Pterostichus Explained

Pterostichus is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer slightly moist, sandy soil although preferences differ between species.^[1]

Species

These species and subgenera are included in the genus Pterostichus.Subgenus Abea Morita, 1992

Pterostichus yamauchii Morita, 1992

Subgenus Adelopterus Reitter, 1886

Pterostichus ambiguus (Fairmaire, 1858)

Subgenus Adelosia Stephens, 1835

Pterostichus macer (Marsham, 1802)

Subgenus Agastillus Reitter, 1892

Pterostichus capitatus (Chaudoir, 1850)
Pterostichus cratocephalus (Tschitscherine, 1897)
Pterostichus cucujinus Reitter, 1892
Pterostichus eriwanicus (Tschitscherine, 1897)

Subgenus Anilloferonia Van Dyke, 1926

Pterostichus diana LaBonte, 2013
Pterostichus malkini (Hatch, 1953)
Pterostichus testaceus (Van Dyke, 1926)

Subgenus Anomostichus Sciaky, 1995

Pterostichus anomostriatus Sciaky, 1995

Subgenus Aphaonus Reitter, 1887

Pterostichus arcanoides (Lorenz, 1998)
Pterostichus compressus (Rost, 1892)
Pterostichus cylindriformis (Reitter, 1887)
Pterostichus koenigianus (Tschitscherine, 1891)
Pterostichus mariamae Zamotajlov; Solodovnikov & Fominykh, 2015
Pterostichus miroshnikovi (Zamotajlov, 1991)
Pterostichus neilgaimani Chaladze & Kalatozishvili, 2017
Pterostichus pseudopercus (Reitter, 1889)
Pterostichus starckianus (Reitter, 1887)
Pterostichus svetlanae Zamotajlov; Solodovnikov & Fominykh, 2015
Pterostichus trubilini Zamotajlov, 1997

Subgenus Argutor Dejean, 1821

Pterostichus aanistschenkoi O. & E.Berlov, 1999
Pterostichus akozyrevi O. & E.Berlov, 1999
Pterostichus bhadravati Jedlicka, 1965
Pterostichus chameleon (Motschulsky, 1866)
Pterostichus commutabilis (Motschulsky, 1866)
Pterostichus cryobioides (Chaudoir, 1868)
Pterostichus cursor (Dejean, 1828)
Pterostichus dulcis (Bates, 1883)
Pterostichus kerzhneri Lafer, 1983
Pterostichus leonisi Apfelbeck, 1904
Pterostichus praetermissus (Chaudoir, 1868)
Pterostichus sulcitarsis A.Morawitz, 1862
Pterostichus vernalis (Panzer, 1796)

Subgenus Asioplatysma Kryzhanovskij, 1968

Pterostichus capito (Tschitscherine, 1900)
Pterostichus darvazicus Kryzhanovskij, 1968
Pterostichus karateghinicus Mikhailov, 1972
Pterostichus klapperichi Jedlicka, 1956
Pterostichus medvedevi Kryzhanovskij & Mikhailov, 1972
Pterostichus mujahedeeni Savich, 1999
Pterostichus rufopiceus Heyden, 1890
Pterostichus talibani Savich, 1999

Subgenus Badistrinus Motschulsky, 1866

Pterostichus arrowi Jedlicka, 1936
Pterostichus arrowianus Jedlicka, 1938
Pterostichus bandotaro Tanaka, 1958
Pterostichus doris Jedlicka, 1962
Pterostichus haptoderoides Tschitscherine, 1889
Pterostichus kajimurai Habu & Tanaka, 1957
Pterostichus laticollis (Motschulsky, 1844)
Pterostichus procephalus Bates, 1873

Subgenus Bothriopterus Chaudoir, 1835

Pterostichus ademidovae O.Berlov, 1996
Pterostichus adstrictus Eschscholtz, 1823
Pterostichus aeneocupreus (Fairmaire, 1887)
Pterostichus bhunetansis Davies, 2004
Pterostichus commixtiformis Roubal, 1920
Pterostichus ferghanicus Kabak, 2001
Pterostichus lustrans LeConte, 1851
Pterostichus mariae (Lutshnik, 1921)
Pterostichus mutus (Say, 1823)
Pterostichus oblongopunctatus (Fabricius, 1787)
Pterostichus oregonus LeConte, 1861
Pterostichus pensylvanicus LeConte, 1873
Pterostichus quadrifoveolatus Letzner, 1852
Pterostichus subovatus (Motschulsky, 1861)
Pterostichus trinarius (Casey, 1918)
Pterostichus tropicalis (Bates, 1882)
Pterostichus zyzzovi O. & E.Berlov, 1996

