Finno-Ugric languages explained

Finno-Ugric
Also Known As:Finno-Ugrian
Region:Eastern, Central and Northern Europe, North Asia
Familycolor:Uralic
Child1:Finno-Permic (traditional grouping)
Child2:Ugric (traditional grouping)
Iso2:fiu
Iso5:fiu
Glotto:none
Map:Finno-Ugric Languages.png
Mapcaption:The Finno-Ugric languages

Finno-Ugric [1] is a traditional grouping of all languages in the Uralic language family except the Samoyedic languages. Its formerly commonly accepted status as a subfamily of Uralic is based on criteria formulated in the 19th century and is criticized by some contemporary linguists such as Tapani Salminen and Ante Aikio.[2] [3] The three most spoken Uralic languages, Hungarian, Finnish, and Estonian, are all included in Finno-Ugric.

The term Finno-Ugric, which originally referred to the entire family, is sometimes used as a synonym for the term Uralic, which includes the Samoyedic languages, as commonly happens when a language family is expanded with further discoveries.[4] [5] [6] Before the 20th century, the language family might be referred to as Finnish, Ugric, Finno-Hungarian or with a variety of other names.[7] The name Finno-Ugric came into general use in the late 19th or early 20th century.[8] [9]

Status

The validity of Finno-Ugric as a phylogenic grouping is under challenge,[10] [11] with some linguists maintaining that the Finno-Permic languages are as distinct from the Ugric languages as they are from the Samoyedic languages spoken in Siberia, or even that none of the Finno-Ugric, Finno-Permic, or Ugric branches has been established. Received opinion is that the easternmost (and last discovered) Samoyed had separated first and the branching into Ugric and Finno-Permic took place later, but this reconstruction does not have strong support in the linguistic data.

Origins

Attempts at reconstructing a Proto-Finno-Ugric proto-language, a common ancestor of all Uralic languages except for the Samoyedic languages, are largely indistinguishable from Proto-Uralic, suggesting that Finno-Ugric might not be a historical grouping but a geographical one, with Samoyedic being distinct by lexical borrowing rather than actually being historically divergent. It has been proposed that the area in which Proto-Finno-Ugric was spoken reached between the Baltic Sea and the Ural Mountains.[12]

Traditionally, the main set of evidence for the genetic proposal of Proto-Finno-Ugric has come from vocabulary. A large amount of vocabulary (e.g. the numerals "one", "three", "four" and "six"; the body-part terms "hand", "head") is only reconstructed up to the Proto-Finno-Ugric level, and only words with a Samoyedic equivalent have been reconstructed for Proto-Uralic. That methodology has been criticised, as no coherent explanation other than inheritance has been presented for the origin of most of the Finno-Ugric vocabulary (though a small number has been explained as old loanwords from Proto-Indo-European or its immediate successors).

The Samoyedic group has undergone a longer period of independent development, and its divergent vocabulary could be caused by mechanisms of replacement such as language contact. (The Finno-Ugric group is usually dated to approximately 4,000 years ago, the Samoyedic a little over 2,000.) Proponents of the traditional binary division note, however, that the invocation of extensive contact influence on vocabulary is at odds with the grammatical conservatism of Samoyedic.

The consonant (voiceless postalveolar fricative, pronounced as /[ʃ]/) has not been conclusively shown to occur in the traditional Proto-Uralic lexicon, but it is attested in some of the Proto-Finno-Ugric material. Another feature attested in the Finno-Ugric vocabulary is that *i now behaves as a neutral vowel with respect to front-back vowel harmony, and thus there are roots such as *niwa- "to remove the hair from hides".[13]

Regular sound changes proposed for this stage are few and remain open to interpretation. Sammallahti (1988)[13] proposes five, following Janhunen's (1981) reconstruction of Proto-Finno-Permic:

Sammallahti (1988) further reconstructs sound changes *oo, *ee*a, (merging with original *a, ) for the development from Proto-Finno-Ugric to Proto-Ugric. Similar sound laws are required for other languages as well. Thus, the origin and raising of long vowels may actually belong at a later stage,[14] and the development of these words from Proto-Uralic to Proto-Ugric can be summarized as simple loss of *x (if it existed in the first place at all; vowel length only surfaces consistently in the Baltic-Finnic languages.) The proposed raising of *o has been alternatively interpreted instead as a lowering *u*o in Samoyedic (PU *lumi*loməProto-Samoyedic *jom).[14]

Janhunen (2007, 2009) notes a number of derivational innovations in Finno-Ugric, including *ńoma "hare" → *ńoma-la, (vs. Samoyedic *ńomå), *pexli "side" → *peel-ka*pelka "thumb", though involving Proto-Uralic derivational elements.

