Zhuganpo Formation Explained
The Zhuganpo Formation is a Triassic geologic unit found in southern China. It has historically been known as the Zhuganpo Member of the Falang Formation. A diverse fossil assemblage known as the Xingyi biota or Xingyi Fauna can be found in the upper part of the Zhuganpo Formation. Fossils of the Xingyi biota include articulated skeletons of marine reptiles, abundant fish, and a plentiful assortment of invertebrates indicating a Ladinian to Carnian age for the sediments of the formation.[1] [2]
Paleobiota
Invertebrates
- Ammonites: Clionitites sp.,[3] Detoniceras sp., Haoceras xingyiense,[4] Parasturia sp., Protrachyceras sp., Ptychites sp., Sinomeginoceras (S. wangi, S. xingyiense), Trachyceras sp., Xenoprotrachyceras cf. primum, Yangites densicostatus,
- Bivalves: Daonella sp.
- Conodonts: Gladigondolella malayensis,[5] Metapolygnathus / Paragondolella / Quadralella (Q. aff. acuminatus, P. foliata, P. inclinata, Q. intermedius, Q. langdaiensis, P. maantangensis, P. navicula, M. nodosus, M. parafoliata, Q. aff. praelindae, P. polygnathiformis, Q. shijiangjunensis, Q. tadpole, Q. uniformis, Q. wanlanensis, Q. aff. wayaoensis, Q. yongueensis, Q. aff. zonneveldi)
- Crinoids: Traumatocrinus hsui
- Crustaceans ("shrimps"): Schimperella acanthocercus (Lophogastrida)[6]
Fish
Reptiles
Notes and References
- Benton. Michael J.. Zhang. Qiyue. Hu. Shixue. Chen. Zhong-Qiang. Wen. Wen. Liu. Jun. Huang. Jinyuan. Zhou. Changyong. Xie. Tao. Tong. Jinnan. Choo. Brian. 2013-10-01. Exceptional vertebrate biotas from the Triassic of China, and the expansion of marine ecosystems after the Permo-Triassic mass extinction. Earth-Science Reviews. en. 125. 199–243. 10.1016/j.earscirev.2013.05.014. 2013ESRv..125..199B . 0012-8252.
- Lu . Hao . Jiang . Da-Yong . Motani . Ryosuke . Ni . Pei-Gang . Sun . Zuo-Yu . Tintori . Andrea . Xiao . Shi-Zhen . Zhou . Min . Ji . Cheng . Fu . Wan-Lu . 2018 . Middle Triassic Xingyi Fauna: Showing turnover of marine reptiles from coastal to oceanic environments . Palaeoworld . en . 27 . 1 . 107–116 . 10.1016/j.palwor.2017.05.005.
- Xiaodong Xu, Marco Balini, Da-Yong Jiang, Andrea Tintori, Zuo-Yu Sun, Yuan-Lin Sun . 2015 . Ammonoids from the Zhuganpo Member of the Falang Formation at Nimaigu and their relevance for dating the Xingyi Fossil-Lagerstatte (Late Ladinian, Guizhou, China). . Rivista Italiana di Paleontologia e Stratigrafia . 121 . 2 . 135–161.
- Sun . Zuoyu . Jiang . Dayong . Ji . Cheng . Hao . Weicheng . 2016 . Integrated biochronology for Triassic marine vertebrate faunas of Guizhou Province, South China . Journal of Asian Earth Sciences . en . 118 . 101–110 . 10.1016/j.jseaes.2016.01.004. 2016JAESc.118..101S .
- Zhang . Z. T. . Sun . Y. D. . Lai . X. L. . Joachimski . M. M. . Wignall . P. B. . 2017-11-15 . Early Carnian conodont fauna at Yongyue, Zhenfeng area and its implication for Ladinian-Carnian subdivision in Guizhou, South China . Palaeogeography, Palaeoclimatology, Palaeoecology . The Palaeozoic-Mesozoic Transition in South China: Oceanic Environments and Life from Late Permian to Late Triassic . en . 486 . 142–157 . 10.1016/j.palaeo.2017.02.011 . 2017PPP...486..142Z . 0031-0182.