Subgenus Calopterus Chaudoir, 1838

Pterostichus pilosus (Host, 1790)
Pterostichus selmanni (Duftschmid, 1812)

Subgenus Carllindrothius Habu, 1984

Pterostichus colonus (Bates, 1883)

Subgenus Cheporus Latreille, 1829

Pterostichus burmeisteri Heer, 1837
Pterostichus dissimilis (A. & G.B.Villa, 1833)
Pterostichus muehlfeldii (Duftschmid, 1812)
Pterostichus transversalis (Duftschmid, 1812)

Subgenus Chinapterus O.Berlov, 1998

Pterostichus lianhuaensis Dorjerem; Shi & Liang, 2020
Pterostichus liupanensis Dorjerem; Shi & Liang, 2020
Pterostichus przewalskyi Tschitscherine, 1888
Pterostichus singularis Tschitscherine, 1889

Subgenus Circinatus Sciaky, 1996

Pterostichus adelphus Shi & Liang, 2015
Pterostichus agilis Allegro & Sciaky, 2010
Pterostichus ailaoicus Shi & Liang, 2015
Pterostichus baenningeri Jedlicka, 1931
Pterostichus batxatensis Fedorenko, 2019
Pterostichus beneshi Sciaky, 1996
Pterostichus bullatus Allegro & Sciaky, 2010
Pterostichus camelus Shi & Liang, 2015
Pterostichus caobang Fedorenko, 2019
Pterostichus cavazzutianus Shi & Liang, 2015
Pterostichus dentifer Allegro & Sciaky, 2010
Pterostichus dimorphus Shi & Liang, 2015
Pterostichus expolitus Fedorenko, 2019
Pterostichus hoanglien Fedorenko, 2019
Pterostichus laocaiensis Fedorenko, 2019
Pterostichus liciniformis Csiki, 1930
Pterostichus maitreya Shi & Liang, 2015
Pterostichus miao Shi & Liang, 2015
Pterostichus phangxipang Fedorenko, 2019
Pterostichus phiaoc Fedorenko, 2019
Pterostichus pohnerti Jedlicka, 1934
Pterostichus subtilissimus Sciaky, 1996
Pterostichus tumulus Shi & Liang, 2015
Pterostichus wangjiani Shi & Liang, 2015
Pterostichus xilingensis Allegro & Sciaky, 2010
Pterostichus yan Shi & Liang, 2015
Pterostichus yuxiaodongi Shi & Liang, 2015
Pterostichus zhygealu Shi & Liang, 2015
Pterostichus zoiai Sciaky, 1996

Subgenus Cophosus Dejean, 1821

Pterostichus cylindricus (Herbst, 1784)