Structural features

The Finno-Ugric group is not typologically distinct from Uralic as a whole: the most widespread structural features among the group all extend to the Samoyedic languages as well.

Classification models

Modern linguistic research has shown that Volgaic languages is a geographical classification rather than a linguistic one, because the Mordvinic languages are more closely related to the Finno-Samic languages than the Mari languages.

The relation of the Finno-Permic and the Ugric groups is adjudged remote by some scholars. On the other hand, with a projected time depth of only 3,000 to 4,000 years, the traditionally accepted Finno-Ugric grouping would be far younger than many major families such as Indo-European or Semitic, and would be about the same age as, for instance, the Eastern subfamily of Nilotic. But the grouping is far from transparent or securely established. The absence of early records is a major obstacle. As for the Finno-Ugric Urheimat, most of what has been said about it is speculation.

Some linguists criticizing the Finno-Ugric genetic proposal, especially Angela Marcantonio,[15] also question the validity of the entire Uralic family, instead proposing a Ural–Altaic hypothesis, within which they believe Finno-Permic may be as distant from Ugric as from Turkic. However, this approach has been rejected by nearly all other specialists in Uralic linguistics.[16] [17] [18] [19] [20] [21]

Common vocabulary

Loanwords

One argument in favor of the Finno-Ugric grouping has come from loanwords. Several loans from the Indo-European languages are present in most or all of the Finno-Ugric languages, while being absent from Samoyedic.

According to Häkkinen (1983) the alleged Proto-Finno-Ugric loanwords are disproportionally well-represented in Hungarian and the Permic languages, and disproportionally poorly represented in the Ob-Ugric languages; hence it is possible that such words have been acquired by the languages only after the initial dissolution of the Uralic family into individual dialects, and that the scarcity of loanwords in Samoyedic results from its peripheric location.[22]

Numbers

The number systems among the Finno-Ugric languages are particularly distinct from the Samoyedic languages: only the numerals "2", "5", and "7" have cognates in Samoyedic, while also the numerals, "1", "3", "4", "6", "10" are shared by all or most Finno-Ugric languages.

Below are the numbers 1 to 10 in several Finno-Ugric languages. Forms in italic do not descend from the reconstructed forms.

NumberBaltic FinnicSámiMordvinicMariPermicUgricProto-
Finno-
Ugric
FinnishEstonianVõroLivonianNorthern SámiInari SámiErzyaMokshaMeadow MariKomi-ZyrianMansiKhantyHungarian
1Finnish: italic=no|yksi
gen. Finnish: italic=no|yhden, part. Finnish: italic=no|yhtä
Estonian: italic=no|üks
gen. Estonian: italic=no|ühe, part. Estonian: italic=no|üht(e)
italic=no|ütśitalic=no|ikšNorthern Sami: italic=no|oktaInari Sami: italic=no|ohtâik/ikyt/iktyt/ikteöťiakwaiHungarian: egy[23] Finno-Ugrian languages: italic=no|*ükte
2kaksi
gen. kahden, part. kahta
kaks
gen. kahe, part. kaht(e)
katśkakšguoktekyeh´tikavtokaftəkok/kokyt/koktytkykkitigkatkettő/két
  • kakta
3kolmekolmkolmkuolmgolbmakulmâkolmokolməkum/kumytkuimxūrumxołəmhárom, harm-
  • kolme
4neljänelinellinēļanjealljeneljińiľeńiľänyl/nylytńoľńilańałnégy
  • neljä
5viisiviisviiśvīžvihttavittâveƭeveťävič/vizytvitatwetöt
  • viite
6kuusikuuskuuśkūžguhttakuttâkotokotəkud/kudytkvajtxōtxothat
  • kuute
7seitsemänseitsesäidseseisčiežačiččâmśiśemśiśämšym/šymytśiźimsātłapəthétśäjććemä
8kahdeksankaheksakatõsakōdõksgávccikäävcikavksokafksəkandaš/kandašekökjamysńololowniwłnyolcN/A
9yhdeksänüheksaütesäīdõksovccioovcevejkseveçksəindeš/indešeökmysontolowjarťaŋkilencN/A
10kymmenenkümmekümmekimlogilovekemeńkeməńludaslowjaŋtízluka/kümmen

The number '2' descends in Ugric from a front-vocalic variant *kektä.

