Leiognathidae Explained

Leiognathidae, the ponyfishes, slipmouths or slimys / slimies, are a small family of fishes in the order Perciformes. They inhabit marine and brackish waters in the Indian and West Pacific Oceans. They can be used in the preparation of bagoong.

Characteristics

Ponyfishes are small and laterally compressed in shape, with a bland, silvery colouration. They are distinguished by highly extensible mouths, and the presence of a mechanism for locking the spines in the dorsal and anal fins. They also possess a highly integrated light organ in their throats that houses symbiotic bioluminescent bacteria that project light through the animal's underside.[1] [2] [3] Typically, the harbored bacterium is only Photobacterium leiognathi, but in the two ponyfish species Photopectoralis panayensis and Photopectoralis bindus, Photobacterium mandapamensis is also present.[4] Two of the most widely studied uses for luminescence in ponyfish are camouflage by ventral counterillumination[5] [6] and species-specific sexual dimorphism.[7] [8] The light organ systems of ponyfishes are highly variable across species and often between sexes.

Taxonomy

Leiognathidae is classified within the suborder Percoidei by the 5th edition of Fishes of the World, but they are placed in an unnamed clade which sits outside the superfamily Percoidea. This clade contains 7 families which appear to have some relationship to Acanthuroidei, Monodactylidae, and Priacanthidae.[9] Other authorities have placed the family in the order Chaetodontiformes alongside the family Chaetodontidae.[10]

Timeline of genera

ImageSize = width:1000px height:auto barincrement:15pxPlotArea = left:10px bottom:50px top:10px right:10px

Period = from:-65.5 till:10TimeAxis = orientation:horizontalScaleMajor = unit:year increment:5 start:-65.5ScaleMinor = unit:year increment:1 start:-65.5TimeAxis = orientation:horAlignBars = justify

Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:NAM2

bar:space bar:period bar:space bar:era

PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:Paleocene from: -55.8 till: -33.9 color:eocene text:Eocene from: -33.9 till: -23.03 color:oligocene text:Oligocene from: -23.03 till: -5.332 color:miocene text:Miocene from: -5.332 till: -2.588 color:pliocene text:Plio. from: -2.588 till: -0.0117 color:pleistocene text:Pleist. from: -0.0117 till: 0 color:holocene text:H.

bar:eratop from: -65.5 till: -23.03 color:paleogene text:Paleogene from: -23.03 till: -2.588 color:neogene text:Neogene from: -2.588 till: 0 color:quaternary text:Q.

PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left

color:eocene bar:NAM1 from: -55.8 till: 0 text: Gazza color:oligocene bar:NAM2 from: -33.9 till: 0 text: Leiognathus

PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25

bar:period from: -65.5 till: -55.8 color:paleocene text:Paleocene from: -55.8 till: -33.9 color:eocene text:Eocene from: -33.9 till: -23.03 color:oligocene text:Oligocene from: -23.03 till: -5.332 color:miocene text:Miocene from: -5.332 till: -2.588 color:pliocene text:Plio. from: -2.588 till: -0.0117 color:pleistocene text:Pleist. from: -0.0117 till: 0 color:holocene text:H.

bar:era from: -65.5 till: -23.03 color:paleogene text:Paleogene from: -23.03 till: -2.588 color:neogene text:Neogene from: -2.588 till: 0 color:quaternary text:Q.

Genera

The following genera are classified within the Leiognathidae:

Notes and References

  1. Book: Paxton, J.R. . Eschmeyer, W.N.. Johnson, G.D. . Gill, A.C. . amp . 1998. Encyclopedia of Fishes. Academic Press. San Diego. 978-0-12-547665-2. 186.
  2. Haneda. Y.. Tsuji. F.I.. 1976. Luminescent systems of pony fishes. Journal of Morphology. 150. 2. 539–552. 10.1002/jmor.1976.150.2.539. 91711575 .
  3. McFall-Ngai. M.J.. Dunlap. P.V.. 1984. External and internal sexual dimorphism in leiognathid fishes: Morphological evidence for sex-specific bioluminescent signaling. Journal of Morphology. en. 182. 1. 71–83. 10.1002/jmor.1051820105. 6492170. 23956409 . 1097-4687.
  4. Kaeding. A.J.. Ast. J.C.. Pearce. M.M.. Urbanczyk. H.. Kimura. S.. Endo. H.. Nakamura. M.. Dunlap. P.V.. 2007. Phylogenetic Diversity and Cosymbiosis in the Bioluminescent Symbioses of "Photobacterium mandapamensis". Applied and Environmental Microbiology. en. 73. 10. 3173–3182. 10.1128/AEM.02212-06. 0099-2240. 1907103. 17369329. 2007ApEnM..73.3173K .
  5. Hastings. J.W.. 1971. Light to Hide by: Ventral Luminescence to Camouflage the Silhouette. Science. en. 173. 4001. 1016–1017. 10.1126/science.173.4001.1016. 0036-8075. 17796582. 1971Sci...173.1016W . 5680392 .
  6. McFall-Ngai. M.J.. Morin. J.G.. 1991. Camouflage by Disruptive Illumination in Leiognathids, a Family of Shallow-Water, Bioluminescent Fishes. Journal of Experimental Biology. en. 156. 1. 119–137. 10.1242/jeb.156.1.119 . 0022-0949.
  7. Sparks. J.S.. Dunlap. P.V.. Smith. W.L.. 2005. Evolution and diversification of a sexually dimorphic luminescent system in ponyfishes (Teleostei: Leiognathidae), including diagnoses for two new genera. Cladistics. en. 21. 4. 305–327. 10.1111/j.1096-0031.2005.00067.x. 34892969 . 1096-0031. 2027.42/72092. 53063530 . free.
  8. Chakrabarty. P.. Davis. M.P.. Smith. W.L.. Berquist. R.. Gledhill. K.M.. Frank. L.R.. Sparks. J.S.. 2011. Evolution of the light organ system in ponyfishes (Teleostei: Leiognathidae). Journal of Morphology. en. 272. 6. 704–721. 10.1002/jmor.10941. 21433053. 9331522 . 1097-4687.
  9. Book: Fishes of the World . 5th . J. S. Nelson . T. C. Grande . M. V. H. Wilson . 2016 . 453 . Wiley . 978-1-118-34233-6 .
  10. R. Betancur-Rodriguez . E. Wiley . N. Bailly . A. Acero . M. Miya . G. Lecointre . G. Ortí . 10.1186/s12862-017-0958-3 . Phylogenetic Classification of Bony Fishes – Version 4 . 2017 . BMC Evolutionary Biology . 17 . 162. 162 . 28683774 . 5501477 . free .
  11. Sparks . J.S. . Chakrabarty . P. . 2015 . Description of a new genus of ponyfishes (Teleostei: Leiognathidae), with a review of the current generic-level composition of the family . Zootaxa . 3947 . 2. 181–190 . 10.11646/zootaxa.3947.2.2. 25947728 .