Y-DNA haplogroups by ethnic group explained

The various ethnolinguistic groups found in the Caucasus, Central Asia, Europe, the Middle East, North Africa and/or South Asia demonstrate differing rates of particular Y-DNA haplogroups.

In the table below, the first two columns identify ethnolinguistic groups. Subsequent columns represent the sample size (n) of the study or studies cited, and the percentage of each haplogroup found in that particular sample.

(Data from studies conducted before 2004 may be inaccurate or a broad estimate, due to obsolete haplogroup naming systems – e.g. the former Haplogroup 2 included members of the relatively unrelated haplogroups known later as Haplogroup G and macrohaplogroup IJ [which comprises haplogroups I and J].)

PopulationLanguage family[1] n[2] R1b[3] n R1an In E1b1bn E1b1an Jn Gn Nn Tn Ln H
AlbaniansIE (Albanian)106
25/106
44
11/44
56
13/56

AlbaniansIE (Albanian)51 51
5/51
51 51 51
12/51
51
1/51
51 51 51
Kosovo Albanians (Pristina)IE (Albanian)114
24/114
114
5/114
114
9/114
114
54/114
114
19/114
Albanians (Tirana)IE (Albanian)30 [4] 30 30 30 30 30
6/30
30
1/30
AlbaniansIE (Albanian)55
10/55
55
5/55
55
12/55
55
14/55
55 55
13/55
55
1/55
55 55 55
Albanians (North Macedonia)IE (Albanian)64
12/64
64
1/64
64
11/64
64
25/64
64 64
14/64
64
1/64
64 64 64
Albanians + Albanians (North Macedonia)IE (Albanian)55+
64
119

22/119
55+
64
119

6/119
55+
64
119

23/119
55+
64
119

39/119
55+
64
119

27/119
55+
64
119

2/119
AbaziniansNorthwest Caucasian14 [5] 14 14 14 14 14 [6] 14
AbkhazNorthwest Caucasian12 12 12 12 12 12 12
AltaiansTurkic, Siberia142
Altaians (Northern)Turkic, Siberian50 [7] 50 50 50 50 50 50
Altaians (Southern)Turkic, Siberia96 96 96 96 96 96
AmbalakararDravidian (Southern)29 29 29 29 29 29 29 29 29 29
AmharaAfro-Asiatic (Semitic)48 [8] 48 48 48 48 48 48 48 48
AndalusiansIE (Romance)29 29 103 76 93 29 29 29 29
AndisNortheast Caucasian49 49 49 49 49 49 49 49 49
Arabs (Algeria)Afro-Asiatic (Semitic)35 [9] 35 32 35
Arabs (Algeria - Oran)Afro-Asiatic (Semitic)102 [10] 102 102 102 102
Arabs (Bedouin) Afro-Asiatic (Semitic)32 32 32 32 32 32
Arabs (Bedouins, Sinai Peninsula)Afro-Asiatic (Semitic)147 [11] 147 147 147 147 147 147 147 147
Arabs (Iraq)Afro-Asiatic (Semitic)218 218 156
Arabs (Palestine)Afro-Asiatic (Semitic)143 143 143 143 143 143
Arabs (Morocco)Afro-Asiatic (Semitic)44 44 44 49 49
Arabs (Oman)Afro-Asiatic (Semitic)121 121 121 121 121 121 121 121 121
Arabs (Qatar)Afro-Asiatic (Semitic)72 [12] 72 72 72 72 72 72 72 72 72
Arabs (Saudi Arabia)Afro-Asiatic (Semitic)157 [13] 157 157 157 157 157 157 157 157 157
Arabs (UAE)Afro-Asiatic (Semitic)164 164 164 164 164 164 164 164 164
Arabs (Yemen)Afro-Asiatic (Semitic)62 62 62 62 62 62 62 62 62 62
Arabs (Syria)Afro-Asiatic (Semitic)20 20 20 20 20 20 20 20 20
Arabs (Lebanon)Afro-Asiatic (Semitic)31 31 31 31 31 31 31 31 31
Arabs (Sudan)Afro-Asiatic (Semitic)102 102 102 102
Arabs (Tunisia)Afro-Asiatic (Semitic)148 148 148 148 148 148 148 148 148
Arabs (Libya)Afro-Asiatic (Semitic)63 [14] 63 63 63 63 63 63 63 63
ArmeniansIE (Armenian)89 89 100 89 89 100 89
ArmeniansIE (Armenian)734 [15] 734 734 734
Aromanians (Dukasi, Albania)IE (Romance)39
1/39
39
1/39
39
7/39
39
7/39
39 39
19/39
39
4/39
39 39 39
Aromanians (Andon Poci, Albania)IE (Romance)19
7/19
19 19
8/19
19
3/19
19 19
1/19
19 19 19 19
Aromanians (Kruševo, North Macedonia)IE (Romance)43
12/43
43
5/43
43
9/43
43
9/43
43 43
5/19
43
3/19
43 43 43
Aromanians (Štip, North Macedonia)IE (Romance)65
15/65
65
14/65
65
11/65
65
12/65
65 65
13/65
65 65 65 65
Aromanians (Romania)IE (Romance)42
10/42
42
1/42
42
8/42
42
3/42
42 42
14/42
42
Aromanians (Balkan)IE (Romance)39+
19+
43+
65+
42=
208