The numbers '9' and '8' in Finnic through Mari are considered to be derived from the numbers '1' and '2' as '10–1' and '10–2'. One reconstruction is *yk+teksa and *kak+teksa, respectively, where *teksa cf. deka is an Indo-European loan; the difference between /t/ and /d/ is not phonemic, unlike in Indo-European. Another analysis is *ykt-e-ksa, *kakt-e-ksa, with *e being the negative verb.

Finno-Ugric Swadesh lists

100-word Swadesh lists for certain Finno-Ugric languages can be compared and contrasted at the Rosetta Project website:Finnish, Estonian, Hungarian, and Erzya.

Speakers

The four largest ethnic groups that speak Finno-Ugric languages are the Hungarians (14.5 million), Finns (6.5 million), Estonians (1.1 million), and Mordvins (0.85 million). Majorities of three (the Hungarians, Finns, and Estonians) inhabit their respective nation states in Europe, i.e. Hungary, Finland, and Estonia, while a large minority of Mordvins inhabit the federal Mordovian Republic within Russia (Russian Federation).[24]

The indigenous area of the Sámi people is known as Sápmi and it consists of the northern parts of the Fennoscandian Peninsula. Some other peoples that speak Finno-Ugric languages have been assigned autonomous republics within Russia. These are the Karelians (Republic of Karelia), Komi (Komi Republic), Udmurts (Udmurt Republic) and Mari (Mari El Republic). The Khanty-Mansi Autonomous Okrug was set up for the Khanty and Mansi of Russia. A once-autonomous Komi-Permyak Okrug was set up for a region of high Komi habitation outside the Komi Republic.

Some of the ethnicities speaking Finno-Ugric languages are:

International Finno-Ugric societies

In the Finno-Ugric countries of Finland, Estonia and Hungary that find themselves surrounded by speakers of unrelated tongues, language origins and language history have long been relevant to national identity. In 1992, the 1st World Congress of Finno-Ugric Peoples was organized in Syktyvkar in the Komi Republic in Russia, the 2nd World Congress in 1996 in Budapest in Hungary, the 3rd Congress in 2000 in Helsinki in Finland, the 4th Congress in 2004 in Tallinn in Estonia, the 5th Congress in 2008 in Khanty-Mansiysk in Russia, the 6th Congress in 2012 in Siófok in Hungary,[25] [26] [27] [28] the 7th Congress in 2016 in Lahti in Finland,[29] and the 8th Congress in 2021 in Tartu in Estonia.[30] The members of the Finno-Ugric Peoples' Consultative Committee include: the Erzyas, Estonians, Finns, Hungarians, Ingrian Finns, Ingrians, Karelians, Khants, Komis, Mansis, Maris, Mokshas, Nenetses, Permian Komis, Saamis, Tver Karelians, Udmurts, Vepsians; Observers: Livonians, Setos.[31] [32]

In 2007, the 1st Festival of the Finno-Ugric Peoples was hosted by President Vladimir Putin of Russia, and visited by Finnish President, Tarja Halonen, and Hungarian Prime Minister, Ferenc Gyurcsány.[33] [34]

The International Finno-Ugric Students' Conference (IFUSCO) is organised annually by students of Finno-Ugric languages to bring together people from all over the world who are interested in the languages and cultures. The first conference was held in 1984 in Göttingen in Germany. IFUSCO features presentations and workshops on topics such as linguistics, ethnography, history and more.[35] [36]

The International Congress for Finno-Ugric Studies is the largest scientific meeting of scientists studying the culture and languages of Finno-Ugric peoples, held every five years.[37] [38] The first congress was organized in 1960 in Budapest, the last congress took place in 2022 in Vienna,[39] the next congress is planned to be held in Tartu, Estonia in 2025.[40]

Population genetics

The linguistic reconstruction of the Finno-Ugric language family has led to the postulation that the ancient Proto-Finno-Ugric people were ethnically related, and that even the modern Finno-Ugric-speaking peoples are ethnically related.[41] Such hypotheses are based on the assumption that heredity can be traced through linguistic relatedness,[42] although it must be kept in mind that language shift and ethnic admixture, a relatively frequent and common occurrence both in recorded history and most likely also in prehistory, confuses the picture and there is no straightforward relationship, if at all, between linguistic and genetic affiliation. Still, the premise that the speakers of the ancient proto-language were ethnically homogeneous is generally accepted.

