole leaf Having lobes with incisions that extend almost to, or up to, the midrib
Latin: plicatus 3-D shape Folded into pleats, usually lengthwise, serving the function of stiffening a large leaf
Latin: reniformis whole leaf Shaped like a kidney, with an inward curve on one side
leaf tip With a shallow notch in a round apex
, Latin: rhomboidalis whole leaf Diamond-shaped
Latin: rotundifolius leaf tip or base Circular, no distinct point
semiterete 3-D shape Rounded on one side and flat on the other
Latin: sinuatus3-D shape Circularly-lobed leaves
Latin: sagittatus whole leaf Arrowhead-shaped with the lower lobes folded, or curled downward
Latin: spathulatus whole leaf Spoon-shaped; having a broad flat end which tapers to the base
-shaped Latin: hastatuswhole leaf See hastate.
Latin: subobtusus leaf tip or base Somewhat blunted; neither blunt nor sharp
Latin: subulatus leaf tip Awl-shaped with a tapering point
3-D shape Cylindrical with a circular or distorted circular cross-section and a single surface wrapping around it with no grooves or ridges. Subterete means the leaves are not completely terete, as seen in various lichens and succulents.
whole leaf Shaped like a masonry trowel
Latin: truncatus leaf tip or base With a squared-off end
Latin: undulatus 3-D shape Wave-like
Latin: unifoliatus compound leaves With a single leaflet; it is distinct from a simple leaf by the presence of two abscission layers and often by petiolules and stipels.

Edge

Leaf margins (edges) are frequently used in visual plant identification because they are usually consistent within a species or group of species, and are an easy characteristic to observe. Edge and margin are interchangeable in the sense that they both refer to the outside perimeter of a leaf.

ImageTermLatinDescription
100pxentireForma
integra
Even; with a smooth margin; without toothing
100pxciliateciliatusFringed with hairs
100pxcrenatecrenatusWavy-toothed; dentate with rounded teeth
crenulate crenulatus Finely crenate
crisped crispus Curly
100pxdentatedentatusToothed; may be coarsely dentate, having large teeth

or glandular dentate, having teeth which bear glands

100pxdenticulatedenticulatusFinely toothed
100pxdoubly serrateduplicato-dentatusEach tooth bearing smaller teeth
100pxserrateserratusSaw-toothed; with asymmetrical teeth pointing forward
100pxserrulateserrulatusFinely serrate
100pxsinuatesinuosusWith deep, wave-like indentations; coarsely crenate
100pxlobatelobatusIndented, with the indentations not reaching the center
100pxundulateundulatusWith a wavy edge, shallower than sinuate
100pxspiny or pungentspiculatusWith stiff, sharp points such as thistles

Leaf folding

Leaves may also be folded, sculpted or rolled in various ways. If the leaves are initially folded in the bud, but later unrolls it is called vernation, ptyxis is the folding of an individual leaf in a bud.

ImageTermLatinDescription
or keeled carinatus With a longitudinal ridge, keel-shaped
Folded upwards, with the surfaces close to parallel
Forming a hood, margins and tip curved downward
Rolled upwards (towards the adaxial surface)
plicatus With parallel folds
Folded downwards, with the surfaces close to parallel
Rolled downwards (towards the abaxial surface)
Opposing left and right halves of lamina folded along longitudinal axis, with one half rolled completely within the other

Latin descriptions

The Latin word for 'leaf', Latin: folium, is neuter. In descriptions of a single leaf, the neuter singular ending of the adjective is used, e.g. Latin: folium lanceolatum 'lanceolate leaf', Latin: folium lineare 'linear leaf'. In descriptions of multiple leaves, the neuter plural is used, e.g. Latin: folia linearia 'linear leaves'. Descriptions commonly refer to the plant using the ablative singular or plural, e.g. Latin: foliis ovatis 'with ovate leaves'.

See also

Bibliography

External links

Notes and References

  1. Book: Vascular Plant Systematics . Radford . A.E. . W.C. . Dickison . J.R. . Massey . C.R. . Bell . 1976 . Harper and Row, New York . Phytography - Morphological Evidence . http://www.ibiblio.org/botnet/glossary/.
  2. Index of Garden Plants, Mark Griffiths, Timber Press, 1992
  3. http://www.answers.com/main/ntquery?s=Mucronate&gwp=13 Mucronate
  4. Web site: palmate (adj. palmately) . 2006 . GardenWeb Glossary of Botanical Terms . iVillage GardenWeb . 19 October 2008 . https://web.archive.org/web/20090213124631/http://glossary.gardenweb.com:80/glossary/palmate.html . 13 February 2009 . dead .
  5. Web site: Nelson . Randal C. . Leaf description glossary . 2009 . 2012 . University of Rochester . 2 November 2017 . https://web.archive.org/web/20200801032721/https://www.cs.rochester.edu/~nelson/wildflowers/glossaries/leaves/index.html . 1 August 2020 . dead.
  6. Web site: Pedate leaf . 24 February 2014.