45/208
208
21/208
208
43/208
208
34/208
208 208
52/208
208
7/208
Ashkenazi JewsAfro-Asiatic (Semitic)79 [16] 79
18/79
79
34/79
79
Ashkenazi JewsAfro-Asiatic (Semitic)442 [17] 442
87/442
442 442
168/442
442
43/442
442 442
AustriansIE (Germanic, West)219 219
AvarsNortheast Caucasian42 [18] 42 42 42 42
30/42
42 42 42 42
AzerbaijanisTurkic, Oghuz72 [19] 72 97
Azerbaijanis (West Azerbaijan Province)Turkic, Oghuz63 [20] 63 63
BagvalalNortheast Caucasian28 28 28 28 28 28 28 28 28
BalkarianTurkic, Kipchak39 16
BalkariansTurkic, Kipchak38 38 38 38 38 38 38 38 38 38
BalochIE (Iranian, NW)25 25 25 25 25 25 25 25 25 25
Bashkirs (Perm)Turkic, Kipchak43
37/43[21]
43 43 43 43 43 43 43 43 43
Basque (France, Spain)Basque (Basque)67
59/67
67 67
5/67
67
1/67
67
2/67
67 67 67 67
German BavariansIE (Germanic, West)80
40/80
80 80 80 80 80
BelgiansIE (Germanic/Romance)92
58/92
92 92
BelarusiansIE (Slavic, East)41 41 147 41 41
BelarusiansIE (Slavic, East)68 [22] 68 68 68 68 68
BelarusiansIE (Slavic, East)306 [23] 306 306 306 306
BearnaisIE (Romance)26 43 26
BejaAfro-Asiatic (Cushitic)42
2/42
42 42 42
22/42
42 42
16/42
42 42 42 42
Berbers (Marrakesh)Afro-Asiatic (Berber)29[24]
Berbers (Moyen Atlas)Afro-Asiatic (Berber)69
Berbers (Mozabite)Afro-Asiatic (Berber)20
Berbers (Morocco)Afro-Asiatic (Berber)64 64 103
Berbers (north Morocco)Afro-Asiatic (Berber)43 43 [25]
Berbers (north-central Morocco)Afro-Asiatic (Berber)63 63
Berbers (central Morocco)Afro-Asiatic (Berber)187 187
Berbers (southern Morocco)Afro-Asiatic (Berber)40 40
Berbers (southern Morocco)Afro-Asiatic (Berber)65 65
Berbers (northern Tunisia)Afro-Asiatic (Berber)32 [26]
Berbers (southern Tunisia)Afro-Asiatic (Berber)27
Borgu (Sudan)Nilo-Saharan (Maban)26 26 26 26 26 26 26 26 26 26
Bosnians (Zenica)IE (Slavic, South)69
1/69
69
17/69
69
37/69
69
7/69
69 69
3/69
Brahmins (Konkanastha)IE (Indo-Aryan)25 25 25 25 25 25 25 25 25 25
BrahuiDravidian (Northern)110 [27] 110 110 110 110 110
BrahuiDravidian (Northern)25 [28] 25 25 25 25 25 25 25 25 25
BritishIE (Germanic, West)32
22/32
32
3/32
BulgariansIE (Slavic, South)[29]
BulgariansIE (Slavic, South)[30]
BulgariansIE (Slavic, South)[31]
BurushoBurushaski (isolate)97 97 97 97 97 97 97 97 97
Spanish (Catalan peopleCatalans)IE (Romance)24
19/24
24 32 33 28 24
Cantabrians (Pasiegos)IE (Romance)56
ChamalinsNortheast Caucasian27 27 27 27 27 27 27 27 27
ChechensNortheast Caucasian330 [32] 330 330 330 330 330 330 330
ChuvashesTurkic, Oghur79 79 79 79 79 79 79 79 79 79
Copts (Sudan)Afro-Asiatic 33 [33] 33 33
Croats (mainland)IE (Slavic, South)108
17/108[34]
108
37/108
108
40/108[35]
108
6/108
108 108
2/108
108
1/108
Croat (mainland)IE (Slavic, South)189
72/189
Cross River (Nigeria)Cross River (NC)1113 [36] 1113 1113 1113
CypriotsIE (Greek)45 45 45
CzechsIE (Slavic, West)257 [37] 257 257 257 257 257
Czechs and SlovaksIE (Slavic, West)45 45 198 45
DanesIE (Germanic, North)12 12 194 35
DarginsNortheast Caucasian68 68 68 68 68 68 68
DarginsNortheast Caucasian26 26 26 26 26 26 26 26 26
DolgansTurkic, Siberia67 67 67 67
Druze (Arabs)Afro-Asiatic (Semitic)28
DutchIE (Germanic, West)27 27 30 84 34
EgyptiansAfro-Asiatic (Semitic)92 [38] 92 92 92 92 92 92 92 92 92
Egyptians (North)Afro-Asiatic (Semitic)43 43 43 43 44 43 43 43
Egyptians (South)Afro-Asiatic (Semitic)29 29 29 29 29 29 29 29
English (Central)IE (Germanic, West)215 215
EstoniansUralic (Finnic)207 118 210 207 207 207
FinnsUralic (Finnic)57 57 57 57
FinnsUralic (Finnic)38 38 38 38
FrenchIE (Romance)23 23 23 40
FrisiansIE (Germanic, West)94 94 94 94 94
Frieslanders (Netherlands)IE (Germanic, West)94 [39] 94 94 94 94
FurNilo-Saharan (Fur)32 32 32 32 32 32 32 32 32 32
Gagauz (Kongaz)Turkic, Oghuz48 [40] 48 48 48 48 48 48 48
Gagauz (Etulia)Turkic, Oghuz41 41 41 41 41 41 41 41
IE (Germanic, West)3,000+ [41]
U106=18
L21=5
U152=9
DF27, DF19=9
others=3
3,000+ 3,000+
I1=13
I2=7
3,000+
V13=5
3,000+
J2=5
J1=1
3,000+ 3,000+
Germans (West)IE (German)48 48 16 16
Germans (East)IE (German)
Germans (Berlin)IE (German)103 [42] 103 103 103 103
GeorgiansKartvelian63 63 63 64 63 63 63 63
GeorgiansKartvelian66 66 66 66 66 66 66 66 66 66
GreeksIE (Greek)92 [43] 92 92 92 92 92 92 92
GreeksIE (Greek)77
9/77
77
12/77
77
15/77
77
16/77
77 77
13/77
77
7/77
77 77 77
others
5/77
GreeksIE (Greek)118 [44] 118 261 84 92
GreeksIE (Greek)171
23/171[45]
171
19/171
171
27/171
171
54/171
171
34/171
171
8/171
171
3/171
Greeks (Crete)IE (Greek)193
33/193
193
17/193
193
25/193
193
17/193
193
75/193
193
21/193
193
4/193
Greeks (Crete)IE (Greek)171+
193=
364

