Taxonomy of the Gastropoda (Bouchet et al., 2017) explained
The taxonomy of the Gastropoda as it was revised in December 2017 by Philippe Bouchet and eight other authors, is a publication which lays out a newly revised system for the scientific classification of gastropod mollusks.[1] The same work also included the taxonomy of monoplacophorans.
The publication is entitled, "Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families"; it was published by the journal Malacologia.
Summary of the taxonomy
The following cladogram shows a summary of the new gastropod taxonomy, according to the work's Contents section:
†Paleozoic molluscs of uncertain position within Gastropoda or Monoplacophora
(Unassigned to Superfamily)
†Superfamily Archinacelloidea Knight, 1952
†Subclass Amphigastropoda
†Superfamily Bellerophontoidea McCoy, 1852
- †Family Bellerophontidae McCoy, 1852
- †Subfamily Bellerophontinae McCoy, 1852 (=Liljevallospiridae Golikov & Starobogatov, 1989)
- †Subfamily Bucanopsinae Wahlman, 1992
- †Subfamily Cymbulariinae Horný, 1963
- †Subfamily Bellerophontinae Knight, 1956
- †Family Bucanellidae Koken, 1925
- †Family Bucaniidae Ulrich & Scofield, 1897
- †Subfamily Bucaniinae Knight, 1956 (=Grandostomatinae Horný, 1962)
- †Subfamily Plectonotinae Boucot & Yochelson, 1966
- †Tribe Plectonotini Boucot & Yochelson, 1966
- †Tribe Boucotonotini Frýda, 1999
- †Subfamily Salpingostomatinae Koken, 1925
- †Subfamily Undulabucaniinae Wahlman, 1992
- †Family Euphemitidae Knight, 1956
- †Subfamily Euphemitinae Knight, 1956
- †Subfamily Paleuphemitinae Frýda, 1999
- †Family Pterothecidae P. Fischer, 1883
- †Subfamily Pterothecinae P. Fischer, 1883
- †Subfamily Carinaropsinae Ulrich & Scofield, 1897
- †Subfamily Pedasiolinae Wahlman, 1992
- †Family Sinuitidae Dall, 1913
- †Subfamily Sinuitinae Dall, 1913 (=Protowarthiidae Ulrich & Scofield, 1897)
- †Subfamily Aiptospirinae Wang, 1980
- †Subfamily Hispanosinuitinae Frýda & Gutierrez-Marco, 1996
- †Family Tremanotidae Naef, 1911
- †Family Tropidodiscidae Knight, 1956 (=Temnodiscinae Horný, 1963)
†Subclass Archaeobranchia
†Superfamily Pelagielloidea Knight, 1956 (=Orthostrophina)
- †Family Pelagiellidae Knight, 1956 (=Proeccyliopteridae Kobayashi, 1962; =Protoscaevogyridae Kobayashi, 1962)
- †Family Aldanellidae Linsley & Kier, 1984
†Superfamily Scenelloidea S. A. Miller, 1889
- †Family Scenellidae S. A. Miller, 1889
- †Subfamily Scenellinae S. A. Miller, 1889 (=Palaeacmaeidae Grabau & Shimer, 1909; =Hampilininae Kobayashi, 1958; =Eosoconidae Yu, 1979; =Merismoconchidae Yu, 1979; =Shelbyoceratidae Stinchcomb, 1986; =Actinoconidae Starobogatov & Moskalev, 1987; =Yangtzemerismatinae Yu, 1987; =Marocellidae Topper, Brock, Skovsted & Paterson, 2009)
- †Subfamily Yangtzeconinae Yu, 1979 (=Ceratoconidae Missarzhevsky, 1989)