Subgenus Cryobius Chaudoir, 1838

Pterostichus abaxoides (Dejean, 1828)
Pterostichus acrogonus (Chaudoir, 1843)
Pterostichus akkusianus Kirschenhofer, 1981
Pterostichus altaiensis (Poppius, 1906)
Pterostichus amaroides (Dejean, 1828)
Pterostichus amblypterus (Chaudoir, 1868)
Pterostichus amoenus (Dejean, 1828)
Pterostichus amurensis (Poppius, 1906)
Pterostichus anatolicus Jedlicka, 1963
Pterostichus apenninus (Dejean, 1831)
Pterostichus aralarensis (Mateu, 1945)
Pterostichus archangaicus Shilenkov, 2000
Pterostichus arcticola (Chaudoir, 1868)
Pterostichus argutoriformis (Poppius, 1906)
Pterostichus auriga Ball, 1962
Pterostichus bargusinicus Shilenkov, 2000
Pterostichus barlensis Straneo, 1935
Pterostichus barryorum Ball, 1962
Pterostichus biocryus Ball, 1962
Pterostichus blandulus L.Miller, 1859
Pterostichus blumenthali Heinz, 1965
Pterostichus brevicornis (Kirby, 1837)
Pterostichus breviusculus (R.F.Sahlberg, 1844)
Pterostichus bryanti (Van Dyke, 1951)
Pterostichus bryantoides Ball, 1962
Pterostichus burjaticus (Poppius, 1906)
Pterostichus cacumensis Ball, 1966
Pterostichus cantabricus (L.Schaufuss, 1862)
Pterostichus caribou Ball, 1962
Pterostichus carradei (Gautier des Cottes, 1866)
Pterostichus champenoisi (Croissandeau, 1893)
Pterostichus chipewyan Ball, 1962
Pterostichus colasi (Jeannel, 1937)
Pterostichus convexus (Gebler, 1847)
Pterostichus coreicus Jedlicka, 1962
Pterostichus davshensis Shilenkov, 2000
Pterostichus dubiosus (Tschitscherine, 1894)
Pterostichus ehlersi (Heyden, 1881)
Pterostichus empetricola (Dejean, 1828)
Pterostichus espanoli (J. & E.Vives, 1977)
Pterostichus euxinus Straneo, 1935
Pterostichus exceptus (J.Sahlberg, 1885)
Pterostichus firmus (Lutshnik, 1933)
Pterostichus fulvescens (Motschulsky, 1844)
Pterostichus gerstlensis Ball, 1962
Pterostichus glacialis (Brisout de Barneville, 1863)
Pterostichus glukhomanka Sundukov, 2013
Pterostichus haftorni Lindroth, 1969
Pterostichus herzi (Poppius, 1906)
Pterostichus homalonotus (Tschitscherine, 1894)
Pterostichus horvatovichi Kirschenhofer, 1991
Pterostichus hudsonicus LeConte, 1863
Pterostichus infimus (Chaudoir, 1868)
Pterostichus inopinatus (Lutshnik, 1933)
Pterostichus iripennis (Chaudoir, 1868)
Pterostichus ishiharai Ishida & Shibata, 1961
Pterostichus jacobsoni (Poppius, 1906)
Pterostichus jaechi Kirschenhofer, 1997
Pterostichus josephi Csiki, 1930
Pterostichus kadjaranze (Jedlicka, 1947)
Pterostichus kaninensis (Poppius, 1906)
Pterostichus kavanaughi Shilenkov, 2000
Pterostichus kaynashensis Schweiger, 1967
Pterostichus kolymensis Erjiomin, 1998
Pterostichus korgei Jedlicka, 1964
Pterostichus kotzebuei Ball, 1962
Pterostichus kultianus Jedlicka, 1946
Pterostichus kuraicus Shilenkov, 2000
Pterostichus kurosai Tanaka, 1958
Pterostichus kurosawai Tanaka, 1958
Pterostichus lamuticus (Poppius, 1906)
Pterostichus latiusculus (Chaudoir, 1868)
Pterostichus longipes (Poppius, 1906)
Pterostichus lucidus (Motschulsky, 1844)
Pterostichus macedonicus Apfelbeck, 1918
Pterostichus macrothorax (Poppius, 1906)
Pterostichus maeklini (Poppius, 1906)
Pterostichus mandibularoides Ball, 1966
Pterostichus marani Jedlicka, 1967
Pterostichus middendorffi (J.Sahlberg, 1875)
Pterostichus negligens (Sturm, 1824)
Pterostichus nemoralis (Graells, 1851)
Pterostichus nigripalpis (Poppius, 1906)
Pterostichus nivalis (R.F.Sahlberg, 1844)
Pterostichus nordqvisti (J.Sahlberg, 1885)
Pterostichus oblongiusculus (Motschulsky, 1850)
Pterostichus ochoticus (R.F.Sahlberg, 1844)
Pterostichus palekhai E. & O.Berlov, 1996
Pterostichus parasimilis Ball, 1962
Pterostichus parviceps (Poppius, 1906)
Pterostichus pinguedineus (Eschscholtz, 1823)
Pterostichus pithyusicus (Lutshnik, 1933)
Pterostichus planus (J.Sahlberg, 1885)
Pterostichus poppiusianus (Jakobson, 1907)
Pterostichus procerulus (Heyden, 1880)
Pterostichus properans (Chaudoir, 1868)
Pterostichus pseudostuxbergi (Poppius, 1906)
Pterostichus pumilio (Dejean, 1828)
Pterostichus punctiger (J.Sahlberg, 1880)
Pterostichus pusillus (Dejean, 1828)
Pterostichus quadraticollis (Chaudoir, 1846)
Pterostichus riparius (Dejean, 1828)
Pterostichus rodiovoni Shilenkov, 2000
Pterostichus rufonitens (Fairmaire, 1866)
Pterostichus safonovae Budarin, 1995
Pterostichus sahlbergi (Tschitscherine, 1894)
Pterostichus scitus (Mäklin, 1878)
Pterostichus shilenkovi Erjiomin & Kabak, 1991
Pterostichus similis Mannerheim, 1852
Pterostichus sojot Shilenkov, 2000
Pterostichus soperi Ball, 1966
Pterostichus splendidus Park, 2013
Pterostichus spornyi Sundukov, 2013
Pterostichus stantonensis Ball, 1966
Pterostichus subgibbus (Motschulsky, 1860)
Pterostichus subiasi (Ortuño & Zaballos, 1992)
Pterostichus subsinuatus (Dejean, 1828)
Pterostichus surgens LeConte, 1878
Pterostichus tareumiut Ball, 1962
Pterostichus taskylensis Shilenkov, 2000
Pterostichus tatricus Kult, 1947
Pterostichus theeli (Mäklin, 1878)
Pterostichus tichomirovi Erjiomin, 1990
Pterostichus tiliaceoradix Ball, 1962
Pterostichus tokmakovae Sundukov, 2013
Pterostichus tunkinensis Shilenkov, 2000
Pterostichus udokanensis Shilenkov, 2000
Pterostichus ulubeyensis Kirschenhofer, 1981
Pterostichus unctulatus (Duftschmid, 1812)
Pterostichus ventricosus (Eschscholtz, 1823)
Pterostichus woodi Ball & Currie, 1997