56/364
364
36/364
364
52/364
364
71/364
364
109/364
364
29/364
364
7/364
Greeks (Peloponnese)IE (Greek)36
17/36
36
5/36
Greeks (Thrace)IE (Greek)41 41 41 41 41 41 41
Greeks (North)IE (Greek)96 =
14/96
96
18/96
96
12/96
96
34/96
96
5/96
96
2/96
96
1/96
Greeks (South)IE (Greek)46
9/46
46
1/46
46
11/46
46
20/46
46
3/46
46
1/46
46
Greeks (North) + (South)IE (Greek)96+
46=
142

23/142
96+
46=
142

19/142
96+
46=
142

23/142
96+
46=
142

54/142
96+
46=
142

8/142
96+
46=
142

3/142
96+
46=
142

1/142
Hausa (Sudan)Afro-Asiatic (Chadic)32 32 32 32 32 32 32 32 32 32
Herzegovinians (Mostar, Široki Brijeg)IE (Slavic, South)141
5/141
141
17/141
141
90/141
141
12/141
141
2/141
HungariansUralic (Ugric)45 113 162 53 49 103 [46] 103
IcelandersIE (Germanic, North)181
75/181
181
43/181
181
62/181
IngushNortheast Caucasian143 143 143 143 143 143 143
Iranians (North Iran)IE (Iranian, West)33 [47] 33 33 33 33 33 33 33 33 33
Iranians (South Iran)IE (Iranian, West)117 117 117 117 117 117 117 117 117 117
IrishIE (Celtic)222
181/222
222 222
33/222
257
ItaliansIE (Romance)50
31/50[48]
332 50 99 50
Italians (Calabria)IE (Romance)37 148 80 57
Italians (Apulia)IE (Romance)78 86 86
Italians (North-central)IE (Romance)50 390 212 52
Italians (South)IE (Romance)68 68 68 68 68 68 68
Italians (Sicily)IE (Romance)51 55 42
Italians (East Sicily)IE (Romance)87 87 87 87 87 87 87
Italians (West Sicily)IE (Romance)125 125 125 125 125 125 125
IyengarDravidian (Southern)30 30 30 30 30 30 30 30 30 30
IyerDravidian (Southern)29 29 29 29 29 29 29 29 29 29
Japan2390 2390 [49]
KabardinianNorthwest Caucasian
Kalash (Pakistan)IE (Dardic)44 44 44 44 44 44 44 44
KarakalpaksTurkic, Kipchak44 44 44 44 44
Kazakhs (more)Turkic1982 [50] 1982 1982 1982 1982
Kazakhs (more)Turkic, Kipchak54 54 54 54 38 54 54
KhantsUralic (Ugric)47
9/47
47
2/47
47 47
36/47
KomiUralic (Finnic)94 94 94 94
Komi (Izhemsky)Uralic (Finnic)54 [51] 54
16/54
54 54 54 54 54 54
37/54
54 54
Komi (Priluzsky)Uralic (Finnic)49 49
16/49
49 49 49 49 49 49
30/49
49 49
KumyksTurkic, Kipchak76 76 76 76 76 76 76 76 76
Kurds (Northern Iraq)IE (Iranian, NW)95 95 95 95 95 95 95
KyrgyzTurkic, Kipchak52 [52] 52 52 52 41 41 52
LatviansIE (Baltic)34 114 86 114 114
LebaneseAfro-Asiatic (Semitic)1,116 & Haber2011[53] 1,116 & Haber20111,116 & Haber20111,116 & Haber20111,116 & Haber20111,116 & Haber20111,116 & Haber20111,116 & Haber20111,116 & Haber20111,116 & Haber2011
LebaneseAfro-Asiatic (Semitic)914 [54] 914 914 914 914 914 914 914 914 914
LezginsNortheast Caucasian31 31 31 31 31 31 31 31 31
LithuaniansIE (Baltic)38 114 [55] 114 114
Macedonians (Skopje)IE (Slavic, South)79 79 79 79 79 [56] [57] 79
Macedonians (Skopje)IE (Slavic, South)52 52 52 52 52 52
Macedonian Romani (Skopje)IE (Indo-Aryan)57 57 57 57 57
MalteseAfro-Asiatic (Semitic)187 187 187 187 187 187 187
MarathaIE (Indo-Aryan)20 20 20 20 20 20
MelanesiaOceanic, Papuan1272
Moldovans (Karahasani)IE (Romance)72 72 72 72 72 72 72 72
Moldovans (Sofia)IE (Romance)54 54 54 54 54 54 54 54
ErzyaUralic (Finnic)46 [58]
MokshaUralic (Finnic)46
MokshaUralic (Finnic)83 83 83 83
MariUralic (Finnic)111 111 111 111
MariUralic (Finnic)48 48 48 48 48
MasalitNilo-Saharan (Maban)32 32 32 32
23/32
32 32 32 32 32 32
MoroccansAfro-Asiatic (Berber)760 [59] 760 760 760 760 760 760
530 [60] 530
NorwegiansIE (Germanic, North)112 112 72
NorwegiansIE (Germanic, North)52 52 52 52 52 52
Nubians (Sudan)Nilo-Saharan (Eastern Sudanic)39 39 39 39 39 39 39 39 39
OrcadiansIE (Celtic/Germanic)71
47/71
71
14/71
OromoAfro-Asiatic (Cushitic)78 78 78 78
62/78
78 78 78 78 78
OssetiansIE (Iranian, NE)47 47 47 47 [61]
PallanDravidian (Southern)29 29 29 29 29 29 29 29 29 29
PashtunsIE (Iranian, SE)96 [62] 96 96 96 96 96 96 96 96
PolesIE (Slavic, West)55 55 191 99 97
PolesIE (Slavic, West)93 93 93 93
PortugueseIE (Romance)303
Portuguese (South)IE (Romance)57 57 57