- Taylor . Rod S. . Schram . Frederick R. . Yan-Bin . Shen . 2001 . A new upper Middle Triassic shrimp (Crustacea: Lophogastrida) from Guizhou, China, with discussion regarding other fossil "mysidaceans" . Journal of Paleontology . en . 75 . 2 . 310–318 . 10.1666/0022-3360(2001)075<0310:ANUMTS>2.0.CO;2 . 131509616 . 0022-3360.
- Li . Jiachun . Sun . Zuoyu . Cuny . Gilles . Ji . Cheng . Jiang . Dayong . Zhou . Min . 2022 . Cavin . Lionel . An unusual shark assemblage from the Ladinian–Carnian interval of South China . Papers in Palaeontology . en . 8 . 1 . 10.1002/spp2.1404 . 2022PPal....8E1404L . 241980979 . 2056-2799.
- Xu . Guang-Hui . Ma . Xin-Ying . 2018-01-27 . Redescription and phylogenetic reassessment of Asialepidotus shingyiensis (Holostei: Halecomorphi) from the Middle Triassic (Ladinian) of China . Zoological Journal of the Linnean Society . 184 . 1 . 95–114 . 10.1093/zoolinnean/zlx105 . 0024-4082.
- Ni . Peigang . Tintori . Andrea . Sun . Zuoyu . Lombardo . Cristina . Jiang . Dayong . 2019-12-01 . Postcranial skeleton of Birgeria liui (Osteichthyes, Actinopterygii) from the Longobardian (Ladinian, Middle Triassic) of Xingyi, Guizhou, South China . Swiss Journal of Geosciences . en . 112 . 2 . 307–324 . 10.1007/s00015-018-0329-0 . 135305199 . 1661-8734.
- Xu . Guang-Hui . Ma . Xin-Ying . Ren . Yi . 2018-12-20 . Fuyuanichthys wangi gen. et sp. nov. from the Middle Triassic (Ladinian) of China highlights the early diversification of ginglymodian fishes . PeerJ . en . 6 . e6054 . 10.7717/peerj.6054 . 30595977 . 6304272 . 2167-8359 . free .
- Guang-Hui . Xu . 2020 . A new species of Luganoia (Luganoiidae, Neopterygii) from the Middle Triassic Xingyi Biota, Guizhou, China . Vertebrata PalAsiatica . 58 . 4 . 267–282 . 10.19615/j.cnki.1000-3118.200624.
- TINTORI . ANDREA . SUN . ZUOYU . NI . PEIGANG . LOMBARDO . CRISTINA . JIANG . DAYONG . MOTANI . RYOSUKE . 2015-11-19 . OLDEST STEM TELEOSTEI FROM THE LATE LADINIAN (MIDDLE TRIASSIC) OF SOUTHERN CHINA . Rivista italiana di Paleontologia e Stratigrafia . en . 121 . 3 . 10.13130/2039-4942/6519.
- Xu Guang-Hi, Ma Xin-Ying, Zhao Li-Jun . 2018 . A large peltopleurid fish (Actinopterygii: Peltopleuriformes) from the Middle Triassic of Yunnan and Guizhou, China . Vertebrata PalAsiatica . 56 . 2 . 106–120 . 10.19615/j.cnki.1000-3118.171225.
- Shen . Chenchen . Arratia . Gloria . 2021-10-02 . Re-description of the sexually dimorphic peltopleuriform fish Wushaichthys exquisitus (Middle Triassic, China): taxonomic implications and phylogenetic relationships . Journal of Systematic Palaeontology . en . 19 . 19 . 1317–1342 . 10.1080/14772019.2022.2029595 . 2021JSPal..19.1317S . 247731689 . 1477-2019.