- †Family Coreospiridae Knight, 1947 (=Archaeospiridae Yu, 1979; =Yangtzespirinae Yu, 1984; =Latouchellidae Golikov & Starobogatov, 1989)
- †Family Carinopeltidae Parkhaev, 2013 (=Igarkiellidae Parkhaev, 2001)
†Superfamily Yochelcionelloidea Runnegar & Jell, 1976
- †Family Yochelcionellidae Runnegar & Jell, 1976 (=Enigmaconidae MacKinnon, 1985)
- †Family Stenothecidae Runnegar & Jell, 1980
- †Subfamily Stenothecinae Runnegar & Jell, 1980 (=Mellopegmidae Parkhaev, 2001)
- †Subfamily Watsonellinae Parkhaev, 2001
- †Family Securiconidae Missarzhevsky, 1989 (=Rugaeconidae Vassiljeva, 1990; =Trenellidae Parkhaev, 2001)
†Paleozoic basal taxa that are certainly Gastropoda
(Unassigned to Superfamily)
- †Family Codonocheilidae S. A. Miller, 1889
- †Family Craspedostomatidae Wenz, 1938
- †Subfamily Craspedostomatinae Wenz, 1938
- †Subfamily Bucanospirinae Wenz, 1938
- †Family Crassimarginatidae Frýda, Blodgett & Lenz, 2002
- †Family Discohelicidae Schröder, 1995
- †Family Isospiridae Wangberg-Eriksson, 1964
- †Family Yuopisthonematidae Nützel, 2017 (=Opisthonematidae Yu, 1976)
- †Family Paraturbinidae Cossmann, 1916
- †Family Pragoserpulinidae Frýda, 1998
- †Family Raphistomatidae Koken, 1896 (=Ceratopeidae Yochelson & Bridge, 1957)
- †Family Rhytidopilidae Starobogatov, 1976
- †Family Scoliostomatidae Frýda, Blodgett & Lenz, 2002
- †Subfamily Scoliostomatinae Frýda, Blodgett & Lenz, 2002
- †Subfamily Mitchelliinae Frýda, Blodgett & Lenz, 2002
- †Family Sinuopeidae Wenz, 1938
- †Subfamily Sinuopeinae Wenz, 1938
- †Subfamily Platyschismatinae Knight, 1956
- †Subfamily Turbonellininae Knight, 1956
†Superfamily Clisospiroidea S. A. Miller, 1889 (=Mimospirina)
- †Family Clisospiridae S. A. Miller, 1889
- †Subfamily Clisospirinae S. A. Miller, 1889
- †Subfamily Atracurinae Horný, 1964
- †Subfamily Progalerinae Knight, 1956
- †Subfamily Trochoclisinae Horný, 1964
- †Family Onychochilidae Koken, 1925
- †Subfamily Onychochilinae Koken, 1925
- †Subfamily Hyperstropheminae Horný, 1964
- †Subfamily Scaevogyrinae Wenz, 1938
†Superfamily Euomphaloidea White, 1877
- †Family Euomphalidae White, 1877
- †Subfamily Euomphalinae White, 1877 (=Schizostomatidae Bronn, 1849; =Polytropidae Ulrich, 1897; =Straparolinae Cossmann, 1916; =Poleumitidae Wenz, 1938)
- †Subfamily Odontomariinae Frýda, Heidelberger & Blodgett, 2006
- †Family Euomphalopteridae Koken, 1896
- †Subfamily Euomphalopterinae Koken, 1896
- †Subfamily Spinicharybdiinae Rohr, Blodgett & Frýda, 2008
- †Family Helicotomidae Wenz, 1938
- †Family Lesueurillidae P. J. Wagner, 2002
- †Family Omphalocirridae Wenz, 1938
- †Family Omphalotrochidae Knight, 1945
- †Family Straparollinidae P. J. Wagner, 2002
†Superfamily Loxonematoidea Koken, 1889
†Superfamily Macluritoidea Carpenter, 1861
†Superfamily Ophiletoidea Koken, 1907
- †Family Ophiletidae Koken, 1907 (=Ecculiomphalinae Wenz, 1938)
†Superfamily Oriostomatoidea Koken, 1896