Portuguese (North)IE (Romance)328 328 328
RajputsIE (Indo-Aryan)29 29 29 29 29 29
Romani (Gypsy)IE (Indo-Aryan)
RomaniansIE (Romance)54 54 54 54 54 54 54 54
RomaniansIE (Romance)361
Romanians (Constanța)IE (Romance)31 31 31 31 31 31 31 31 31 31
Romanians (Ploiești)IE (Romance)36 36 36 36 36 36 36 36 36 36
RussiansIE (Slavic, East)122 122 122 122 122
RussiansIE (Slavic, East)414 [63] 414 414 414 414 414 414 414
Russians (Northern)(Arkhangelsk region)IE (Slavic, East)114 114 114 114 114 114 114 114 114 114
Russians (Central)IE (Slavic, East)364 [64] 364 364 364 364 364 364 364 364
Russians (Southerns)IE (Slavic, East)484 484 484 484 484 484 484 484
Russians(Oryol region)IE (Slavic, East)110 110 110 110 110 110 110 110
Russians(Voronezh region)IE (Slavic, East)96 96 96 96 96 96 96 96
Russians(Tver region)IE (Slavic, East)73 73 73 73 73 73 73 73
Russians (Kuban Cossacks)IE (Slavic, East)90 90 90 90 90 90 90 90
Russians (Novgorod region)IE (Slavic, East)38 38 38
Sami (Sweden)Uralic (Finnic)38 [65] 38
6/38
38
12/38
38 38 38 38 38
17/38
38 38
SamiUralic (Finnic)127 127 35 [66] 127
60/127
Saharawis (Western Sahara)Afro-Asiatic (Semitic)29
23/29
29 29
Sardinians (Italy)IE (Romance)77 142 139 144
Sardinians (Northern Sardinia)IE (Romance)86 86 86 86 86 86 86
Sardinians (Southern Sardinia)IE (Romance)187 187 187 187 187 187 187
ScotsIE (Celtic)61
47/61
61 178
Serbs (Serbia)IE (Slavic, South)113 113 113
41/113
113 113
Serbs (Bosnia)IE (Slavic, South)81 81 81
33/81
81
8/81
81 81 81 81 81 81
SloveniansIE (Slavic, South)75
16/75
75
29/75
75
23/75
75 75 75 75 75 75 75
SlovenianIE (Slavic, South)70
26/70[67]
70 70 70 70 70
SlovenianIE (Slavic, South)55
21/55[68]
Dir and IsaaqAfro-Asiatic (Cushitic)201 [69] 201 201 201
156/201
201 201 201 201 201 201
SpanishIE (Romance)126 126 126
Spanish (Ibiza)IE (Romance)54
31/54[70]
54 54 54 54 54 54
Spanish (Majorca)IE (Romance)62
41/62
62 62 62 62 62 62
Spanish (Menorca)IE (Romance)37
27/37
37 37 37 37 37 37
Spanish (South)IE (Romance)162 162 162 162 162 162 162
Spanish (Valencia)IE (Romance)73 73 73 73 73 73 73
Swedes (Northern)IE (Germanic, North)48 48 57 48 48 48
SwedesIE (Germanic, North)110 110 225
SwedesIE (Germanic, North)160 [71] 160 160 160 160 160
SwissIE (German/Romance)144
TabassaransNortheast Caucasian43 43 43 43 43 43 43 43 43
Tamil Nadu (India)Dravidian1680 1680 [72] 1680 1680 1680 1680 1680 1680
TatarsTurkic, Kipchak126 [73] 126 126 126
TurkmensTurkic, Oghuz30 30 30 30 21 21 30
TurksTurkic, Oghuz523 [74] 523 741 523 523 523 523 523 523
TurksTurkic, Oghuz167 167 167 167 167
TurksTurkic, Oghuz59 59 59 59 59 59 59 59
Turks (Central Anatolia)Turkic, Oghuz61
Turks (Istanbul)Turkic, Oghuz46 73
Turks (Konya)Turkic, Oghuz117 129
Turks (Cypriot)Turkic, Oghuz46
Turks (Southeastern)Turkic, Oghuz24
Turks (Erzurum)Turkic, Oghuz25
UdmurtUralic (Finnic)87 87 87 87
UkrainiansIE (Slavic, East)50 50 50 50 50 50 50 50 50
UkrainiansIE (Slavic, East)53 53 53 93 53 53 53 53
Uyghurs (Kazakhstan)Turkic, Karluk41 41 41 41 41 33 [75] 41 41 41
Uyghurs (Xinjiang)Turkic, Karluk68 [76] 68 68 68 68 68 68 67 [77] 67 68
Uyghurs (Ürümqi)Turkic, Karluk49 [78] 49 49 49 49
Uyghurs (Northwest China)Turkic, Karluk50 [79] doi:10.1038/jhg.2010.30, Link to the table of haplogroup distribution and Y-chromosome diversity in 14 northwestern populations50 50
Uyghurs (Ürümqi)Turkic, Karluk31 [80] 31 31 31 31 31 31
Uyghurs (Yili)Turkic, Karluk39 39 39 39 39
UzbeksTurkic, Karluk366 366 366 366 28 28 366
VanniyarDravidian (Southern)25 25 25 25 25 25
VellalarDravidian (Southern)31 31 31 31 31 31 31 31 31 31
Welsh (Anglesey)IE (Celtic)88
78/88[81]
88 196 88
YagnobisIE (Iranian, NE)31
10/31
31
5/31
31 31 31
10/31
31 31
3/31
YakutsTurkic, Siberia155 155 155 155
137/155
YakutsTurkic, Siberia10
8/10[82]
PopulationLanguage familyn R1bn R1an In E1b1bn E1b1an Jn Gn Nn Tn Ln H

