List of threatened rays explained

See also: Threatened sharks.

Threatened rays are those vulnerable to endangerment (extinction) in the near future. The International Union for Conservation of Nature (IUCN) ranks threatened species in three categories:^[1]

Vulnerable species

Endangered species

Critically endangered species

The term threatened strictly refers to these three categories (critically endangered, endangered and vulnerable), while vulnerable is used to refer to the least at risk of these categories. The terms can be used somewhat interchangeably, as all vulnerable species are threatened, all endangered species are vulnerable and threatened, and all critically endangered species are endangered, vulnerable and threatened. Threatened species are also referred to as a red-listed species, as they are listed in the IUCN Red List of Threatened Species.^[2]

Together rays and sharks make up the class of modern cartilaginous fishes. Modern fish are either cartilaginous or bony. Cartilaginous fishes have skeletons made of cartilage while bony fishes have skeletons made of bone. Because rays and sharks are closely related, they are often studied together. In 2010 a global IUCN study of vertebrates found that of 1,044 cartilaginous (ray and shark) species examined, 345 or 33% were threatened with extinction.^{[3] [4]}

There are four orders of rays: stingrays, skates, electric rays and sawfishes. Like sharks, rays are relatively long living and thrive in stable populations. They are K-strategists which grow slowly, mature late sexually and produce few offspring. They cannot recover as rapidly as many faster growing fish can if their populations are depleted.^[5] As with sharks, rays are increasingly becoming vulnerable because of commercial and recreational fishing pressures, the impact of non-ray fisheries on the seabed and ray prey species, and other habitat alterations such as damage and loss from coastal development and marine pollution.^[6] Most particularly, the continuing decline of threatened rays and sharks is the consequence of unregulated fishing,^[7] as illustrated by a recent international survey which listed only 38 species of skates and rays still subsisting in the highly impacted Mediterranean Sea.^[8]

Manta rays are largest rays in the world, with wingspans reaching 7 metres.^[9] They have one of the highest brain-to-body mass ratios of all fish.^[10] Manta populations suffer when they are caught as bycatch by fishermen fishing for other species, but fisheries which target manta rays are even more harmful. Manta rays use their gills to filter plankton from the sea. Demand for their dried gill rakers, cartilaginous structures protecting the gills, has been growing in traditional Chinese medicine practices.^[11] The market is "bogus" since dried manta gills have never been used historically in Chinese medicine, and there is no evidence that the gills have any medicinal value.^{[12] [13]} The flesh is edible and is consumed in some countries, but is tough and unattractive compared to other fish. To fill the growing demand in Asia for gill rakers, targeted fisheries have developed in other parts of the world, including Sri Lanka, Indonesia, West Africa and Central and South America.^[14] Each year, thousands of manta rays, primarily the giant manta ray, are being caught and killed purely for their gill rakers. A fisheries study in Sri Lanka estimated that over a thousand of these were being sold in the country's fish markets each year.^[15]

In 2011, manta rays became strictly protected in international waters thanks to their recent inclusion in the Convention on Migratory Species of Wild Animals. The CMS is an international treaty organization concerned with conserving migratory species and habitats on a global scale. Although individual nations were already protecting manta rays, the fish often migrate through unregulated waters, putting them at increased risk from overfishing.^[16] In 2013, the Convention on International Trade in Endangered Species (CITES) listed both species of manta rays as CITES Appendix II species. This means that the international trade of manta rays will now be monitored and regulated.^{[17] [18]}

Sawfish are a less well known family of rays which have a long rostrum resembling a saw. Some species can reach 7m (23feet) in length. All species of sawfish are either endangered or critically endangered as a result of habitat destruction and overfishing. Their young stay close to shore, and are particularly affected by coastal developments.^[19] Because their rostrum is easily entangled, sawfishes can easily become bycatch in fishing nets. They are also exploited for the novelty value of their rostrum, their fins are eaten as a delicacy in China, and their liver oil used as a food supplement. While arguing for a global ban on international commerce in 2007, a representative from the National Museums of Kenya stated, "Only the meat is consumed locally; and artisanal fishermen can retire after catching one sawfish due to the high value of a single rostrum, up to $1,450." In 2013 CITES uplisted the largetooth sawfish to Appendix I. This is CITES highest protection level, and means that all international trade of the species is banned.^{[20] [21]}