†Superfamily Palaeotrochoidea† Knight, 1956
†Superfamily Trochonematoidea Zittel, 1895
Superfamily Eoacmaeoidea Nakano & Ozawa, 2007
Superfamily Patelloidea Rafinesque, 1815
Superfamily Lottioidea Gray, 1840
- Family Lottiidae Gray, 1840
- Subfamily Lottiinae Gray, 1840
- Tribe Lottiini Gray, 1840
- Tribe Patelloidini Chapman & Gabriel, 1923
- Subfamily Tecturinae Gray, 1847
- Family Acmaeidae Forbes, 1850 (=Rhodopetalinae Lindberg, 1981; =Erginini Lindberg, 1990)
- †Family Damilinidae Horný, 1961
- Family Lepetidae Gray, 1850
- Subfamily Lepetinae Gray, 1850
- Subfamily Propilidiinae Thiele, 1891
- †Family Lepetopsidae McLean, 1990
- Family Nacellidae Thiele, 1891 (=Bertiniidae Jousseaume, 1883)
- Family Neolepetopsidae McLean, 1990
- Family Pectinodontidae Pilsbry, 1891
Superfamily Neomphaloidea McLean, 1981
Superfamily Cocculinoidea Dall, 1882
†Paleozoic taxa of uncertain position
- †Family Holopeidae Cossmann, 1908
- †Family Micromphalidae J. A. Harper, 2016
Order Pleurotomariida
†Superfamily Eotomarioidea Wenz, 1938
- †Family Eotomariidae Wenz, 1938
- †Subfamily Eotomariinae Wenz, 1938 (=Liospirinae Knight, 1956)
- †Tribe Eotomariini Wenz, 1938
- †Tribe Deseretospirini Gordon & Yochelson, 1987
- †Tribe Glabrocingulini Gordon & Yochelson, 1987
- †Subfamily Neilsoniinae Knight, 1956
- †Tribe Neilsoniini Knight, 1956
- †Tribe Spirovallini Waterhouse, 2001
- †Family Gosseletinidae Wenz, 1938
- †Subfamily Gosseletininae Wenz, 1938
- †Subfamily Coelozoninae Knight, 1956
- †Tribe Coelozonini Knight, 1956 (=Euryzoninae P. J. Wagner, 2002)
- †Tribe Planozonini Knight, 1956
- †Subfamily Triangulariinae Vostokova, 1960
- †Family Luciellidae Knight, 1956
- †Family Phanerotrematidae Knight, 1956
- †Family Pseudoschizogoniidae Bandel, 2009
- †Family Wortheniellidae Bandel, 2009
†Superfamily Murchisonioidea Koken, 1896
- †Family Murchisoniidae Koken, 1896
- †Subfamily Murchisoniinae Koken, 1896 (=Hormotomidae Wenz, 1938)
- †Subfamily Cheeneetnukiinae Blodgett & Cook, 2002
- †Family Farewelliidae Mazaev, 2011
- †Family Plethospiridae Wenz, 1938 (=Pithodeinae Wenz, 1938)
- †Family Ptychocaulidae Mazaev, 2011
Superfamily Pleurotomarioidea Swainson, 1840
- Family Pleurotomariidae Swainson, 1840
- †Family Catantostomatidae Wenz, 1938
- †Family Lancedelliidae Bandel, 2009
- †Family Phymatopleuridae Batten, 1956
- †Family Polytremariidae Wenz, 1938
- †Family Portlockiellidae Batten, 1956
- †Family Rhaphischismatidae Knight, 1956
- †Family Stuorellidae Bandel, 2009
- †Family Trochotomidae Cox, 1960 (1934) (=Ditremariinae Haber, 1934)
- †Family Zygitidae Cox, 1960
†Superfamily Porcellioidea Koken, 1895
- †Family Porcelliidae Koken, 1895
- †Subfamily Porcelliinae Koken, 1895
- †Subfamily Agnesiinae Knight, 1956
- †Tribe Agnesiini Knight, 1956
- †Tribe Anoriostomatini Frýda & Farrell, 2005