See also

External links

Notes and References

  1. IE=Indo-European
  2. First column gives the amount of total Sample Size studied
  3. Second column gives the Percentage of the particular haplogroup among the Sample Size
  4. Bosch . E. . Calafell . F. . González-Neira . A. . Flaiz . C . Mateu . E. . Scheil . H. G. . Huckenbeck . W. . Efremovska . L. . Mikerezi . I. . Xirotiris . N. . Grasa . C. . Schmidt . H. . Comas . D. . 8 . 2006 . Paternal and maternal lineages in the Balkans show a homogeneous landscape over linguistic barriers, except for the isolated Aromuns . Annals of Human Genetics . 70 . Pt 4 . 459–87 . 10.1111/j.1469-1809.2005.00251.x . 16759179. 23156886 .
  5. Nasidze et al., (2004)Mitochondrial DNA and Y-Chromosome Variation in the Caucasus
  6. Although the chart here says J, this group has mainly only J2
  7. Khar'kov . VN . Stepanov . VA . Medvedeva . OF . Spiridonova . MG . Voevoda . MI . Tadinova . VN . Puzyrev . VP . 2007 . Gene pool differences between Northern and Southern Altaians inferred from the data on Y-chromosomal haplogroups . Genetika . 43 . 5 . 675–87 . 17633562 . 10.1134/S1022795407050110. 566825 .
  8. Semino . O . Santachiara-Benerecetti . AS . Falaschi . F . Cavalli-Sforza . LL . Underhill . PA . 2002 . Ethiopians and Khoisan Share the Deepest Clades of the Human Y-Chromosome Phylogeny . American Journal of Human Genetics . 70 . 1 . 265–8 . 10.1086/338306 . 384897 . 11719903.
  9. Arredi . B . Poloni . ES . Paracchini . S . Zerjal . T . Fathallah . DM . Makrelouf . M . Pascali . VL . Novelletto . A . Tyler-Smith . C . 2004 . A Predominantly Neolithic Origin for Y-Chromosomal DNA Variation in North Africa . American Journal of Human Genetics . 75 . 2 . 338–45 . 10.1086/423147 . 1216069 . 15202071.
  10. Robino . C . Crobu . F . Di Gaetano . C . Bekada . A . Benhamamouch . S . Cerutti . N . Piazza . A . Inturri . S . Torre . C . 2008 . Analysis of Y-chromosomal SNP haplogroups and STR haplotypes in an Algerian population sample . International Journal of Legal Medicine . 122 . 3 . 251–5 . 10.1007/s00414-007-0203-5 . 17909833. 11556974 .
  11. Luis . JR . Rowold . DJ . 532–44 . Regueiro . M . Caeiro . B . Cinnioğlu . C . Roseman . C . Underhill . PA . Cavalli-Sforza . LL . Herrera . RJ . 2004 . The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations . American Journal of Human Genetics . 74 . 3 . 10.1086/382286 . 1182266 . 14973781 . https://web.archive.org/web/20120216123633/http://hpgl.stanford.edu/publications/AJHG_2004_v74_p000-0130.pdf . February 16, 2012 . dead. (Errata)
  12. Cadenas . Alicia M . Zhivotovsky . Lev A . Cavalli-Sforza . Luca L . Underhill . PA . Herrera . RJ . 2008 . Y-chromosome diversity characterizes the Gulf of Oman . European Journal of Human Genetics . 16 . 3 . 374–86 . 17928816 . 10.1038/sj.ejhg.5201934. free .
  13. Abu-Amero . KK . Hellani . A . González . AM . Larruga . JM . Cabrera . VM . Underhill . PA . 2009 . Saudi Arabian Y-Chromosome diversity and its relationship with nearby regions . BMC Genetics . 10 . 59 . 2759955 . 19772609 . 10.1186/1471-2156-10-59 . free .
  14. U.-D. Immela, M. Erhumab, T. Mustafac, M. Kleibera and M. Klintschara, "Population genetic analysis in a Libyan population using the PowerPlex 16 system," International Congress Series Volume 1288, April 2006, Pages 421-423
  15. Michael E. Weale, Levon Yepiskoposyan, Rolf F. Jager, Nelli Hovhannisyan, Armine Khudoyan, Oliver Burbage-Hall, Neil Bradman, Mark G. Thomas, "Armenian Y chromosome haplotypes reveal strong regional structure within a single ethno-national group," Human Genetics (2001) 109 : 659–674.
  16. Nebel . A . Filon . D . Brinkmann . B . Majumder . PP . Faerman . M . Oppenheim . A . 2001 . The Y Chromosome Pool of Jews as Part of the Genetic Landscape of the Middle East . American Journal of Human Genetics . 69 . 5 . 1095–112 . 10.1086/324070 . 1274378 . 11573163 .
  17. Doron M. Behar, Daniel Garrigan, Matthew E. Kaplan et al., "Contrasting patterns of Y chromosome variation in Ashkenazi Jewish and host non-Jewish European populations," Human Genetics (2004) 114 : 354–365.
  18. Yunusbaev 2006
  19. Nasidze2001
  20. grugni2012
  21. Lobov. Y chromosome analysis in subpopulations of Bashkirs from Russia
  22. V. N. Kharkov, V. A. Stepanov, S. P. Feshchenko, S. A. Borinskaya, N. K. Yankovsky, and V. P. Puzyrev, "Frequencies of Y Chromosome Binary Haplogroups in Belarusians," Russian Journal of Genetics, Vol. 41, No. 8, 2005, pp. 928–931.
  23. Behar . Doron M. . Thomas . Mark G. . Skorecki . Karl . Hammer . Michael F. . Bulygina . Ekaterina . Rosengarten . Dror . Jones . Abigail L. . Held . Karen . Moses . Vivian . Goldstein . David . Bradman . Neil . Weale . Michael E. . 3 . 2003 . Multiple Origins of Ashkenazi Levites: Y Chromosome Evidence for Both Near Eastern and European Ancestries . American Journal of Human Genetics . 73 . 4 . 768–79 . 1180600 . 13680527 . 10.1086/378506.