Subgenus Cylindrocharis Casey, 1918

Pterostichus acutipes Barr, 1971
Pterostichus hypogeus Barr, 1971
Pterostichus rostratus (Newman, 1838)

Subgenus Eosteropus Tschitscherine, 1902

Pterostichus abensis Morita, 2010
Pterostichus abudarini O. & E.Berlov, 1997
Pterostichus aenescens (Chaudoir, 1850)
Pterostichus aereipennis (Solsky, 1872)
Pterostichus aethiops (Panzer, 1796)
Pterostichus ainus Plutenko, 2005
Pterostichus alacer A.Morawitz, 1862
Pterostichus bituberculatus (Tschitscherine, 1899)
Pterostichus circulosus Lindroth, 1966
Pterostichus coruscus (Tschitscherine, 1895)
Pterostichus creper (Tschitscherine, 1902)
Pterostichus discrepans A.Morawitz, 1862
Pterostichus dudkoi Sundukov, 2013
Pterostichus faldermanni Schatzmayr, 1929
Pterostichus funakoshii Morita, 2007
Pterostichus hidanus Morita, 2010
Pterostichus hiramatsui Morita, 2010
Pterostichus hirasawai Morita, 2010
Pterostichus imurai Morita, 2007
Pterostichus japonicus (Motschulsky, 1861)
Pterostichus karasawai Tanaka, 1958
Pterostichus kimioi Morita, 2011
Pterostichus makolskii G.Müller, 1933
Pterostichus mandzhuricus (Lutshnik, 1916)
Pterostichus mannerheimii (Dejean, 1831)
Pterostichus matsunagai Morita, 2011
Pterostichus maurusiacus (Mannerheim, 1825)
Pterostichus mizunoyai Morita, 2007
Pterostichus moestus (Say, 1823)
Pterostichus nasuensis Morita, 2007
Pterostichus noburai Morita, 2008
Pterostichus ohkawai Morita, 2010
Pterostichus orientalis (Motschulsky, 1844)
Pterostichus rengensis Morita, 2010
Pterostichus rufitarsis (Dejean, 1828)
Pterostichus sakuraii Morita, 2008
Pterostichus shimizui Morita, 2010
Pterostichus sudai Morita, 1991
Pterostichus superciliosus (Say, 1823)
Pterostichus tenuimarginatus (Chaudoir, 1868)
Pterostichus tokui Morita, 2007
Pterostichus tuberculiger (Tschitscherine, 1897)
Pterostichus virescens (Gebler, 1833)
Pterostichus yoshizawai Morita, 2010

Subgenus Ethira Andrewes, 1936

Pterostichus cometes (Andrewes, 1936)
Pterostichus depilatus (Bates, 1889)
Pterostichus heinzianus Sciaky, 1996
Pterostichus multiseta (Straneo, 1984)
Pterostichus piliferus (Bates, 1878)
Pterostichus pseudopilifer (Straneo, 1957)
Pterostichus rugiceps (Straneo, 1984)
Pterostichus seticeps (Straneo, 1984)
Pterostichus sharanus (Straneo, 1984)
Pterostichus variseta (Straneo, 1984)
Pterostichus viridellus Straneo, 1984