- †Family Cirridae Cossmann, 1916
- †Subfamily Cirrinae Cossmann, 1916
- †Subfamily Platyacrinae Wenz, 1938 (=Hesperocirrinae O. Haas, 1953)
- †Subfamily Cassianocirrinae Bandel, 1993
- †Family Pavlodiscidae Frýda, 1998
†Superfamily Pseudophoroidea S. A. Miller, 1889
†Superfamily Ptychomphaloidea Wenz, 1938
- †Family Ptychomphalidae Wenz, 1938 (=Ptychomphalinini Wenz, 1938; =Mourloniini Yochelson & Dutro, 1960)
- †Family Rhaphistomellidae Bandel, 2009
†Superfamily Schizogonioidea Cox, 1960
- †Family Schizogoniidae Cox, 1960
- †Family Pseudowortheniellidae Bandel, 2009
†Superfamily Sinuspiroidea Mazaev, 2011
- †Family Sinuspiridae Mazaev, 2011
Order Seguenziida
Superfamily Seguenzioidea Verrill, 1884
- Family Seguenziidae Verrill, 1884
- Subfamily Seguenziinae Verrill, 1884
- Tribe Seguenziini Verrill, 1884
- Tribe Fluxinellini B. A.Marshall, 1991 (=Ancistrobasidae Bandel, 2010)
- Subfamily Asthelysinae B. A.Marshall, 1991
- Subfamily Davisianinae Egorova, 1972 (=Putillinae F. Nordsieck, 1972; =Oligomeriinae Egorov, 2000)
- Subfamily Guttulinae Goryachev, 1987
- Family Cataegidae McLean & Quinn, 1987
- Family Chilodontaidae Wenz, 1938
- Family Choristellidae Bouchet & Warén, 1979
- Family Eucyclidae Koken, 1896 (=Amberleyidae Wenz, 1938; =Calliotropini Hickman & McLean, 1990; =Turcicidae Bandel, 2010)
- †Family Eucycloscalidae Gründel, 2007
- Family Eudaroniidae Gründel, 2007
- †Family Eunemopsidae Bandel, 2010
- †Family Lanascalidae Bandel, 1992
- †Family Laubellidae Cox, 1960
- Family Pendromidae Warén, 1991
- †Family Pseudoturcicidae Bandel, 2010
- †Family Sabrinellidae Bandel, 2010
- Family Trochaclididae Thiele, 1928 (=Acremodontinae B. A. Marshall, 1983)
Superfamily Lepetelloidea Dall, 1882
- Family Lepetellidae Dall, 1882
- Family Addisoniidae Dall, 1882
- Subfamily Addisoniinae Dall, 1882
- Subfamily Helicopeltinae B. A. Marshall, 1996
- Family Bathyphytophilidae B. A. Marshall, 1986
- Family Caymanabyssiidae Moskalev, 1978
- Family Cocculinellidae Moskalev, 1971
- Family Osteopeltidae B. A. Marshall, 1987
- Family Pseudococculinidae Hickman, 1983
- Family Pyropeltidae McLean & Haszprunar, 1987
Superfamily Fissurelloidea Fleming, 1822
- Family Fissurellidae Fleming, 1822
- Subfamily Fissurellinae Fleming, 1822
- Subfamily Diodorinae Odhner, 1932
- Subfamily Emarginulinae Children, 1834 (=Rimulidae Anton, 1838; =Fissurellideini Pilsbry, 1890; =Zeidoridae Naef, 1911; =Scutini Christiaens, 1973; =Clypidinidae Golikov & Starobogatov, 1989
- Subfamily Hemitominae Kuroda, Habe & Oyama, 1971
Superfamily Haliotoidea Rafinesque, 1815
Superfamily Lepetodriloidea McLean, 1988
Superfamily Scissurelloidea Gray, 1847
Superfamily Trochoidea Rafinesque, 1815
- Family Trochidae Rafinesque, 1815
- Subfamily Trochinae Rafinesque, 1815 (=Pyramidinae Gray, 1847)
- Subfamily Alcyninae Williams, Donald, Spencer & Nakano, 2010