  24. Cruciani . La Fratta . Roberta . Santolamazza . Piero . Sellitto . Daniele . Pascone . Roberto . Moral . Pedro . Watson . Elizabeth . Guida . Valentina . Colomb . Eliane Beraud . Zaharova . Boriana . Lavinha . João . Vona . Giuseppe . Aman . Rashid . Calì . Francesco . Akar . Nejat . Richards . Martin . Torroni . Antonio . Novelletto . Andrea . Scozzari . Rosaria . etal . May 2004 . Phylogeographic Analysis of Haplogroup E3b (E-M215) Y Chromosomes Reveals Multiple Migratory Events Within and Out Of Africa . Am J Hum Genet . 74 . 5 . 1014–1022 . 1181964 . 15042509 . 10.1086/386294.
  25. Rouba . Hassan . Barakat . Abdelhamid . Harich . Nourdin . Reguig . Ahmed . April 2014 . Phylogeography of E1b1b1b-M81 Haplogroup and Analysis of Its Subclades in Morocco . Human Biology . 86 . 2 . 105–113 . 10.3378/027.086.0204 . 0018-7143 . 25397701. 11334895 .
  26. Fadhlaoui-Zid . Karima . Martinez-Cruz . Begona . Khodjet-el-khil . Houssein . Mendizabal . Isabel . Elgaaied . Amel . Comas . David . 1 October 2011 . Genetic Structure of Tunisian Ethnic Groups Revealed by Paternal Lineages . American Journal of Physical Anthropology . 146 . 2 . 271–80 . 10.1002/ajpa.21581 . 21915847.
  27. Qamar . Raheel . etal . May 2002 . Y-Chromosomal DNA Variation in Pakistan . American Journal of Human Genetics . 70 . 1107–1124 . 1107–24 . 10.1086/339929 . 447589 . 11898125.
  28. Sengupta . S . Zhivotovsky . LA . King . R . Mehdi . SQ . Edmonds . CA . Chow . CE . Lin . AA . Mitra . M . Sil . SK . Ramesh . A. . Usha Rani . M.V. . Thakur . Chitra M. . Cavalli-Sforza . L. Luca . Majumder . Partha P. . Underhill . Peter A. . 8 . 2006 . Polarity and Temporality of High-Resolution Y-Chromosome Distributions in India Identify Both Indigenous and Exogenous Expansions and Reveal Minor Genetic Influence of Central Asian Pastoralists . American Journal of Human Genetics . 78 . 2 . 202–21 . 10.1086/499411 . 1380230 . 16400607.
  29. Web site: Bulgarian_Y_Table .
  30. Karachanak . Sena . Fornarino . Simona . Grugni . Viola . Semino . Ornella . Toncheva . Draga . Galabov . Angel . Atanasov . Boris . 2009 . Y-Chromosomal Haplogroups in Bulgarians . Comptes Rendus de l'Académie Bulgare des Sciences . 62 . 3 . 393–400 . 1310-1331 . .
  31. Martinez-Cruz B, Ioana M, Calafell F, Arauna LR, Sanz P, Ionescu R, Boengiu S, Kalaydjieva L, Pamjav H, Makukh H, Plantinga T, van der Meer JW, Comas D, Netea MG . 2012 . Kivisild . 2012PLoSO...741803M . Toomas . Y-chromosome analysis in individuals bearing the Basarab name of the first dynasty of Wallachian kings . PLOS ONE . 7 . 7 . e41803 . 10.1371/journal.pone.0041803 . 3404992 . 22848614. free .
  32. Oleg Balanovsky et al., "Parallel Evolution of Genes and Languages in the Caucasus Region," Molecular Biology and Evolution 2011
  33. 97/445, O. Hassan et al. (2008), Y-Chromosome Variation Among Sudanese: Restricted Gene Flow, Concordance With Language, Geography, and History, American journal of physical anthropology
  34. Peričić M, Lauc LB, Klaric IM, Rootsi S, Janicijevic B, Rudan I, Terzic R, Colak I, Kvesic A, Popovic D, Sijacki A, Behluli I, Dordevic D, Efremovska L, Bajec DD, Stefanovic BD, Villems R, Rudan P . High-resolution phylogenetic analysis of southeastern Europe traces major episodes of paternal gene flow among Slavic populations . . October 2005 . 22 . 10 . 1964–75 . 15944443 . 10.1093/molbev/msi185. free . Haplogroup frequency data in table 1
  35. Barac et al. (2003)
  36. Little genetic differentiation as assessed by uniparental markers in the presence of substantial language variation in peoples of the Cross River region of Nigeria. BMC Evolutionary Biology. 2010. Krishna R. Veeramah. 10. 1 . 92. 20356404. 2867817. 10.1186/1471-2148-10-92. etal . free . 2010BMCEE..10...92V . & Supplementary material
  37. Luca . F. . Di Giacomo . F. . Benincasa . T. . etal . 2007 . Y-Chromosomal Variation in the Czech Republic . American Journal of Physical Anthropology . 132 . 1 . 132–139 . 10.1002/ajpa.20500 . 17078035. 2108/35058 . free .
  38. Elizabeth T Wood, Daryn A Stover, Christopher Ehret et al., "Contrasting patterns of Y chromosome and mtDNA variation in Africa: evidence for sex-biased demographic processes," European Journal of Human Genetics (2005) 13, 867–876. cf. "Appendix A: Y Chromosome Haplotype Frequencies."
  39. http://mbe.oxfordjournals.org/cgi/reprint/19/7/1008
  40. Alexander Varzari, "Population History of the Dniester-Carpathians: Evidence from Alu Insertion and Y-Chromosome Polymorphisms" (2006)
  41. Web site: Distribution of Y-DNA haplogroups.
  42. Web site: Significant genetic differentiation between Poland and Germany follows present-day political borders, as revealed by Y-chromosome analysis . 2011-10-14 . https://web.archive.org/web/20111002034940/http://vetinari.sitesled.com/poland.pdf . 2011-10-02 . dead .
