Cimolodonta Explained

Cimolodonta is a clade of multituberculate mammals that lived from the Cretaceous to the Eocene. They probably lived something of a rodent-like existence until their ecological niche was assumed by true rodents. The more basal multituberculates are found in a different suborder, "Plagiaulacida", a paraphyletic group containing all non cimolodontan multituberculates.

Cimolodonta is apparently a natural (monophyletic) suborder. Remains have been identified from across the Northern Hemisphere. They first appeared during the Aptian, and completely replaced the more primitive plagiaulacidans by the early Late Cretaceous.^[1] The taxon is recognized as the informal Paracimexomys group and the superfamilies Djadochtatherioidea, Taeniolabidoidea, and Ptilodontoidea. Additionally, and of uncertain affinities, are the families Cimolomyidae, Boffiidae, Eucosmodontidae, Kogaionidae, Microcosmodontidae and the two genera Uzbekbaatar and Viridomys. More precise placement of these types awaits further discoveries and analysis.

Taxonomy

Suborder †Cimolodonta McKenna, 1975

• Superfamily Incertae sedis
  • Family Incertae sedis
    • Subfamily Incertae sedis
      • Genus ?Ameribaatar Eaton & Cifelli, 2001
      • Genus ?Bryceomys Eaton, 1995
      • Genus Cedaromys Eaton & Cifelli, 2001
      • Genus ?Dakotamys Eaton, 1995; (Early Cretaceous, Central North America)
      • Genus ?Fractinus Higgins, 2003
      • Genus Halodon Marsh, 1889
      • Genus Ptilodus (Marsh, 1889) Gidley, 1909
      • Genus ?Uzbekbaatar Kielan-Jaworowska & Nesov, 1992
      • Genus ?Barbatodon Rãdulescu & Samson, 1986
• "Paracimexomys group" Archibald, 1982
  • Genus Paracimexomys Archibald, 1982
  • Genus ?Cimexomys Fox, 1971
  • Genus Cimexomys Sloan & Van Valen, 1965 sensu stricto?
• Family Boffiidae Hahn & Hahn, 1983
  • Genus Boffius Vianey-Liaud, 1979
• Family Cimolomyidae Marsh, 1889 sensu Kielan-Jaworowska & Hurum, 2001
  • Genus Essonodon