Subgenus Falsargutor Kryzhanovskij, 1983

Pterostichus kataevi Kryzhanovskij, 1989
Pterostichus ponticus Kirschenhofer, 1987
Pterostichus pseudopedius Reitter, 1887

Subgenus Feronidius Jeannel, 1942

Pterostichus brevipennis (Chevrolat, 1840)
Pterostichus fornicatus (Kolenati, 1845)
Pterostichus hungaricus (Dejean, 1828)
Pterostichus incommodus Schaum, 1858
Pterostichus melas (Creutzer, 1799)

Subgenus Feronina Casey, 1918

Pterostichus barri Bousquet, 2006
Pterostichus palmi Schaeffer, 1910

Subgenus Fukienostichus Lafer, 1983

Pterostichus shaovuensis Jedlicka, 1956

Subgenus Gastrosticta Casey, 1918

Pterostichus enodis Bousquet, 1992
Pterostichus mutoides Bousquet, 1992
Pterostichus obesulus LeConte, 1873
Pterostichus ophryoderus (Chaudoir, 1878)
Pterostichus punctiventris (Chaudoir, 1878)
Pterostichus putus Casey, 1913
Pterostichus sayanus Csiki, 1930
Pterostichus subacutus (Casey, 1918)
Pterostichus tumescens LeConte, 1863
Pterostichus ventralis (Say, 1823)

Subgenus Georgeballius Habu, 1984

Pterostichus hoplites (Bates, 1883)

Subgenus Gutta Wrase & J.Schmidt, 2006

Pterostichus adulterinus Wrase & J.Schmidt, 2006
Pterostichus gaoligongensis Wrase & J.Schmidt, 2006
Pterostichus kongshuhensis B.Gueorguiev, 2015
Pterostichus phungaraziensis Wrase & J.Schmidt, 2006

Subgenus Haplomaseus Reitter, 1896

Pterostichus abagonensis Reitter, 1896
Pterostichus andreae (Tschitscherine, 1897)
Pterostichus arator (Faldermann, 1836)
Pterostichus armenus (Faldermann, 1836)
Pterostichus astutus (Tschitscherine, 1903)
Pterostichus balikliensis Hovorka & Skoupy, 2007
Pterostichus caucasicola (Tschitscherine, 1893)
Pterostichus caucasicus (Ménétriés, 1832)
Pterostichus chasautianus Vysoky, 1984
Pterostichus chydaeus (Tschitscherine, 1897)
Pterostichus consanguineus (Chaudoir, 1878)
Pterostichus cristicaudis Kurnakov, 1962
Pterostichus denticaudis Kurnakov, 1962
Pterostichus giresuni Jedlicka, 1965
Pterostichus goriensis (Tschitscherine, 1897)
Pterostichus heinzi Jedlicka, 1965
Pterostichus ikizderensis Hovorka & Skoupy, 2007
Pterostichus jugicola (Lutshnik, 1916)
Pterostichus krulikovskyi (Lutshnik, 1933)
Pterostichus meskheticus Belousov, 1991
Pterostichus novotnyorum Vysoky, 1981
Pterostichus olegi Belousov, 1991
Pterostichus phaeus (Lutshnik, 1922)
Pterostichus porcellus Kurnakov, 1962
Pterostichus pseudoastutus Hovorka & Skoupy, 2007
Pterostichus rousi Kirschenhofer, 1982
Pterostichus skoupyi Hovorka & Skoupy, 2007
Pterostichus staveni Hovorka & Skoupy, 2007
Pterostichus tamsii (Dejean, 1831)
Pterostichus tanjae Hovorka & Skoupy, 2007
Pterostichus woronowi (Lutshnik, 1916)

Subgenus Haptotapinus Reitter, 1886

Pterostichus aksekianus (Straneo, 1988)
Pterostichus besucheti (Straneo, 1988)
Pterostichus bythiniensis (Maran, 1944)
Pterostichus crassiusculus (Chaudoir, 1868)
Pterostichus dipojranus (Straneo, 1988)
Pterostichus matchai (Jedlicka, 1930)
Pterostichus oblongoparallelus Maran, 1932
Pterostichus odontocnemis G.Müller, 1931
Pterostichus orduensis (Straneo, 1988)
Pterostichus pavani (Straneo, 1988)
Pterostichus reissi G.Müller, 1931
Pterostichus vseteckai (Maran, 1944)

Subgenus Huaius Tian; Huang; Chen & Ding, 2019

Pterostichus caoi Tian; Huang; Chen & Ding, 2019
Pterostichus hanwang Tian & He, 2020
Pterostichus tiankeng Tian; Huang; Chen & Ding, 2019
Pterostichus yuae Tian; Huang; Chen & Ding, 2019