  43. Battaglia . Vincenza . Fornarino . Simona . Al-Zahery . Nadia . Olivieri . Anna . Pala . Maria . Myres . Natalie M . King . Roy J . Rootsi . Siiri . Marjanovic . Damir . Primorac . Dragan . Hadziselimovic . Rifat . Vidovic . Stojko . Drobnic . Katia . Durmishi . Naser . Torroni . Antonio . Santachiara-Benerecetti . A Silvana . Underhill . Peter A . Semino . Ornella . 8 . 24 December 2008 . Y-chromosomal evidence of the cultural diffusion of agriculture in southeast Europe . European Journal of Human Genetics . 17 . 6 . 820–30 . 10.1038/ejhg.2008.249 . 2947100 . 19107149.
  44. Helgason . Agnar . Sigureth Ardóttir . S . Nicholson . J . Sykes . B . Hill . E. W. . Bradley . D. G. . Bosnes . V . Gulcher . J. R. . Ward . R . Stefánsson . K . etal . 2000 . Estimating Scandinavian and Gaelic Ancestry in the Male Settlers of Iceland . Am. J. Hum. Genet. . 67 . 3 . 697–717 . 1287529 . 10931763 . 10.1086/303046 . https://web.archive.org/web/20081011075349/http://www.ajhg.org/AJHG/abstract/S0002-9297(07)63256-X . 2008-10-11 . dead.
  45. King . R. J. . Özcan . S. S. . Carter . T. . Kalfoğlu . E. . Atasoy . S. . Triantaphyllidis . C. . Kouvatsi . A. . Lin . A. A. . Chow . C-E. T. . Zhivotovsky . L. A. . Michalodimitrakis . M. . Underhill . P. A. . 2008 . Differential Y-chromosome Anatolian Influences on the Greek and Cretan Neolithic . Annals of Human Genetics . 72 . 2. 205–214 . 10.1111/j.1469-1809.2007.00414.x . 18269686 . 22406638 .
  46. Hungarian Bukovina DNA Project
  47. M. Regueiro et al. (2006), "Iran: Tricontinental Nexus for Y-Chromosome Driven Migration"
  48. Web site: Archived copy . 2008-03-24 . https://web.archive.org/web/20061212173158/http://www.ebc.ee/tymri00/PhD/2004/SRootsi_thesis.pdf . 2006-12-12 . dead .
  49. Youichi Sato et al. 2014, Overview of genetic variation in the Y chromosome of modern Japanese males J-STAGEトップ/Anthropological Science/122 巻 (2014) 3 号/書誌/全文 The Anthropological Society of Nippon
  50. Жабагин Максат Кизатович 2017,АНАЛИЗ СВЯЗИ ПОЛИМОРФИЗМА Y-ХРОМОСОМЫ И РОДОПЛЕМЕННОЙ СТРУКТУРЫ В КАЗАХСКОЙ ПОПУЛЯЦИИ ДИССЕРТАЦИЯ, ФЕДЕРАЛЬНОЕ ГОСУДАРСТВЕННОЕ БЮДЖЕТНОЕ УЧРЕЖДЕНИЕ, НАУКИ ИНСТИТУТ ОБЩЕЙ ГЕНЕТИКИ им. Н.И. ВАВИЛОВА, РОССИЙСКОЙ АКАДЕМИИ НАУК. Москва
  51. Mirabal S . Regueiro M . Cadenas AM . Cavalli-Sforza . L Luca . Underhill . Peter A . Verbenko . Dmitry A . Limborska . Svetlana A . Herrera . Rene J . 3 . vanc . March 2009 . Y-Chromosome distribution within the geo-linguistic landscape of northwestern Russia . Eur. J. Hum. Genet. . 17 . 10 . 1260–73 . 10.1038/ejhg.2009.6 . 2986641 . 19259129.
  52. R. Spencer Wells et al., "The Eurasian Heartland: A continental perspective on Y-chromosome diversity," Proceedings of the National Academy of Sciences of the United States of America (August 28, 2001)
  53. Haber . Marc . Platt . Daniel . Badro . Danielle . Xue . Yali . El-Sibai . Mirvat . Ashrafian Bonab . Maziar . Youhanna . Sonia . Saade . Stephanie . Soria-Hernanz . David . Royyuru . Ajay . Wells . Spencer . Tyler-Smith . Chris . Zalloua . Pierre . Influences of history, geography, and religion on genetic structure: the Maronites in Lebanon . European Journal of Human Genetics . December 2010 . 19 . 3 . 334–340 . 10.1038/ejhg.2010.177 . 21119711 . 3062011 . free .
  54. Pierre A. Zalloua et al., PY-Chromosomal Diversity in Lebanon Is Structured by Recent Historical Events (2008)
  55. Laitinen . Virpi . Lahermo . Päivi . Sistonen . Pertti . Savontaus . Marja-Liisa . 2002 . Y-Chromosomal Diversity Suggests that Baltic Males Share Common Finno-Ugric-Speaking Forefathers . Human Heredity . 53 . 2 . 68–78 . 10.1159/000057985 . 12037406. 29863635 .
  56. Semino A, Passarino G, Oefner PJ, Lin AA, Arbuzova S, Beckman LE, De Benedictis G, Francalacci P, Kouvatsi A, Limborska S, Marcikiae M, Mika A, Mika B, Primorac D, Santachiara-Benerecetti AS, Cavalli-Sforza LL, Underhill PA . 2000 . The Genetic Legacy of Paleolithic *** sapiens sapiens in Extant Europeans: A Y Chromosome Perspective . Science . 290 . 5494 . 1155–9 . 2000Sci...290.1155S . 10.1126/science.290.5494.1155 . 11073453 . https://web.archive.org/web/20031125151213/http://hpgl.stanford.edu/publications/Science_2000_v290_p1155.pdf . November 25, 2003 . dead.
  57. Web site: Semino et al. (2004). . 2008-03-21 . https://web.archive.org/web/20110709064219/http://hpgl.stanford.edu/publications/AJHG_2004_v74_p1023-1034.pdf . 2011-07-09 . dead .
  58. Malaspina et al. (2003).
  59. Bekada A, Fregel R, Cabrera VM, Larruga JM, Pestano J, Benhamamouch S, González AM . Introducing the Algerian mitochondrial DNA and Y-chromosome profiles into the North African landscape . PLOS ONE . 8 . 2 . e56775 . 2013 . 23431392 . 3576335 . 10.1371/journal.pone.0056775 . 2013PLoSO...856775B . free .