According to a 2021 study published in the journal Nature, relative fishing pressure in the oceans has increased by a factor of 18 since 1970.^[22] This overfishing has resulted in the number of oceanic sharks and rays declining globally by 71%, and has increased the global extinction risk to the point where three-quarters of these species are now threatened with extinction. Precautionary science-based catch limits and strict prohibitions are now needed urgently if population collapse is to be avoided,^{[23] [24]} if the disruption of ecological functions is to be averted,^[25] and if a start is to be made on rebuilding global fisheries.^[26]

List

Order	Image	Scientific name	Common name	Population trend	IUCN status	Fish Base	FAO	Comment
Stingray		Aetobatus flagellum	Longheaded eagle ray	align=center	decreasing	Endangered^[27]
Stingray		Aetomylaeus maculatus	Mottled eagle ray	align=center	decreasing	Endangered
Stingray		Aetomylaeus nichofii	Banded eagle ray	align=center	decreasing	Vulnerable^[28]
Stingray		Aetomylaeus vespertilio	Ornate eagle ray	align=center	decreasing	Endangered^[29]
Skate		Amblyraja radiata	Thorny skate	align=center	decreasing	Vulnerable^[30]
Sawfish		Anoxypristis cuspidata	Knifetooth sawfish	align=center	decreasing	Endangered^[31]
Stingray		Aptychotrema timorensis	Spotted shovelnose ray	align=center	unknown	Vulnerable^[32]
Skate		Atlantoraja castelnaui	Spotback skate	align=center	decreasing	Endangered^[33]
Skate		Atlantoraja cyclophora	Eyespot skate	align=center	decreasing	Vulnerable^[34]
Skate		Atlantoraja platana	La Plata skate	align=center	decreasing	Vulnerable^[35]
Skate		Bathyraja griseocauda	Graytail skate	align=center	decreasing	Endangered^[36]
Electric ray		Benthobatis kreffti	Brazilian blind electric ray	align=center	unknown	Vulnerable^[37]
Stingray		Dasyatis colarensis	Colares stingray	align=center	unknown	Vulnerable^[38]
Stingray		Dasyatis fluviorum	Estuary stingaree	align=center	decreasing	Vulnerable^[39]
Stingray		Dasyatis garouaensis	Smooth freshwater stingray	align=center	decreasing	Vulnerable^[40]			freshwater
Stingray		Dasyatis laosensis	Mekong freshwater stingray	align=center	decreasing	Endangered^[41]			freshwater
Stingray		Dasyatis margarita	Daisy stingray	align=center	decreasing	Endangered^[42]
Stingray		Dasyatis ukpam	Daisy stingray	align=center	unknown	Endangered^[43]			freshwater
Electric ray		Diplobatis colombiensis	Colombian electric ray	align=center	unknown	Vulnerable^[44]
Electric ray		Diplobatis guamachensis	Brownband numbfish	align=center	unknown	Vulnerable^[45]
Electric ray		Diplobatis ommata	Ocellated electric ray	align=center	unknown	Vulnerable^[46]
Electric ray		Diplobatis pictus	Variegated electric ray	align=center	unknown	Vulnerable^[47]
Skate		Dipturus australis	Sydney skate	align=center	unknown	Vulnerable^[48]
Skate	align=center		Dipturus batis	Blue skate	align=center	decreasing	Critically endangered^[49]
Skate		Dipturus crosnieri	Madagascar skate	align=center	unknown	Vulnerable^[50]
Skate		Dipturus laevis	Barndoor skate	align=center	stable	Endangered^[51]
Skate		Dipturus mennii	South Brazilian skate	align=center	unknown	Vulnerable^[52]
Skate		Dipturus trachydermus	Roughskin skate	align=center	decreasing	Vulnerable^[53]
Skate		Glaucostegus granulatus	Sharpnose guitarfish	align=center	decreasing	Vulnerable^[54]