- Subfamily Cantharidinae Gray, 1857 (=Pagodatrochidae Bandel, 2010)
- Subfamily Chrysostomatinae Williams, Donald, Spencer & Nakano, 2010
- Subfamily Fossarininae Bandel, 2009
- Subfamily Halistylinae Keen, 1958
- Subfamily Kaiparathininae B. A. Marshall, 1993
- Subfamily Monodontinae Gray, 1857
- Subfamily Stomatellinae Gray, 1840 (=Stomatiidae Carpenter, 1861)
- Subfamily Umboniinae H. Adams & A. Adams, 1854 (1840) (=Rotellinae Swainson, 1840; =Talopiidae Finlay, 1928; =Bankiviini Hickman & McLean, 1990; =Lirulariinae Hickman & McLean, 1990; =Monileini Hickman & McLean, 1990; =Isandini Hickman, 2003)
- Family Angariidae Gray, 1857 (=Delphinulinae Stoliczka, 1868)
- †Family Anomphalidae Wenz, 1938
- †Family Araeonematidae Nützel, 2012
- Family Areneidae McLean, 2012
- Family Calliostomatidae Thiele, 1924 (1847)
- Subfamily Calliostomatinae Thiele, 1924 (1847) (=Ziziphininae Gray, 1847)
- †Subfamily Callotrochinae Szabó, 2011
- Subfamily Fautricinae B. A. Marshall, 1995
- Subfamily Margarellinae Williams, 2013
- Subfamily Thysanodontinae B. A. Marshall, 1988
- Subfamily Xeniostomatinae McLean, 2012
- Family Colloniidae Cossmann, 1917
- Subfamily Colloniinae Cossmann, 1917 (=Bothropomatinae Thiele, 1924; =Homalopomatinae Keen, 1960)
- †Subfamily Crossostomatinae Cox, 1960
- †Tribe Tadeorbisinini Monari, Conti & Szabó, 1995
- †Tribe Costataphrini Gründel, 2008
- †Tribe Crossostomatini Cox, 1960
- †Tribe Helicocryptini Cox, 1960
- †Subfamily Lewisiellinae Gründel, 2008
- Subfamily Liotipomatinae McLean, 2012
- Subfamily Moelleriinae Hickman & McLean, 1990
- †Subfamily Petropomatinae Cox, 1960
- Family Conradiidae Golikov & Starobogatov, 1987 (=Crosseolidae Hickman, 2013)
- †Family Nododelphinulidae Cox, 1960
- †Family Elasmonematidae Knight, 1956
- †Family Epulotrochidae Gründel, Keupp & Lang, 2017
- †Family Eucochlidae Bandel, 2002
- Family Liotiidae Gray, 1850
- Subfamily Liotiinae Gray, 1850 (=Cyclostrematidae P. Fischer, 1885)
- †Subfamily Brochidiinae Yochelson, 1956
- †Subfamily Dichostasiinae Yochelson, 1956
- Family Margaritidae Thiele, 1924 (=Margaritinae Stoliczka, 1868; =Gazidae Hickman & McLean, 1990)
- †Family Metriomphalidae Gründel, Keupp & Lang, 2017
- †Family Microdomatidae Wenz, 1938
- †Subfamily Microdomatinae Wenz, 1938
- †Subfamily Decorospirinae Blodgett & Frýda, 1999
- Family Phasianellidae Swainson, 1840
- Subfamily Phasianellinae Swainson, 1840 (=Eutropiinae Gray, 1847)
- Subfamily Gabrieloninae Hickman & McLean, 1990
- Subfamily Tricoliinae Woodring, 1928
- †Family Proconulidae Cox, 1960 (=Parataphrinae Calzada, 1989)
- †Family Sclarotrardidae Gründel, Keupp & Lang, 2017
- Family Skeneidae W. Clark, 1851 (=Delphinoideinae Thiele, 1924)
- Family Solariellidae Powell, 1951 (=Minoliinae Kuroda, Habe & Oyama, 1971)
- Family Tegulidae Kuroda, Habe & Oyama, 1971