Subgenus Hypherpes Chaudoir, 1838

Pterostichus adoxus (Say, 1823)
Pterostichus algidus LeConte, 1853
Pterostichus amethystinus Mannerheim, 1843
Pterostichus annosus Casey, 1913
Pterostichus arcanus Casey, 1913
Pterostichus baldwini (Casey, 1924)
Pterostichus barbarinus Casey, 1913
Pterostichus brachylobus Kavanaugh & LaBonte, 2006
Pterostichus californicus (Dejean, 1828)
Pterostichus canallatus Casey, 1913
Pterostichus castaneus (Dejean, 1828)
Pterostichus castanipes (Ménétriés, 1843)
Pterostichus congestus (Ménétriés, 1843)
Pterostichus craterensis (Hatch, 1949)
Pterostichus crenicollis LeConte, 1873
Pterostichus ecarinatus Hatch, 1936
Pterostichus esuriens Casey, 1913
Pterostichus gliscans Casey, 1913
Pterostichus gracilior LeConte, 1873
Pterostichus herculaneus Mannerheim, 1843
Pterostichus hornii LeConte, 1873
Pterostichus illustris LeConte, 1851
Pterostichus inermis Fall, 1901
Pterostichus isabellae LeConte, 1851
Pterostichus jacobinus Casey, 1913
Pterostichus laborans Casey, 1913
Pterostichus lacertus Casey, 1913
Pterostichus lama (Ménétriés, 1843)
Pterostichus lassulus (Casey, 1920)
Pterostichus lattini LaBonte, 2006
Pterostichus luscus (Casey, 1918)
Pterostichus menetriesii LeConte, 1873
Pterostichus mercedianus (Casey, 1918)
Pterostichus miscellus Casey, 1913
Pterostichus morionides (Chaudoir, 1868)
Pterostichus neobrunneus Lindroth, 1966
Pterostichus nigrocaeruleus Van Dyke, 1926
Pterostichus obsidianus Casey, 1913
Pterostichus occultus Casey, 1913
Pterostichus ordinarius Casey, 1913
Pterostichus ovalipennis Casey, 1913
Pterostichus panticulatus Casey, 1913
Pterostichus pergracilis (Casey, 1920)
Pterostichus planctus LeConte, 1853
Pterostichus protensiformis (Casey, 1924)
Pterostichus protractus LeConte, 1860
Pterostichus restrictus (Casey, 1918)
Pterostichus scutellaris LeConte, 1873
Pterostichus sejungendus (Chaudoir, 1868)
Pterostichus serripes (LeConte, 1875)
Pterostichus setosus Hatch, 1951
Pterostichus sierranus Casey, 1913
Pterostichus sponsor Casey, 1913
Pterostichus spraguei LeConte, 1873
Pterostichus suffusus Casey, 1913
Pterostichus tarsalis LeConte, 1873
Pterostichus tristis (Dejean, 1828)
Pterostichus tuberculofemoratus Hatch, 1936
Pterostichus vandykei Schaeffer, 1910
Pterostichus vicinus Mannerheim, 1843
Pterostichus ybousqueti O.Berlov, 1999