Skate		Glaucostegus thouin	Clubnose guitarfish	align=center	unknown	Vulnerable^[55]
Skate		Glaucostegus typus	Common shovelnose ray	align=center	decreasing	Vulnerable^[56]
Skate		Gurgesiella dorsalifera	Onefin skate	align=center	decreasing	Vulnerable^[57]
Stingray		Gymnura altavela	Spiny butterfly ray	align=center	decreasing	Vulnerable^[58]
Stingray		Gymnura zonura	Zonetail butterfly ray	align=center	decreasing	Vulnerable^[59]
Electric ray		Heteronarce garmani	Natal electric ray	align=center	unknown	Vulnerable^[60]
Stingray		Himantura gerrardi	Whitespotted whipray	align=center	unknown	Vulnerable^[61]			freshwater
Stingray		Himantura hortlei	Hortle's whipray	align=center	decreasing	Vulnerable^[62]			freshwater
Stingray		Himantura kittipongi	Roughback whipray	align=center	unknown	Endangered^[63]			freshwater
Stingray		Himantura leoparda	Leopard whipray	align=center	decreasing	Vulnerable^[64]			freshwater
Stingray		Himantura lobistoma	Tubemouth whipray	align=center	decreasing	Vulnerable^[65]			freshwater
Stingray		Himantura oxyrhyncha	Longnose marbled whipray	align=center	unknown	Endangered^[66]			freshwater
Stingray		Himantura pastinacoides	Round whipray	align=center	decreasing	Vulnerable^[67]			freshwater
Stingray		Urogymnus polylepis	Giant freshwater stingray	align=center	decreasing	Endangered^[68]			freshwater
Stingray		Himantura signifer	White-edge freshwater whipray	align=center	unknown	Endangered^[69]			freshwater
Stingray		Himantura uarnacoides	Bleeker's whipray	align=center	decreasing	Vulnerable^[70]			freshwater
Stingray		Himantura uarnak	Reticulate whipray	align=center	decreasing	Vulnerable^[71]
Stingray		Himantura undulata	Bleeker's variegated whipray	align=center	decreasing	Vulnerable^[72]
Skate		Leucoraja circularis	Sandy ray	align=center	unknown	Vulnerable^[73]
Skate		Leucoraja melitensis	Maltese skate	align=center	decreasing	Critically endangered^[74]
Skate		Leucoraja ocellata	Winter skate	align=center	decreasing	Endangered^[75]
Stingray		Myliobatis hamlyni	Purple eagle ray	align=center	decreasing	Endangered^[76]
Skate		Malacoraja senta	Smooth skate	align=center	decreasing	Endangered^[77]
Stingray		Manta alfredi	Reef manta ray	align=center	decreasing	Vulnerable^[78]
Stingray		Manta birostris	Giant manta ray	align=center	decreasing	Endangered^[79]
Stingray		Mobula mobular	Giant devil ray	align=center	decreasing	Endangered^[80]
Stingray		Mobula rochebrunei	Lesser Guinean devil ray	align=center	unknown	Vulnerable^[81]
Electric ray		Narcine brevilabiata	Shortlip electric ray	align=center	unknown	Vulnerable^[82]
Electric ray		Narke japonica	Japanese sleeper ray	align=center	unknown	Vulnerable^[83]
Skate		Okamejei pita	Pita skate	align=center	unknown	Critically endangered^[84]
Stingray		Pastinachus solocirostris	Roughnose stingray	align=center	unknown	Endangered^[85]
Stingray		Platyrhina sinensis	Fanray	align=center	unknown	Vulnerable^[86]
Sawfish		Pristis clavata	Queensland sawfish	align=center	decreasing	Critically endangered^[87]
Sawfish		Pristis pectinata	Smalltooth sawfish	align=center	decreasing	Critically endangered^[88]
Sawfish		Pristis pristis	Common sawfish	align=center	decreasing	Critically endangered^[89]
Sawfish		Pristis zijsron	Narrowsnout sawfish	align=center	decreasing	Critically endangered^[90]
Skate		Raja pulchra	Mottled skate	align=center	decreasing	Vulnerable^[91]