- Family Turbinidae Rafinesque, 1815
- Subfamily Turbininae Rafinesque, 1815 (=Senectinae Swainson, 1840; =Imperatorinae Gray, 1847; =Astraliinae H. Adams & A. Adams, 1854; =Astraeinae Davies, 1935; =Bolmidae Delpey, 1941)
- †Subfamily Moreanellinae J. C. Fischer & Weber, 1997
- Subfamily Prisogastrinae Hickman & McLean, 1990
- †Family Tychobraheidae Horný, 1992
- †Family Velainellidae Vasseur, 1880
†Paleozoic taxa of uncertain position
†Superfamily Nerrhenoidea Bandel & Heidelberger, 2001
†Superfamily Platyceratoidea Hall, 1879
- †Family Platyceratidae Hall, 1879 (=Cyclonematidae P. Fischer, 1885; =Platyostomatidae S. A. Miller, 1889; =Strophostylidae Grabau & Shimer, 1909; =Palaeocapulidae Grabau, 1936)
- †Family Orthonychiidae Bandel & Frýda, 1999
- †Family Vltaviellidae Bandel & Frýda, 1999
- †Subfamily Vltaviellinae Bandel & Frýda, 1999
- †Subfamily Krameriellinae Frýda & Heidelberger, 2003
Superfamily Helicinoidea
Superfamily Hydrocenoidea
†Superfamily Naticopsoidea
- †Family Naticopsidae
- †Subfamily Naticopsinae
- †Subfamily Ampezzonaticopsinae
- †Subfamily Hologyrinae
- †Family Scalaneritinidae
- †Family Trachyspiridae
- †Family Tricolnaticopsidae
Superfamily Neritoidea
- Family Neritidae
- Subfamily Neritinae (=Neritellinae; =Protoneritidae)
- Subfamily Neritininae (=Orthopomatini; =Stenopomatini; =Septariini; =Theodoxinae)
- Subfamily Smaragdiinae
- †Subfamily Velatinae
- †Family Cortinellidae
- †Family Neridomidae
- †Family Neritariidae
- †Subfamily Neritariinae
- †Subfamily Oncochilinae
- †Subfamily Trachyneritariinae
- †Family Otostomidae
- †Family Parvulatopsidae
- Family Phenacolepadidae
- †Family Pileolidae
Superfamily Neritopsoidea
†Superfamily Symmetrocapuloidea
†Fossil taxa of uncertain position
Unassigned to Superfamily
†Superfamily Dendropupoidea
†Superfamily Peruneloidea
†Superfamily Subulitoidea
Superfamily Ampullarioidea
Superfamily Cyclophoroidea
Superfamily Viviparoidea
Unassigned to Superfamily
†Superfamily Acteoninoidea
†Superfamily Orthonematoidea
†Superfamily Palaeostyloidea
†Superfamily Pseudozygopleuroidea
†Superfamily Soleniscoidea
Subcohort Campanilimorpha
Superfamily Campaniloidea
Taxa of uncertain position
=Taxa of uncertain position
=
=
Cohort Ringipleura
=Order Ringiculida
=
=
=Order Nudibranchia
=Suborder Doridina
Suborder Cladobranchia
=
=
=
=
=Suborder EuthecosomataSuborder PseudothecosomataSuborder Gymnosomata
=
=Superorder Siphonarimorpha
=
=Superorder Pylopulmonata
=
=Superorder Acochlidimorpha
=
=
=
See also
Notes and References
- Bouchet. Philippe. Jean-Pierre Rocroi . Rocroi. Jean-Pierre. Hausdorf. Bernhard. Kaim. Andrzej. Kano. Yasunori. Nützel. Alexander. Parkhaev. Pavel. Schrödl. Michael. Strong. Ellen E.. 2017. Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families. Malacologia. 1–2. 0076-2997. 1–526. 61. 10.4002/040.061.0201. 91051256 . Philippe Bouchet .