Subgenus Jedlickaia Sciaky, 1997

Pterostichus sterbai Jedlicka, 1934

Subgenus Koreonialoe Park & Kwon, 1996

Pterostichus adatarasanus Sasakawa, 2005
Pterostichus akitai Morita, 2004
Pterostichus apiculatiphallus Nemoto, 1988
Pterostichus asahinus Habu & Baba, 1960
Pterostichus bellatrix (Tschitscherine, 1895)
Pterostichus bifidiphallus Nemoto, 1988
Pterostichus chokaisanus Sasakawa, 2009
Pterostichus eboshiyamanus Sasakawa, 2009
Pterostichus falcispinus Sasakawa, 2005
Pterostichus fukube Sugimura, 2005
Pterostichus gassanus Sasakawa, 2009
Pterostichus gujoensis Toda, 2012
Pterostichus ishikawai Nemoto, 1988
Pterostichus ishikawaioides Sasakawa; J.L.Kim; J.K.Kim & Kubota, 2008
Pterostichus isolatus Sasakawa; Kim & Kubota, 2005
Pterostichus iwakiensis Sasakawa, 2009
Pterostichus iwasakii Kasahara, 1990
Pterostichus ixion (Tschitscherine, 1902)
Pterostichus jiricola Sasakawa; J.L.Kim; J.K.Kim & Kubota, 2008
Pterostichus kitakamisanus Sasakawa, 2005
Pterostichus koheii Nakane, 1963
Pterostichus kuraiyamanus Morita & Ohkawa, 2010
Pterostichus kurikomasanus Sasakawa, 2005
Pterostichus kuwolsanensis Li & Zhang, 2014
Pterostichus macrogenys Bates, 1883
Pterostichus microps Heyden, 1887
Pterostichus miyazawai Morita & Ohkawa, 2009
Pterostichus momuranus Morita; Ohkawa & Kurihara, 2013
Pterostichus monolineatus Sasakawa; Mitsuduka & H.Ito, 2020
Pterostichus nagasawai N.Ito & Ogai, 2015
Pterostichus ohsawacavus Sasakawa, 2005
Pterostichus opacipennis Jedlicka, 1934
Pterostichus palgongsanus Nemoto, 1988
Pterostichus shikatai Toda, 2012
Pterostichus shinbodakensis Sasakawa & H.Ito, 2017
Pterostichus shirakamisan Sasakawa, 2009
Pterostichus shirakamisanus Sasakawa, 2005
Pterostichus shojii Sugimura, 2006
Pterostichus sumondakensis Sasakawa, 2005
Pterostichus syleus Kirschenhofer, 1997
Pterostichus taebaegsanus Nemoto, 1988
Pterostichus takadateyamanus Sasakawa, 2009
Pterostichus tanakai Ishida, 1964
Pterostichus tateishiyamanus Sasakawa & H.Ito, 2017
Pterostichus teretis Park & Kwon, 1996
Pterostichus todai Morita & Kanie, 1997
Pterostichus togyusanus Park & Kwon, 1996
Pterostichus toyodai Morita & Kurosa, 1998
Pterostichus uedaorum Morita & Hirasawa, 1996
Pterostichus vicinus Park & Kwon, 1996
Pterostichus woongbii Park & Kwon, 1996
Pterostichus yahikosanus Sasakawa, 2009
Pterostichus yamizosanus Sasakawa, 2005
Pterostichus yorikoae Sugimura, 2007

Subgenus Kozyrevius O.Berlov, 1998

Pterostichus militaris (Tschitscherine, 1893)

Subgenus Lamenius Bousquet, 1999

Pterostichus caudicalis (Say, 1823)

Subgenus Lenapterus O.Berlov, 1996

Pterostichus agonus G.Horn, 1880
Pterostichus cancellatus (Motschulsky, 1860)
Pterostichus costatus (Ménétriés, 1851)
Pterostichus galae Farkac & Plutenko, 1996
Pterostichus gromykoi Sundukov, 2005
Pterostichus marginatus Matsumura, 1911
Pterostichus punctatissimus (Randall, 1838)
Pterostichus rugosipennis Jedlicka, 1932
Pterostichus saxicola (Tschitscherine, 1899)
Pterostichus vermiculosus (Ménétriés, 1851)

Subgenus Leptoferonia Casey, 1918

Pterostichus angustus (Dejean, 1828)
Pterostichus beyeri Van Dyke, 1926
Pterostichus blodgettensis Will, 2007
Pterostichus caligans G.Horn, 1891
Pterostichus cochlearis Hacker, 1968
Pterostichus deino Will, 2007
Pterostichus enyo Will, 2007
Pterostichus falli Van Dyke, 1926
Pterostichus fenyesi Csiki, 1930
Pterostichus fuchsi Schaeffer, 1910
Pterostichus hackerae Will, 2016
Pterostichus hatchi Hacker, 1968
Pterostichus humilis Casey, 1913
Pterostichus idahoae Csiki, 1930
Pterostichus inanis G.Horn, 1891
Pterostichus infernalis Hatch, 1936
Pterostichus inopinus (Casey, 1918)
Pterostichus lobatus Hacker, 1968
Pterostichus marinensis Hacker, 1968
Pterostichus mattolensis Hacker, 1968
Pterostichus pemphredo Will, 2007
Pterostichus pumilus Casey, 1913
Pterostichus rothi (Hatch, 1951)
Pterostichus sphodrinus LeConte, 1863
Pterostichus stapedius Hacker, 1968
Pterostichus trinitensis Hacker, 1968
Pterostichus yosemitensis Hacker, 1968