Skate		Raja undulata	Undulate ray	align=center	decreasing	Endangered^[92]
Sawfish		Rhina ancylostoma	Bowmouth guitarfish	align=center	decreasing	Critically endangered^[93]
Sawfish		Rhinobatos albomaculatus	White-spotted guitarfish	align=center	decreasing	Vulnerable^[94]
Sawfish		Rhinobatos cemiculus	Blackchin guitarfish	align=center	decreasing	Endangered^[95]
Sawfish		Rhinobatos formosensis	Taiwan guitarfish	align=center	unknown	Vulnerable^[96]
Sawfish		Rhinobatos horkelii	Brazilian guitarfish	align=center	decreasing	Critically endangered^[97]
Sawfish		Rhinobatos irvinei	Spineback guitarfish	align=center	decreasing	Vulnerable^[98]
Sawfish		Rhinobatos jimbaranensis	Jimbaran shovelnose ray	align=center	decreasing	Vulnerable^[99]
Sawfish		Rhinobatos obtusus	Widenose guitarfish	align=center	decreasing	Vulnerable^[100]
Sawfish		Rhinobatos penggali	Indonesian shovelnose ray	align=center	decreasing	Vulnerable^[101]
Sawfish		Rhinobatos rhinobatos	Common guitarfish	align=center	decreasing	Endangered^[102]
Stingray		Rhinoptera brasiliensis	Brazilian cownose ray	align=center	decreasing	Endangered^[103]
Stingray		Rhinoptera javanica	Javanese cownose ray	align=center	unknown	Vulnerable^[104]
Skate		Rhinoraja albomaculata	White-dotted skate	align=center	unknown	Vulnerable^[105]
Sawfish		Rhynchobatus australiae	White-spotted guitarfish	align=center	decreasing	Vulnerable^[106]
Sawfish		Rhynchobatus djiddensis	Giant guitarfish	align=center	decreasing	Vulnerable^[107]
Sawfish		Rhynchobatus laevis	Smoothnose wedgefish	align=center	unknown	Vulnerable^[108]
Sawfish		Rhynchobatus luebberti	Lubbert's guitarfish	align=center	decreasing	Endangered^[109]
Sawfish		Rhynchobatus sp. nov. A	Roughnose wedgefish	align=center	decreasing	Vulnerable^[110]
Sawfish		Rhynchobatus springeri	Broadnose wedgefish	align=center	decreasing	Vulnerable^[111]
Skate		Rioraja agassizii	Rio skate	align=center	unknown	Vulnerable^[112]
Skate		Rostroraja alba	Bottlenose skate	align=center	decreasing	Endangered^[113]
Skate		Spiniraja whitleyi	Melbourne skate	align=center	unknown	Vulnerable^[114]
Skate		Sympterygia acuta	Bignose fanskate	align=center	decreasing	Vulnerable^[115]
Stingray		Taeniura meyeni	Round ribbontail ray	align=center	unknown	Vulnerable^[116]
Electric ray		Temera hardwickii	Finless sleeper ray	align=center	unknown	Vulnerable^[117]
Sawfish		Trygonorrhina melaleuca	Magpie fiddler ray	align=center	unknown	Endangered^[118]
Stingray		Urogymnus asperrimus	Porcupine ray	align=center	unknown	Vulnerable^[119]
Stingray		Urogymnus ukpam	Pincushion ray	align=center	unknown	Endangered^[120]
Stingray		Urolophus bucculentus	Sandyback stingaree	align=center	decreasing	Vulnerable^[121]
Stingray		Urolophus orarius	Coastal stingaree	align=center	unknown	Endangered^[122]
Stingray		Urolophus sufflavus	Yellowback stingaree	align=center	decreasing	Vulnerable^[123]
Stingray		Urolophus viridis	Greenback stingaree	align=center	decreasing	Vulnerable^[124]
Stingray		Urotrygon reticulata	Reticulate round stingray	align=center	unknown	Vulnerable^[125]
Stingray		Urotrygon simulatrix	Fake round ray	align=center	unknown	Vulnerable^[126]
Sawfish		Zapteryx brevirostris	Shortnose guitarfish	align=center	decreasing	Vulnerable^[127]
Skate		Zearaja chilensis	Yellownose skate	align=center	decreasing	Vulnerable^[128]
Skate		Zearaja maugeana	Maugean skate	align=center	unknown	Endangered^[129]