- Xu . G. -H. . Zhao . L. -J. . Gao . K. -Q. . Wu . F. -X. . 2012 . A new stem-neopterygian fish from the Middle Triassic of China shows the earliest over-water gliding strategy of the vertebrates . Proceedings of the Royal Society B: Biological Sciences . 280 . 1750 . 20122261 . 10.1098/rspb.2012.2261 . 3574442 . 23118437.
- Xu . Guang-Hui . Zhao . Li-Jun . Shen . Chen-Chen . 2015 . A Middle Triassic thoracopterid from China highlights the evolutionary origin of overwater gliding in early ray-finned fishes . Biology Letters . en . 11 . 1 . 20140960 . 10.1098/rsbl.2014.0960 . 1744-9561 . 4321159 . 25568155.
- Cheng . L. . Chen, X. . Wang, C. . 2007 . A new species of Late Triassic Anshunsaurus (Reptilia: Thalattosauria) from Guizhou Province . Acta Geologica Sinica . zh . 81 . 10 . 1345–1351.
- Long . Cheng . Xiaohong . Chen . Baomin . Zhang . Yongjian . Cai . 2011 . New Study of Anshunsaurus huangnihensis Cheng, 2007 (Reptilia: Thalattosauria): Revealing its Transitional Position in Askeptosauridae . Acta Geologica Sinica - English Edition . en . 85 . 6 . 1231–1237 . 10.1111/j.1755-6724.2011.00584.x. 2011AcGlS..85.1231C . 129819570 .
- Rieppel . O. . Liu, J. . Li, C. . 2006 . A new species of the thalattosaur genus Anshunsaurus (Reptilia: Thalattosauria) from the Middle Triassic of Guizhou Province, southwestern China . Vertebrata PalAsiatica . 44 . 285–296.
- Liu . J. . 2007 . A juvenile specimen of Anshunsaurus (Reptilia: Thalattosauria) . American Museum Novitates . 3582 . 1–9 . 10.1206/0003-0082(2007)3582[1:ajsoar]2.0.co;2 . 0003-0082 . 85916769.
- Shang . Qing-Hua . Wu . Xiao-Chun . Li . Chun . 2020-05-03 . A New Ladinian Nothosauroid (Sauropterygia) from Fuyuan, Yunnan Province, China . Journal of Vertebrate Paleontology . 40 . 3 . e1789651 . 10.1080/02724634.2020.1789651 . 2020JVPal..40E9651S . 227241106 . 0272-4634. free .
- Li . C. . Wu . X. C. . Zhao . L. J. . Sato . T. . Wang . L. T. . 2012 . A new archosaur (Diapsida, Archosauriformes) from the marine Triassic of China . Journal of Vertebrate Paleontology . 32 . 5 . 1064 . 10.1080/02724634.2012.694383 . 2012JVPal..32.1064L . 86797826.
- Michelle R. Stocker . Li-Jun Zhao . Sterling J. Nesbitt . Xiao-Chun Wu . Chun Li . 2017 . A Short-Snouted, Middle Triassic Phytosaur and its Implications for the Morphological Evolution and Biogeography of Phytosauria . Scientific Reports . 7 . Article number 46028 . 2017NatSR...746028S . 10.1038/srep46028 . 5385495 . 28393843.
- Wang . Xue . Lu . Hao . Jiang . Da-Yong . Zhou . Min . Sun . Zuo-Yu . 2020 . A new specimen of Yunguisaurus (Reptilia; Sauropterygia) from the Ladinian (Middle Triassic) Zhuganpo Member, Falang Formation, Guizhou, China and the restudy of Dingxiaosaurus . Palaeoworld . en . 29 . 1 . 137–150 . 10.1016/j.palwor.2019.05.006. 181711576 .
- Fraser . Nicholas C. . Rieppel . Olivier . Chun . Li . 2013 . A long-snouted protorosaur from the Middle Triassic of southern China . Journal of Vertebrate Paleontology . en . 33 . 5 . 1120–1126 . 10.1080/02724634.2013.764310 . 2013JVPal..33.1120F . 83521468 . 0272-4634.