Modern genetic studies have shown that the Y-chromosome haplogroup N3, and sometimes N2, is almost specific though certainly not restricted to Uralic- or Finno-Ugric-speaking populations, especially as high frequency or primary paternal haplogroup.[43] [44] These haplogroups branched from haplogroup N, which probably spread north, then west and east from Northern China about 12,000–14,000 years before present from father haplogroup NO (haplogroup O being the most common Y-chromosome haplogroup in Southeast Asia).

A study of the Finno-Ugric-speaking peoples of northern Eurasia (i.e., excluding the Hungarians), carried out between 2002 and 2008 in the Department of Forensic Medicine at the University of Helsinki, showed that the Finno-Ugric-speaking populations do not retain genetic evidence of a common founder. Most possess an amalgamation of West and East Eurasian gene pools that may have been present in central Asia, with subsequent genetic drift and recurrent founder effects among speakers of various branches of Finno-Ugric. Not all branches show evidence of a single founder effect. North Eurasian Finno-Ugric-speaking populations were found to be genetically a heterogeneous group showing lower haplotype diversities compared to more southern populations. North Eurasian Finno-Ugric-speaking populations possess unique genetic features due to complex genetic changes shaped by molecular and population genetics and adaptation to the areas of Boreal and Arctic North Eurasia.[45]