See also

• List of threatened sharks

Further reading

• The Conservation Status of Pelagic Sharks and Rays . IUCN Shark Specialist Group . April 3, 2012 . Camhi, M.D . Valenti, S.V. . Fordham, S.V. . Fowler, S.L. . Gibson, C. . Pelagic Shark Red List Workshop . February 2007 . Oxford, England . 978-0-9561063-1-5 . https://web.archive.org/web/20110114184614/http://cmsdata.iucn.org/downloads/ssg_pelagic_report_final.pdf . January 14, 2011 . dead .
• Dulvy, N.K., Baum, J.K., Clarke, S., Compagno, L.J.V., Cortés, E., Domingo, A., Fordham, S., Fowler, S., Francis, M.P., Gibson, C., Martínez, J., Musick, J.A., Soldo, A., Stevens, J.D. and Valenti, S. (2008) "You can swim but you can’t hide: The global status and conservation of oceanic pelagic sharks and rays" Aquatic Conservation: Marine and Freshwater Ecosystems, 18 (5): 459–482.
• FAO (2000) Conservation and Management of Sharks Technical Guidelines for Responsible Fisheries, Rome. .
• Faria VV, McDavitt MT, Charvet P, Wiley TR, Simpfendorfer CA and Naylor GJP (2013) "Species delineation and global population structure of Critically Endangered sawfishes (Pristidae)" Zoological Journal of the Linnean Society, 167 (1): 136–164.
• Fowler SL, Cavanagh RD, Camhi M, Burgess GH, Cailliet GM, Fordham SV, Simpfendorfer CA and Musick JA (comp. and ed.) (2005) Sharks, Rays and Chimaeras: The Status of the Chondrichthyan Fishes IUCN Shark Specialist Group, Status Survey. .
• Musick, John A and Bonfil, Ramón (2005) Management techniques for elasmobranch fisheries Fisheries Technical Paper 474, FAO, Rome.
• Teutscher, Frans (2004) "Sharks (Chondrichthyes)" In: World markets and industry of selected commercially-exploited aquatic species with an international conservation profile, Camillo Catarci. Fisheries Circular 990, FAO
• Endangered and Threatened Species Under NMFS’ Jurisdiction National Marine Fisheries Service, NOAA. Updated: 28 February 28, 2013.

External links

• Protecting the fragile manta rays of the Maldives BBC, 5 March 2012.

