Cercopidae Explained
Cercopidae are the largest family of Cercopoidea, a xylem-feeding insect group, commonly called froghoppers.[1] They belong to the hemipteran suborder Auchenorrhyncha. A 2023 phylogenetic study of the family suggested the elevation of subfamily Ischnorhininae to full family status as Ischnorhinidae, leaving a monophyletic Cercopinae.[2]
Taxonomy
Based on a 2023 molecular phylogenetic analysis, the Cercopidae senso stricto was divided into two monophyletic subfamilies, Cercopinae and Cosmoscartinae, each with a number of tribes.[2]
- Subfamily CercopinaeTribe Bandusiini
Tribe Callitetixini
- Abidama
- Callitettix
- Caloscarta
- Moultoniella
- Rhinastria
- Tadascarta
Tribe Cercopini
Tribe Haematoscartini
- Clypeocarta
- Haematoscarta
- Lieftinckana
- Straelenia
Tribe Hemiaufidini
Tribe Locrisini
- Anyllis
- Dulitana
- Locris
- Nesaphrogeneia
- Paralocris
Tribe Rhinaulacini
- Amberana
- Asiacercopis
- Aufidus
- Aufiterna
- Augustohahnia
- Baibarana
- Bourgoinrana
- Circulocercopis
- Colsa
- Eoscarta
- Euryaulax
- Hemibandusia
- Hemicercopis
- Hemiliterna
- Heteroliterna
- Jacobsoniella
- Jeanneliensa
- Kanozata
- Lehina
- Leptoliterna
- Liorhinella
- Literna
- Machadoa
- Mioscarta [3]
- Nesaulax
- Neocercopis
- Paracercopis
- Paracercopoides
- Paraliterna
- Paramioscarta
- Parapisidice
- Petyllis
- Poeciloterpa [4]
- Rhinaulax
- Sounama
- Synavea
- Telogmometopius
- Thoodzata
- Trigonoschema [3]
- Villiersana
- Wawi
Tribe incertae sedis
- Alluaudensia
- Ambonga
- Blötea
- Cercopicesa
- Euglobiceps
- Euryliterna
- Janssensia
- Lamprochlamys
- Leptynis
- Orthorhinella
- Paramonecphora
- Phlebarcys
- Pisianax
- Pogonorhinella
- Pseudocercopis
- Subfamily CosmoscartinaeTribe Considiini
Tribe Cosmoscartini
- Cosmoscarta
- Ectemnonotops
- Ectemnonotum
- Eubakeriella
- Gynopygocarta
- Gynopygolax
- Helioscarta
- Homalostethus
- Kanoscarta
- Kotozata
- Okiscarta
- Oxymegaspis
- Oxypeltarion
- Platynotoscarta
Tribe Phymatostethini
Tribe Suracartini
- Anoplosnastus
- Ectemnocarta
- Leptataspis
- Megastethodon
- Opistharsostethus
- Serapita
- Simeliria
- Suracarta
Tribe Trichoscartini
- Hemitrichoscarta
- Neoporpacella
- Paratrichoscarta
- Pisidice
- Porpacella
- Trichoscarta
Tribe incertae sedis
Invalid genera
Crispolon Jr et al. (2023) considered certain described genera as invalid without discussion and did not include them in the family as prescribed by their phylogeny.[2]
- Delassor
- Funkhouseria
- Guarania
- Hemieoscarta
- Kuscarta
- Luederwaldtia
- Pachypterinella
- Pseudeoscarta
- Stenaulophrys
- Vigilantius
Fossil genera
- †Allocercopis
- †Berro
- †Cercopites
- †Cicadellites
- †Dawsonites
- †Eocercopidium
- †Lithecphora
- †Locrites
- †Megacercopis
- †Nangamostethos [5]
- †Nisocercopis
- †Palecphora
- †Parafitopteryx
- †Prinecphora
- †Stenecphora
- †Stenolocris
Taxa removed from Cercopidae
Crispolon Jr. et al.s 2023 molocular phylogenetic analysis recommended removing the following genera from Cercopidae and transferring them to Aphrophoridae:[2]
- Hemitriecphora
- Microsargane
Additionally they recommended the elevation of the subfamily Ischnorhininae, endemic to the Americas, to full family status as Ischnorhinidae. This was based on the closer phylogenetic ties to Machaerotidae then to subfamily Cercopinae found exclusively in Europe, Africa, and Asia, Australasia:[2]
- Family IschnorhinidaeTribe Ischnorhinini
- Baetkia
- Choconta
- Homalogrypota
- Huaina
- Iphirhina
- Ischnorhina
- Laccogrypota
- Neolaccogrypota
- Neosphenorhina
- Schistogonia
- Sphenoclypeana
- Tiodus
- Typeschata
Tribe Neaenini
- Helioptera
- Hemitomaspis
- Marcion
- Menytes
- Neaenus
- Simorhina
- Tomaspisinella
- Zuata
Tribe Tomaspini
- Aeneolamia
- Aracamunia
- Bradypteroscarta
- Carachata
- Carpentiera
- Catrimania
- Chinana
- Deois
- Deoisella
- Ferorhinella
- Gervasiella
- Hemiplagiophleboptena
- Hyalotomaspis
- Hyboscarta
- Isozulia
- Kanaima
- Korobona
- Lujana
- Mahanarva
- Makonaima
- Maxantonia
- Monecphora
- Neomonecphora
- Notozulia
- Ocoaxo
- Olcotomaspis
- Orodamnis
- Pachacanthocnemis
- Panabrus
- Plagiophleboptena
- Prosapia
- Sinopia
- Sphenorhina
- Tapaiuna
- Tomaspis
- Tomaspisina
- Tropidorhinella
- Tunaima
- Urubaxia
- Vorago
- Zulia
Notes and References
- Carvalho C, Webb M . Cercopid Spittle Bugs of the New World (Hemiptera, Auchenorrhyncha, Cercopidae) . Pensoft Series Faunistica . 49 . 280pp . 2005 . 954-642-246-0.
- Crispolon Jr . E. S. . Soulier‐Perkins . A. . Guilbert . E. . 2023 . Molecular phylogeny of Cercopidae (Hemiptera, Cercopoidea). . Zoologica Scripta . 52 . 5 . 494–516 . 10.1111/zsc.12597. free .
- Crispolon Jr . E. S. . Guilbert . E. . Yap . S. A. . Soulier‐Perkins . A. . 2021 . New genus and new species of spittlebugs (Hemiptera: Cercopidae) from the Philippines . European Journal of Taxonomy . 778 . 90–135 . 10.5852/ejt.2021.778.1571. free .
- Crispolon Jr . E. S. . Yap . S. A. . Soulier‐Perkins . A. . 2019 . Revision of the endemic Philippine Poeciloterpa Stål (Hemiptera: Cercopidae) with description of four new species. . Zootaxa . 4608 . 2 . 291–328 . 10.11646/zootaxa.4608.2.6.
- Xu . X.-T. . Szwedo . J. . Huang . D.-Y. . Deng . W.-Y.-D. . Obroślak . M. . Wu . F.-X. . Su . T. . 2022 . A New Genus of Spittlebugs (Hemiptera, Cercopidae) from the Eocene of Central Tibetan Plateau . Insects . 13 . 770 . 770 . 10.3390/insects13090770. 36135471 . 9503688 . free .