See also

Further reading

External links

Notes and References

  1. Collins English Dictionary – Complete & Unabridged 11th Edition. Retrieved 4 September 2012 from website: http://www.collinsdictionary.com/dictionary/english/Finno-Ugric
  2. Tapani Salminen, "The rise of the Finno-Ugric language family." In Carpelan, Parpola, & Koskikallio (eds.), Early contacts between Uralic and Indo-European: linguistic and archaeological considerations. Mémoires de la Société Finno-Ougrienne 242; Helsinki 2001. 385–396.http://www.helsinki.fi/~tasalmin/tvarminne.html
  3. Encyclopedia: Aikio . Ante . Proto-Uralic . Bakró-Nagy . Marianne . Marianne Bakró-Nagy. Laakso . Johanna . Skribnik . Elena . Oxford Guide to the Uralic Languages . 2019 . Oxford University Press . Oxford, UK . 3–4 . 18 May 2020 . 10 December 2021 . https://web.archive.org/web/20211210230034/https://www.academia.edu/40193033 . live .
  4. Book: Tommola, Hannu . Mood in the Languages of Europe . John Benjamins Publishing Company . 2010 . 978-90-272-0587-2 . 155 . Finnish among the Finno-Ugrian languages . https://books.google.com/books?id=o3L8oKcbZtoC&pg=PA511.
  5. 24 March 2022 . Introduction . The Oxford Guide to the Uralic Languages . liv–lvi . 10.1093/oso/9780198767664.002.0013. 978-0-19-876766-4 .
  6. Bakró-Nagy . Marianne . Marianne Bakró-Nagy. 2012 . The Uralic Languages . Revue belge de Philologie et d'Histoire . 90 . 3 . 1001–1027 . 10.3406/rbph.2012.8272.
  7. Web site: Sommer . Łukasz . 1 January 2023 . Conceptualizing language kinship or How Finnocentric is Finno-Ugricity? . Academia.edu.
  8. Book: Guyla. Décsy. Otto Harrassowitz . Wiesbaden . Einführung in die finnisch-ugrische Sprachwissenschaft . 1965. 1. de.
  9. Book: 125 éves a Budapesti Finnugor Tanszék: jubileumi kötet . 1998 . ELTE, BFT . 978-963-463-213-9 . Domokos . Péter . Urálisztikai tanulmányok . Budapest . Csepregi. Márta. 56–62. Péter. Hajdú. hu. A magyar–ugor vs. altaji összehasonlítótól az uráli nyelvészetig (via finnugor). From the Hungarian-Ugric vs. Altaic comparative study to Uralic linguistics (via Finno-Ugric).
  10. http://www.helsinki.fi/~tasalmin/kuzn.html Salminen, Tapani (2002): Problems in the taxonomy of the Uralic languages in the light of modern comparative studies
  11. Encyclopedia: Aikio . Ante . Ante Aikio . Chapter 1: Proto-Uralic . Bakró-Nagy . Marianne . Laakso . Johanna . Skribnik . Elena . The Oxford Guide to the Uralic Languages . 2019 . . Oxford, UK . 3ff. . 9780198767664 . 18 May 2020 . 10 December 2021 . https://web.archive.org/web/20211210230034/https://www.academia.edu/40193033 . live .
  12. Book: Campbell, Lyle. Historical linguistics: an introduction. 2004. MIT Press. 978-0-262-53267-9. 405 .
  13. Book: Sammallahti, Pekka. The Uralic languages – Description, history and foreign influences. https://books.google.com/books?id=TM2NQ78dP2wC&pg=PA492. 1988. BRILL. 978-90-04-07741-6. 478–554. Historical Phonology of the Uralic languages. Denis, Sinor .
  14. Häkkinen, Jaakko 2009: Kantauralin ajoitus ja paikannus: perustelut puntarissa. – Suomalais-Ugrilaisen Seuran Aikakauskirja 92. http://www.sgr.fi/susa/92/hakkinen.pdf
  15. Book: Marcantonio, Angela. The Uralic Language Family: Facts, Myths and Statistics. Blackwell. 2002. 978-0-631-23170-7. Publications of the Philological Society. 35. Oxford. 803186861.
  16. Aikio . Ante . 2003 . Angela Marcantonio, The Uralic Language Family: Facts, Myths and Statistics . Word . 54 . 3 . 401–412 . Book review . 10.1080/00437956.2003.11432539 . free .
  17. Bakro-Nagy . Marianne . The Uralic Language Family. Facts, Myths and Statistics . 2005 . Lingua . 115 . 7 . 1053–1062 . Book review . 10.1016/j.lingua.2004.01.008 .
  18. Stefan. Georg. Marcantonio, Angela: The Uralic Language Family. Facts, Myths and Statistics. Book review. 2004. Finnisch-Ugrische Mitteilungen. 26/27. 155–168.
  19. Petri. Kallio. The Uralic Language Family: Facts, Myths and Statistics. Angela Marcantonio.. Book review. 2004. Anthropological Linguistics. 46. 486–490.
  20. Ulla-Maija. Kulonen. Myyttejä uralistiikasta. Angela Marcantonio. The Uralic Language Family: Facts, Myths and Statistics. Book review. 2004. Virittäjä. 2/2004. 314–320.
  21. Laakso . Johanna . 2004 . Sprachwissenschaftliche Spiegelfechterei (Angela Marcantonio: The Uralic language family. Facts, myths and statistics) . Finnisch-ugrische Forschungen . 58 . 296–307 . Book review . de .
  22. Kaisa. Häkkinen. 1983. Suomen kielen vanhimmasta sanastosta ja sen tutkimisesta. fi. PhD. Turun yliopisto. 951-642-445-7.
  23. According to Book: Zaich, Gábor. Etimológiai szótár. 2006. hu. 978-963-7094-01-9. 167 . Tinta., the Hungarian word for "one" is an internal development, i.e. it is not related to the Proto-Finno-Ugric Finno-Ugrian languages: *ükte