Notes and References

1. Book: Guidelines for Using the IUCN Red List Categories and Criteria . . August 2010 . 2012-09-05 . https://web.archive.org/web/20110221235804/http://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf . 21 February 2011 .
2. Web site: Red List Overview . IUCN Red List . . dead . https://web.archive.org/web/20120527175928/http://www.iucnredlist.org/about/red-list-overview . 2012-05-27 .
3. https://www.sciencedaily.com/releases/2010/10/101026184209.htm Third of Shark and Ray Species Are Threatened, Study Suggests
4. Hoffmann M, C Hilton-Taylor and 173 others (2010) "The impact of conservation on the status of the world's vertebrates" Science, 330: 1503–1509.
5. http://www.cites.org/common/com/ac/18/E18i-01.doc Conservation of Sharks
6. Web site: The Greatest Threats to Sharks. 2007. Oceana. 2009-08-29. https://web.archive.org/web/20090603052356/http://oceana.org/sharks/threats/ . 2009-06-03.
7. http://www.cites.org/common/com/ac/25/E25i-06.pdf The Future of Sharks: A Review of Action and Inaction
8. Guide of Mediterranean Skates and Rays. Mendez L., Bacquet A. and F. Briand. Oct. 2022.https://www.researchgate.net/publication/364309239_Guide_of_Mediterranean_Skates_and_Rays
9. https://www.newscientist.com/blogs/shortsharpscience/2013/03/manta-ray-mouth.html Glider of the sea threatened by 'traditional' medicine
10. Ari, C.. 2011. Encephalization and brain organization of mobulid rays (Myliobatiformes, Elasmobranchii) with ecological perspectives. Open Anatomy Journal. 3. 1–13. 10.2174/1877609401103010001. free.
11. News: No species is safe from burgeoning wildlife trade. 2013-03-13. The New York Times. 2013-03-12. Wassener, Bettina.
12. http://blogs.scientificamerican.com/extinction-countdown/2012/01/17/manta-rays-endangered-by-sudden-demand-from-chinese-medicine/ Manta Rays Endangered by Sudden Demand from Chinese Medicine
13. https://sharksavers.box.com/s/44nbx900qajxghzb4oh8 "The global threat to manta and mobula rays"
14. Web site: Manta Fisheries . The Manta Trust . 2011 . 2013-02-21 . https://web.archive.org/web/20130127060340/http://www.mantatrust.org/threats/manta-fisheries/ . 2013-01-27 . dead .
15. Web site: Sri Lanka Manta Project . The Manta Trust . 2011 . 2013-02-21 . https://web.archive.org/web/20130127165733/http://www.mantatrust.org/in-the-field/sri-lanka/ . 2013-01-27 . dead .
16. Web site: COP10 Outcome: Migratory Manta Ray under CMS Protection . 2011-11-25 . Convention on Migratory Species . UNEP; CMS . 2013-02-19.
17. http://www.cites.org/eng/cop/16/prop/E-CoP16-Prop-46.pdf Proposal 46: Genus Manta
18. http://www.cites.org/eng/news/pr/2013/20130314_cop16.php CITES conference takes decisive action to halt decline of tropical timber, sharks, manta rays and a wide range of other plants and animals
19. https://web.archive.org/web/20051110135951/http://news.nationalgeographic.com/news/2003/06/0604_030604_sawfish_2.html Sawfish Is First Sea Fish on U.S. Endangered List
20. http://www.cites.org/eng/cop/16/prop/E-CoP16-Prop-45.pdf Proposal 45: Pristis microdon
21. Web site: CITES Appendices I, II and III . Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES) . 7 March 2013.
