Taldycupedidae Explained

Taldycupedidae is an extinct family of beetles primarily known from the Permian period.

Taxonomy

Kirejtshuk (2020) included the following genera in Taldycupedidae, preliminarily treating it as a subfamily Taldycupedinae of the family Permocupedidae.^[1]

• Afrotaldycupes – Beaufort Group, South Africa, Late Permian (Changhsingian)
  • Afrotaldycupes africanus (Type species)
  • Afrotaldycupes lidgettoniensis
• Mesothoris – Blackstone Formation, Australia Late Triassic (Norian)
  • Mesothoris clathrata (Type species)
  • Mesothoris grandis
  • Mesothoris punctomarginum
• Proterocupes (syn Lobanovia)
  • Proterocupes major – Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian)
  • Proterocupes nedubrovensis – Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian)
  • Proterocupes permiana (Type species) – Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian)
• Simmondsia – Blackstone Formation, Australia Late Triassic (Norian)
  • Simmondsia cylindrica
  • Simmondsia subpiriformis (Type species)
• Taldycupes (syn Cryptocupes, Taldycupidium)
  • Taldycupes bergi – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Taldycupes cellulosus – Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian)
  • Taldycupes khalfini (Type species) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Taldycupes moltshanovi – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Taldycupes pingi – Yinping Formation, Anhui, China, Middle Permian (Capitanian)
  • Taldycupes reticulatus – Akkolka Formation, Kazakhstan, Late Permian (Changhsingian)
  • Taldycupes rosanovi – Limptekon Formation, Krasnoyarsk, Russia, Late Permian (Changhsingian)
  • Taldycupes vasjuchitshevi – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
• Tecticupes (syn Stegocupes)
  • Tecticupes heckeri (Type species) – Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian) Mitina Formation, Kemerovo, Russia, Middle Permian (Roadian)
  • Tecticupes indistinctus – Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian)
  • Tecticupes martynovi – Amanak Formation. Orenburg, Middle Permian (Capitanian)
• Tychticupes – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Tychticupes beljanini
  • Tychticupes radtshenkoi (Type species)
  • Tychticupes ragozini
  • "Tychticupes" ussovi (nomen dubium)
• Tychticupoides – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Tychticupoides grjazevi (Type species)
• Uskaticupes (nomen dubium) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Uskaticupes javorskyi (nomen dubium)

The genera Alveolacupes, Clathropenna and Penecupes were previously placed in the family by some authors, but were all excluded from it by Kirejtshuk (2020). The genera Wuchangia and Yuxianocoleus, both from the Lower Jurassic of Hubei, China, were also excluded from the family by Zhao et al. (2021) (as the subfamily Taldycupedinae).^[2]

Notes and References

1. Kirejtshuk. Alexander G.. 2020-02-17. Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea. Geosciences. 10. 2. 73. 2020Geosc..10...73K. 10.3390/geosciences10020073. 2076-3263. free.
2. Zhao . Xianye . Yu . Yilun . Clapham . Matthew E . Yan . Evgeny . Chen . Jun . Jarzembowski . Edmund A . Zhao . Xiangdong . Wang . Bo . 2021-11-08 . Perry . George H . Fikacek . Martin . Early evolution of beetles regulated by the end-Permian deforestation . eLife . 10 . e72692 . 10.7554/eLife.72692 . 34747694 . 8585485 . 2050-084X . free .