  24. Iurchenkov . Valerii . March 2001 . The Mordvins: Dilemmas of Mobilization in a Biethnic Community . Nationalities Papers . en . 29 . 1 . 85–95 . 10.1080/00905990120036394 . 0090-5992.
  25. Web site: 7th World Congress of the Finno-Ugric Peoples . World Congress of the Finno-Ugric Peoples . 15 July 2016 . https://web.archive.org/web/20190317164710/http://lahti2016.fucongress.org/en . 17 March 2019 . dead.
  26. Web site: Statutes of the Consultative Committee of Finno-Ugrian peoples . 13 July 2016 . Finno-Ugric Peoples' Consultative Committee . 31 July 2020 . https://web.archive.org/web/20200731164926/http://www.suri.ee/doc/statutes.html . live .
  27. Web site: The Congress of the Finno-Ugric Peoples . Russia . 15 July 2016 . 6 August 2020 . https://web.archive.org/web/20200806125723/https://rkomi.ru/en/print/razdelpseudo/337/ . dead .
  28. Web site: Fenno-Ugria . Estonia . 15 July 2016 . 19 September 2016 . https://web.archive.org/web/20160919044725/http://jabadaba.eki.ee/index.php?id=10973 . live .
  29. Web site: The VII (7th) World Congress of Finno-Ugric Peoples . Fenno-Ugria . 17 August 2021 . 17 August 2021 . https://web.archive.org/web/20210817233025/https://fennougria.ee/en/representative-bodies/worldcongresses/seventh/ . live .
  30. Web site: The VIII(8th) World Congress of Finno-Ugric Peoples . Fenno-Ugria . 17 August 2021 . 11 July 2021 . https://web.archive.org/web/20210711202852/https://fennougria.ee/en/representative-bodies/worldcongresses/eighth/ . live .
  31. Web site: Finno-Ugric Peoples' Consultative Committee, Members . 15 July 2016 . Finno-Ugric Peoples' Consultative Committee . World Congresses of the Finno-Ugric Peoples . 2 February 2017 . https://web.archive.org/web/20170202002051/http://www.suri.ee/coco.html . live .
  32. Web site: Suomalais-Ugrilainen Seura (in Finnish) . Finno-Ugrian Society (in English) . 15 July 2016 . 3 March 2018 . https://web.archive.org/web/20180303004401/http://www.sgr.fi/english/index.html . live .
  33. Web site: International Festival of the Finno-Ugric Peoples . Press Release from the Kremlin in Russia . 19 July 2007 . 15 July 2016 . 31 July 2020 . https://web.archive.org/web/20200731151016/http://en.kremlin.ru/events/president/transcripts/24422 . live .
  34. News: Press Release from the Kremlin in Russia . Press Statements by President Vladimir Putin, leaders of Finland and Hungary . 19 July 2007 . 15 July 2016 . 31 July 2020 . https://web.archive.org/web/20200731175452/http://en.kremlin.ru/events/president/transcripts/24418 . live .
  35. Web site: IFUSCO comes back to "fatherland" from Syktyvkar . dead . https://web.archive.org/web/20180616223234/http://finugor.ru/en/node/41040 . 16 June 2018.
  36. Web site: IFUSCO XXXVII Prague 2022 FAQ . ifusco2022.ff.cuni.cz/faq/ . 28 May 2023 . 28 May 2023 . https://web.archive.org/web/20230528190840/https://ifusco2022.ff.cuni.cz/faq/ . live .
  37. Gulyás . Nikolett F. . Janurik . Boglárka . Mus . Nikolett . Tánczos . Orsolya . 2011 . The 11th International Congress for Finno-Ugric Studies: Finno-Ugric Peoples and Languages in the 21st Century . Finno-Ugric Languages and Linguistics . 1 . 1–2.
  38. Georgieva . Ekaterina . Mus . Nikolett . 2015 . The 12th International Congress for Finno-Ugric Studies . Finno-Ugric Languages and Linguistics . 4 . 1–2.
  39. Web site: 13th International Congress for Finno-Ugric Studies . The official site of Estonian Non-Profit Organisation Fenno-Ugria.
  40. Web site: Congressus XIV Internationalis Fenno-Ugristarum. August 2025, Tartu. . 12 March 2024 . The official site of 14th International Congress for Finno-Ugric Studies.
  41. Book: Sámuel Gyarmathi. Grammatical Proof of the Affinity of the Hungarian Language with Languages of Fennic Origin: (Gottingen Dieterich, 1799). 1983. John Benjamins Publishing. 978-90-272-0976-4 .
  42. Web site: Where do Finnish come from?. 19 November 2014. 22 March 2010. 17 November 2010. https://web.archive.org/web/20101117094021/http://finland.fi/public/default.aspx?contentid=160056. live.
  43. European Journal of Human Genetics – Abstract of article: A counter-clockwise northern route of the Y-chromosome haplogroup N from Southeast Asia towards Europe. European Journal of Human Genetics. February 2007. 15. 2. 204–211. 10.1038/sj.ejhg.5201748. Rootsi. Siiri. Zhivotovsky. L. A.. Baldovic. M.. Kayser. M.. Kutuev. I. A.. Khusainova. R.. Bermisheva. M. A.. Gubina. M.. Fedorova. S. A.. Ilumäe. A. M.. Khusnutdinova. E. K.. Voevoda. M. I.. Osipova. L. P.. Stoneking. M.. Lin. A. A.. Ferak. V.. Parik. J.. Kivisild. T.. Underhill. P. A.. Villems. R.. 17149388. 19265287. free.
  44. Web site: Journals Home.
  45. Web site: Pimenoff, Ville. Living on the edge : Population genetics of Finno-Ugric-speaking humans in North Eurasia. University of Helsinki, Finland. 2008. 12 April 2019. 19 March 2021. https://web.archive.org/web/20210319005220/https://helda.helsinki.fi/handle/10138/20330. live. PhD thesis