  60. Zegura . S. L. . High-Resolution SNPs and Microsatellite Haplotypes Point to a Single, Recent Entry of Native American Y Chromosomes into the Americas . Molecular Biology and Evolution . 31 October 2003 . 21 . 1 . 164–175 . 10.1093/molbev/msh009 . 14595095 . free .
  61. Nasidze I. et al 2004, Genetic evidence concerning the origins of South and North Ossetians.
  62. Firasat . Sadaf . Khaliq . Shagufta . Mohyuddin . Aisha . Papaioannou . Myrto . Tyler-Smith . Chris . Underhill . Peter A . Ayub . Qasim . 2007 . Y-chromosomal evidence for a limited Greek contribution to the Pathan population of Pakistan . European Journal of Human Genetics . 15 . 1. 121–6 . 10.1038/sj.ejhg.5201726 . 2588664 . 17047675.
  63. B. A. Malyarchuk and M. V. Derenko - Gene Pool Structure of Russian Populations from the European Part of Russia Inferred from the Data on Y Chromosome Haplogroups Distribution // Russian Journal of Genetics, 2008, Vol. 44, No. 2, pp. 187–192.
  64. Oleg Balanovsky, Siiri Rootsi, Andrey Pshenichnov et al., "Two Sources of the Russian Patrilineal Heritage in Their Eurasian Context," American Journal of Human Genetics 82, 236–250, January 2008.
  65. Karlsson . Andreas O . Wallerström . Thomas . Götherström . Anders . Holmlund . Gunilla . 2006 . Y-chromosome diversity in Sweden – A long-time perspective . European Journal of Human Genetics . 14 . 8 . 963–70 . 10.1038/sj.ejhg.5201651 . 16724001. free .
  66. Raito et al.
  67. Rosser et al. (2000)
  68. Web site: Rootsi et al. (2004). . 2008-03-06 . 2020-09-05 . https://web.archive.org/web/20200905162020/http://evolutsioon.ut.ee/publications/Rootsi2004.pdf . dead .
  69. Sanchez . Juan J . Hallenberg . Charlotte . Børsting . Claus . Hernandez . Alexis . Morling . Niels . 2005 . High frequencies of Y chromosome lineages characterized by E3b1, DYS19-11, DYS392-12 in Somali males . European Journal of Human Genetics . 13 . 7 . 856–66 . 10.1038/sj.ejhg.5201390 . 15756297. free .
  70. Zalloua et al. (2008), Identifying Genetic Traces of Historical Expansions: Phoenician Footprints in the Mediterranean, The American Journal of Human Genetics, Volume 83, Issue 5, 633-642 17 November 2008
  71. T. Lappalainen, V. Laitinen, E. Salmela et al., "Migration Waves to the Baltic Sea Region," Annals of Human Genetics (2008).
  72. ArunKumar G, Soria-Hernanz DF, Kavitha VJ, Arun VS, Syama A, Ashokan KS, et al. (2012) Population Differentiation of Southern Indian Male Lineages Correlates with Agricultural Expansions Predating the Caste System. PLoS ONE 7(11): e50269. https://doi.org/10.1371/journal.pone.0050269
  73. Web site: Tambets et al. (2004) . 2008-03-26 . 2019-08-19 . https://web.archive.org/web/20190819054721/http://evolutsioon.ut.ee/publications/Tambets2004PhD.pdf . dead .
  74. http://hpgl.stanford.edu/publications/HG_2004_v114_p127-148.pdf Cinnioglu et al. (2004)
  75. Zerjal . Tatiana . Wells . R. Spencer . Yuldasheva . Nadira . Ruzibakiev . Ruslan . Tyler-Smith . Chris . September 2002 . A Genetic Landscape Reshaped by Recent Events: Y-Chromosomal Insights into Central Asia . American Journal of Human Genetics . 71 . 466–482 . 466–82 . 10.1086/342096 . 419996 . 12145751.
  76. Karafet . Tatiana . Xu . Liping . Du . Ruofu . etal . September 2001 . Paternal Population History of East Asia: Sources, Patterns, and Microevolutionary Processes . American Journal of Human Genetics . 69 . 615–628 . 615–28 . 10.1086/323299 . 1235490 . 11481588.
  77. Michael F. Hammer, Tatiana M. Karafet, Hwayong Park, Keiichi Omoto, Shinji Harihara, Mark Stoneking and Satoshi Horai, "Dual origins of the Japanese: common ground for hunter-gatherer and farmer Y chromosomes," Journal of Human Genetics Volume 51, Number 1 / January, 2006.
  78. Ruixia Zhou, Daqun Yang, Hua Zhang, Weiping Yu, Lizhe An, Xilong Wang, Hong Li, Jiujin Xu, and Xiaodong Xie, "Origin and evolution of two Yugur sub-clans in Northwest China: a case study in paternal genetic landscape," Annals of Human Biology (2008), 35:2, 198 — 211.
  79. Shou et al (2010), Y-chromosome distributions among populations in Northwest China identify significant contribution from Central Asian pastoralists and lesser influence of western Eurasians, Journal of Human Genetics 55, 314-322 (May 2010)
  80. Yali Xue, Tatiana Zerjal, Weidong Bao, Suling Zhu, Qunfang Shu, Jiujin Xu, Ruofu Du, Songbin ***, Pu Li, Matthew E. Hurles, Huanming Yang, Chris Tyler-Smith, "Male demography in East Asia: a north-south contrast in human population expansion times," Genetics 2006.
  81. http://www.pnas.org/cgi/reprint/98/9/5078.pdf
  82. Web site: Y-chromosome haplogroup N dispersals from south Siberia to Europe. Derenko et al. . 2008-03-05 . https://web.archive.org/web/20140606011243/https://www.familytreedna.com/pdf/Derenko2007.pdf . 2014-06-06 . dead .