22. 10.1038/s41586-020-03173-9. Half a century of global decline in oceanic sharks and rays. 2021. Pacoureau. Nathan. Rigby. Cassandra L.. Kyne. Peter M.. Sherley. Richard B.. Winker. Henning. Carlson. John K.. Fordham. Sonja V.. Barreto. Rodrigo. Fernando. Daniel. Francis. Malcolm P.. Jabado. Rima W.. Herman. Katelyn B.. Liu. Kwang-Ming. Marshall. Andrea D.. Pollom. Riley A.. Romanov. Evgeny V.. Simpfendorfer. Colin A.. Yin. Jamie S.. Kindsvater. Holly K.. Dulvy. Nicholas K.. Nature. 589. 7843. 567–571. 33505035. 2021Natur.589..567P . 10871/124531 . 231723355. free.
23. 10.1111/faf.12119. Why have global shark and ray landings declined: Improved management or overfishing?. 2016. Davidson. Lindsay N K.. Krawchuk. Meg A.. Dulvy. Nicholas K.. Fish and Fisheries. 17. 2. 438–458.
24. 10.1016/j.cub.2017.04.038. Challenges and Priorities in Shark and Ray Conservation. 2017. Dulvy. Nicholas K.. Simpfendorfer. Colin A.. Davidson. Lindsay N.K.. Fordham. Sonja V.. Bräutigam. Amie. Sant. Glenn. Welch. David J.. Current Biology. 27. 11. R565–R572. 28586694. 20632998. free.
25. 10.1016/j.cub.2016.12.017. Bright spots of sustainable shark fishing. 2017. Simpfendorfer. Colin A.. Dulvy. Nicholas K.. Current Biology. 27. 3. R97–R98. 28171764. 46839247. free.
26. 10.1371/journal.pone.0040542. Benefits of Rebuilding Global Marine Fisheries Outweigh Costs. 2012. Sumaila. Ussif Rashid. Cheung. William. Dyck. Andrew. Gueye. Kamal. Huang. Ling. Lam. Vicky. Pauly. Daniel. Srinivasan. Thara. Swartz. Wilf. Watson. Reginald. Zeller. Dirk. PLOS ONE. 7. 7. e40542. 22808187. 3396648. 2012PLoSO...740542S . free.
27. White, W.T. . 2006 . Aetobatus flagellum . 2006 . e.T60119A12306888 . 10.2305/IUCN.UK.2006.RLTS.T60119A12306888.en . 17 January 2020.
28. Kyne, P.M. . Compagno, L.J.V. . Bennett, M.B. . 2016 . Aetomylaeus nichofii . 2016 . e.T84784918A68607458 . 10.2305/IUCN.UK.2016-1.RLTS.T84784918A68607458.en . 12 November 2021.
29. White, W.T. . Kyne, P.M. . 2016 . Aetomylaeus vespertilio . 2016 . e.T60121A68607665 . 10.2305/IUCN.UK.2016-1.RLTS.T60121A68607665.en . 17 January 2020.
30. Kulka, D.W. . Sulikowski, J. . Gedamke, J. . Pasolini, P. . Endicott, M. . 2009 . Amblyraja radiata . 2009 . e.T161542A5447511 . 10.2305/IUCN.UK.2009-2.RLTS.T161542A5447511.en . 17 January 2020.
31. D'Anastasi, B. . Simpfendorfer, C. . van Herwerden, L. . 2013 . Anoxypristis cuspidata . 2019 . 2013 . e.T39389A141789456 . 10.2305/IUCN.UK.2013-1.RLTS.T39389A141789456.en . 17 January 2020.
32. Last, P.R. . Kyne, P.M. . Sherman, C.S. . 2015 . Aptychotrema timorensis . 2015 . e.T60179A68609203 . 10.2305/IUCN.UK.2015-4.RLTS.T60179A68609203.en . 17 January 2020.
33. Hozbor, N. . Massa, A. . Vooren, C.M. . 2004 . Atlantoraja castelnaui . 2004 . e.T44575A10921544 . 10.2305/IUCN.UK.2004.RLTS.T44575A10921544.en . 17 January 2020.
34. Massa, A. . Hozbor, N. . Vooren, C.M. . 2006 . Atlantoraja cyclophora . 2006 . e.T61398A12462475 . 10.2305/IUCN.UK.2006.RLTS.T61398A12462475.en . 17 January 2020.
35. San Martín, J.M. . Stehmann, M.F.W. . Kyne, P.M. . 2007 . Atlantoraja platana . 2007 . e.T63110A12608554 . 10.2305/IUCN.UK.2007.RLTS.T63110A12608554.en . 17 January 2020.
36. McCormack, C. . Lamilla, J. . San Martín, M.J. . Stehmann, M.F.W. . 2007 . Bathyraja griseocauda . 2007 . e.T63113A12609854 . 10.2305/IUCN.UK.2007.RLTS.T63113A12609854.en . 17 January 2020.
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