- Ezcurra . Martín D. . Butler . Richard J. . 2018-06-13 . The rise of the ruling reptiles and ecosystem recovery from the Permo-Triassic mass extinction . Proc. R. Soc. B . en . 285 . 1880 . 20180361 . 10.1098/rspb.2018.0361 . 0962-8452 . 6015845 . 29899066.
- Spiekman . Stephan N. F. . Fraser . Nicholas C. . Scheyer . Torsten M. . 2021-05-03 . A new phylogenetic hypothesis of Tanystropheidae (Diapsida, Archosauromorpha) and other "protorosaurs", and its implications for the early evolution of stem archosaurs . PeerJ . en . 9 . e11143 . 10.7717/peerj.11143 . 33986981 . 8101476 . 2167-8359 . free .
- Zhao . L.-J. . Li, C. . Liu, J. . He, T. . 2008 . A new armored placodont from the Middle Triassic of Yunnan Province, Southwestern China . Vertebrata PalAsiatica . 46 . 3 . 171–177 . https://web.archive.org/web/20160303233554/http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/200811/W020090813369302050276.pdf . 3 March 2016.
- Jiang . D. Y. . Motani . R. . Tintori . A. . Rieppel . O. . Ji . C. . Zhou . M. . Wang . X. . Lu . H. . Li . Z. G. . 2020 . Evidence supporting predation of 4-m marine reptile by Triassic megapredator . iScience . 23 . 9 . 101347 . 2020iSci...23j1347J . 10.1016/j.isci.2020.101347 . 7520894 . 32822565.
- Cheng . Yen-nien . Wu . Xiao-chun . Ji . Qiang . 2004 . Triassic marine reptiles gave birth to live young . Nature . en . 432 . 7015 . 383–386 . 10.1038/nature03050 . 15549103 . 2004Natur.432..383C . 4391810 . 1476-4687.
- Holmes . Robert . Cheng . Yen-Nien . Wu . Xiao-Chun . 2008-03-12 . New information on the skull of Keichousaurus hui (Reptilia: Sauropterygia) with comments on sauropterygian interrelationships . Journal of Vertebrate Paleontology . 28 . 1 . 76–84 . 10.1671/0272-4634(2008)28[76:NIOTSO]2.0.CO;2 . 85868385 . 0272-4634.
- Rieppel . Olivier . Jinling . Li . Jun . Liu . 2003 . Lariosaurus xingyiensis (Reptilia: Sauropterygia) from the Triassic of China . Canadian Journal of Earth Sciences . 40 . 4 . 621–634 . en . 10.1139/e02-067. 2003CaJES..40..621R .
- Wen-Bin Lin . Da-Yong Jiang . Olivier Rieppel . Ryosuke Motani . Cheng Ji . Andrea Tintori . Zuo-Yu Sun . Min Zhou . 2017 . A new specimen of Lariosaurus xingyiensis (Reptilia, Sauropterygia) from the Ladinian (Middle Triassic) Zhuganpo Member, Falang Formation, Guizhou, China . Journal of Vertebrate Paleontology . 37 . 2 . e1278703 . 10.1080/02724634.2017.1278703. 2017JVPal..37E8703L . 90181915 .
- Li . Chun . Wu . Xiao-chun . Zhao . Li-jun . Nesbitt . Sterling J. . Stocker . Michelle R. . Wang . Li-Ting . 2016-11-09 . A new armored archosauriform (Diapsida: Archosauromorpha) from the marine Middle Triassic of China, with implications for the diverse life styles of archosauriforms prior to the diversification of Archosauria . The Science of Nature . en . 103 . 11 . 95 . 10.1007/s00114-016-1418-4 . 27830290 . 2016SciNa.103...95L . 253634068 . 1432-1904.
- Jiang . Da-Yong . Rieppel . Olivier . Fraser . Nicholas C. . Motani . Ryosuke . Hao . Wei-Cheng . Tintori . Andrea . Sun . Yuan-Lin . Sun . Zuo-Yu . 2011 . New information on the protorosaurian reptile Macrocnemus fuyuanensis Li et al., 2007, from the Middle/Upper Triassic of Yunnan, China . Journal of Vertebrate Paleontology . en . 31 . 6 . 1230–1237 . 10.1080/02724634.2011.610853 . 2011JVPal..31.1230J . 131615836 . 0272-4634.