Simpson, 1927

• • Genus Buginbaatar Kielan-Jaworowska & Sochava, 1969
  • Genus Meniscoessus Cope, 1882 (syn. Dipriodon Marsh, 1889, Tripriodon Marsh, 1889, Selenacodon Marsh, 1889, Halodon Marsh, 1889, Oracodon Marsh, 1889)
  • Genus Cimolomys Marsh, 1889 (syn. ?Allacodon Marsh, 1889; Meniscoessus; Ptilodus; Selenacodon Marsh, 1889)
• Superfamily Ptilodontoidea Cope, 1887 sensu McKenna & Bell, 1997 & Kielan-Jaworowska & Hurum, 2001
  • Family Cimolodontidae Marsh, 1889 sensu Kielan-Jaworowska & Hurum, 2001
    • Genus Liotomus Lemoine, 1882
    • Genus Essonodon Simpson, 1927
    • Genus Anconodon Jepsen, 1940
    • Genus Cimolodon Marsh, 1889 (syn. Nanomys Marsh, 1889, Nonomyops Marsh, 1892)
    • Genus Neoliotomus Jepsen, 1930
  • Family Ptilodontidae Cope, 1887 (syn. Ptilodontinae Cope, 1887 sensu McKenna & Bell, 1997)
    • Genus Kimbetohia Simpson, 1936
    • Genus Ptilodus Cope, 1881 (Chirox Cope, 1884)
    • Genus Baiotomeus Krause, 1987
    • Genus Prochetodon Jepsen, 1940
  • Family Neoplagiaulacidae Ameghino, 1890 (syn. Neoplagiaulacinae Ameghino, 1890 sensu McKenna & Bell, 1997)
    • Genus Mesodma Marsh, 1889
    • Genus Ectypodus Matthew & Cranger, 1921 (syn. Charlesmooria Kühne, 1969)
    • Genus Mimetodon Jepsen, 1940
    • Genus Neoplagiaulax Lemoine, 1882
    • Genus Parectypodus Jepsen, 1930
    • Genus Cernaysia Vianey-Liaud, 1986
    • Genus Krauseia Vianey-Liaud, 1986
    • Genus Xyronomys Rigby, 1980
    • Genus Xanclomys Rigby, 1980
    • Genus Mesodmops Tong & Wang, 1994
  • Family Kogaionidae Rãdulescu & Samson, 1996
    • Genus Barbatodon Rãdulescu & Samson, 1996
    • Genus Hainina Vianey-Liaud, 1979
    • Genus Kogaionon Rãdulescu & Samson, 1996
  • Family Eucosmodontidae Jepsen, 1940 sensu Kielan-Jaworowska & Hurum, 2001 (syn. Eucosmodontinae Jepsen, 1940 sensu McKenna & Bell, 1997)
    • Genus Clemensodon Krause, 1992
    • Genus Eucosmodon Matthew & Granger, 1921
    • Genus Stygimys Sloan & Van Valen, 1965
  • Family Microcosmodontidae Holtzman & Wolberg, 1977 (syn. Microcosmodontinae Holtzman & Wolberg, 1977 McKenna & Bell, 1997)
    • Genus Pentacosmodon Jepsen, 1940
    • Genus Acheronodon Archibald, 1982
    • Genus Microcosmodon Jepsen, 1930
• Superfamily Djadochtatherioidea Kielan-Jaworowska & Hurum, 1997 sensu Kielan-Jaworowska & Hurum, 2001 (syn. Djadochtatheria Kielan-Jaworowska & Hurum, 1997)
  • Genus ?Bulganbaatar Kielan-Jaworowska, 1974
  • Genus ?Chulsanbaatar Kielan-Jaworowska, 1974
  • Genus Nemegtbaatar Kielan-Jaworowska, 1974
  • Family Sloanbaataridae Kielan-Jaworowska, 1974
    • Genus Kamptobaatar Kielan-Jaworowska, 1970
    • Genus Nessovbaatar Kielan-Jaworowska & Hurum, 1997
    • Genus Sloanbaatar Kielan-Jaworowska, 1974
  • Family Djadochtatheriidae Kielan-Jaworowska & Hurum, 1997
    • Genus Djadochtatherium Simpson, 1925
    • Genus Catopsbaatar Kielan-Jaworowska, 1974
    • Genus Tombaatar Kielan-Jaworowska, 1974
    • Genus Kryptobaatar Kielan-Jaworowska, 1970 (syn. Gobibaatar Kielan-Jaworowska, 1970, Tugrigbaatar Kielan-Jaworowska & Dashzeveg, 1978)
• Superfamily Taeniolabidoidea Granger & Simpson, 1929 sensu Kielan-Jaworowska & Hurum, 2001
  • Genus Prionessus Matthew & Granger, 1925
  • Family Taeniolabididae Granger & Simpson, 1929
    • Genus Kimbetopsalis Sarah, 2015
    • Genus Taeniolabis Cope, 1882
  • Family Lambdopsalidae Chow & Qi, 1978
    • Genus Sphenopsalis Matthew, Granger & Simpson, 1928
    • Genus Lambdopsalis Chow & Qi, 1978

Further reading

• Kielan-Jaworowska Z. and Hurum J.H. (2001), "Phylogeny and Systematics of multituberculate mammals". Paleontology 44, p. 389-429.
• Much of this information has been derived from https://web.archive.org/web/20120204044911/http://home.arcor.de/ktdykes/multis.htm Multituberculata Cope, 1884.

External links

• Web site: Introduction to Multituberculates. Berkeley. https://web.archive.org/web/20230220122512/https://ucmp.berkeley.edu/mammal/multis/multis.html. February 20, 2023. live.
• Web site: Multituberculates: Heyday of the longest lived mammalian order. Paleocene-mammals. https://web.archive.org/web/20221208052324/http://www.paleocene-mammals.de/multis.htm. December 8, 2022. live.
• Web site: Genera and species of Paleocene mammals - Part 1. Paleocene-mammals. https://web.archive.org/web/20230606031509/http://paleocene-mammals.de/pal1.htm. June 6, 2023. live.
• Web site: Mammals. Paleofile. https://web.archive.org/web/20230412043149/http://www.paleofile.com/Lists/Mammalia.asp. April 12, 2023. dead.

Notes and References

1. Weaver. Lucas N.. Wilson. Gregory P.. Krumenacker. L. J.. Mclaughlin. Kayla. Moore. Jason R.. Varricchio. David J.. 2019-03-04. New multituberculate mammals from the mid-Cretaceous (lower Cenomanian) Wayan Formation of southeastern Idaho and implications for the early evolution of Cimolodonta. Journal of Vertebrate Paleontology. en. 39. 2. e1604532. 10.1080/02724634.2019.1604532. 196655261 . 0272-4634.