- Ji . Cheng . Jiang . Da-Yong . Rieppel . Olivier . Motani . Ryosuke . Tintori . Andrea . Sun . Zuo-Yu . 2014-03-01 . A new specimen of Nothosaurus youngi from the Middle Triassic of Guizhou, China . Journal of Vertebrate Paleontology . 34 . 2 . 465–470 . 10.1080/02724634.2013.808204 . 2014JVPal..34..465J . 84886121 . 0272-4634.
- Yang, P.F., Ji, C., Jiang, D.Y., Motani, R., Tintori, A., Sun, Y.L., Sun, Z.Y. . 2013 . A new species of Qianichthyosaurus (Reptilia: Ichthyosauria) from Xingyi Fauna (Ladinian, Middle Triassic) of Guizhou. . Acta Scientiarum Naturalium Universitatis Pekinensis . Chinese . 49 . 1002–1008.
- Cheng . Y. N. . Wu . X. C. . Sato . T. . Shan . H. Y. . 2012 . A new eosauropterygian (Diapsida, Sauropterygia) from the Triassic of China . Journal of Vertebrate Paleontology . 32 . 6 . 1335 . 10.1080/02724634.2012.695983 . 2012JVPal..32.1335C . 85253142.
- Spiekman . Stephan N. F. . Neenan . James M. . Fraser . Nicholas C. . Fernandez . Vincent . Rieppel . Olivier . Nosotti . Stefania . Scheyer . Torsten M. . 2020-11-20 . The cranial morphology of Tanystropheus hydroides (Tanystropheidae, Archosauromorpha) as revealed by synchrotron microtomography . PeerJ . en . 8 . e10299 . 10.7717/peerj.10299 . 2167-8359 . 7682440 . 33240633 . free .
- Rieppel . Olivier . Jiang . Da-Yong . Fraser . Nicholas C. . Hao . Wei-Cheng . Motani . Ryosuke . Sun . Yuan-Lin . Sun . Zuo-Yu . 2010 . Tanystropheus cf. T. Longobardicus from the early Late Triassic of Guizhou Province, southwestern China . Journal of Vertebrate Paleontology . 30 . 4 . 1082–1089 . 10.1080/02724634.2010.483548 . 40864387 . 2010JVPal..30.1082R . 86315078.
- Ma . Le-Tian . Jiang . Da-Yong . Rieppel . Olivier . Motani . Ryosuke . Tintori . Andrea . 2015-01-02 . A new pistosauroid (Reptilia, Sauropterygia) from the late Ladinian Xingyi marine reptile level, southwestern China . Journal of Vertebrate Paleontology . 35 . 1 . e881832 . 10.1080/02724634.2014.881832 . 2015JVPal..35E1832M . 130696958 . 0272-4634.
- Li . Z.-G. . Jiang . D.-Y. . Rieppel . O. . Motani . R. . Tintori . A. . Sun . Z.-Y. . Ji . C. . 2016-11-01 . A new species of Xinpusaurus (Reptilia, Thalattosauria) from the Ladinian (Middle Triassic) of Xingyi, Guizhou, southwestern China . Journal of Vertebrate Paleontology . 36 . 6 . e1218340 . 10.1080/02724634.2016.1218340 . 2016JVPal..36E8340L . 0272-4634 . 132418823.
- Sato . Tamaki . Zhao . Li-Jun . Wu . Xiao-Chun . Li . Chun . 2014 . Ruta . Marcello . A new specimen of the Triassic pistosauroid Yunguisaurus, with implications for the origin of Plesiosauria (Reptilia, Sauropterygia) . Palaeontology . en . 57 . 1 . 55–76 . 10.1111/pala.12048. 2014Palgy..57...55S . 129457774 . free .