Subgenus Lianoe Gozis, 1882

Pterostichus amanoi Nakane, 1968
Pterostichus arribasi Ortuño, 1991
Pterostichus asturicus (Jeanne, 1966)
Pterostichus cordatissimus (Straneo, 1937)
Pterostichus drescoi Nègre, 1957
Pterostichus dufourii (Dejean, 1828)
Pterostichus microphthalmus Delarouzée, 1857
Pterostichus muelleri (Straneo, 1936)
Pterostichus nadari (Vuillefroy, 1893)
Pterostichus nakamiyorinus Morita; Ohkawa & Kurihara, 2013
Pterostichus rousselli Colas, 1963
Pterostichus sakagamii Morita, 2001
Pterostichus sudrei Sainte-Claire Deville, 1922

Subgenus Lyrothorax Chaudoir, 1838

Pterostichus amagisanus Tanaka & Ishida, 1972
Pterostichus caspius (Ménétriés, 1832)
Pterostichus eoyoritomus Sasakawa, 2009
Pterostichus fujitai Tanaka & Ishida, 1972
Pterostichus yoritomus Bates, 1873

Subgenus Megabea Sciaky, 1997

Pterostichus tienmushanus Sciaky, 1997

Subgenus Melanius Bonelli, 1810

Pterostichus aterrimus (Herbst, 1784)
Pterostichus castor Goulet & Bousquet, 1983
Pterostichus corvinus (Dejean, 1828)
Pterostichus ebeninus (Dejean, 1828)
Pterostichus elongatus (Duftschmid, 1812)
Pterostichus licenti Jedlicka, 1939
Pterostichus noguchii Bates, 1873

Subgenus Metallophilus Chaudoir, 1838

Pterostichus dandongensis Kirschenhofer, 1997
Pterostichus interruptus (Dejean, 1828)
Pterostichus kamtschaticus Motschulsky, 1860
Pterostichus mirus (Tschitscherine, 1894)
Pterostichus orion (Tschitscherine, 1901)
Pterostichus pfizenmayeri Poppius, 1906
Pterostichus rugosus (Gebler, 1823)
Pterostichus sublaevis (J.Sahlberg, 1880)

Subgenus Micronialoe Park; Kwon & Lafer, 1996

Pterostichus bifoveolatus Park; Kwon & Lafer, 1996
Pterostichus chogyesanus Park; Kwon & Lafer, 1996
Pterostichus kaniei Morita, 2005

Subgenus Monoferonia Casey, 1918

Pterostichus carolinus Darlington, 1932
Pterostichus diligendus (Chaudoir, 1868)
Pterostichus mancus (LeConte, 1853)
Pterostichus primus Darlington, 1932

Subgenus Morphohaptoderus Tschitscherine, 1898

Pterostichus bowanus Sciaky, 1997
Pterostichus cervenkai Sciaky, 1994
Pterostichus chungkingi Jedlicka, 1932
Pterostichus confucius Sciaky & Wrase, 1997
Pterostichus dentellus Facchini & Sciaky, 2003
Pterostichus dundai Sciaky, 1994
Pterostichus emei Sciaky, 1994
Pterostichus expeditus (Tschitscherine, 1898)
Pterostichus geberti Sciaky & Wrase, 1997
Pterostichus giacomazzoi Sciaky, 1994
Pterostichus gongga Sciaky, 1997
Pterostichus guizhouensis Sciaky, 1997
Pterostichus huashanus Sciaky, 1994
Pterostichus hubeicus Facchini & Sciaky, 2003
Pterostichus irideus Sciaky, 1994
Pterostichus janatai Sciaky & Wrase, 1997
Pterostichus kalabi Sciaky, 1994
Pterostichus kucerai Sciaky, 1997
Pterostichus lingshanus Sciaky & Wrase, 1997
Pterostichus maximus Tschitscherine, 1889
Pterostichus megaloderus Sciaky, 1994
Pterostichus ming Sciaky & Wrase, 1997
Pterostichus miroslavi Sciaky & Wrase, 1997
Pterostichus muellermotzfeldi Wrase & J.Schmidt, 2006
Pterostichus parvicollis Sciaky & Wrase, 1997
Pterostichus pseudoplatyderus Sciaky, 1994
Pterostichus saueri Sciaky, 1994
Pterostichus schuelkei Sciaky & Wrase, 1997
Pterostichus shennongjianus Facchini & Sciaky, 2003
Pterostichus straneellus Jedlicka, 1938
Pterostichus toledanoi Facchini & Sciaky, 2003
Pterostichus wenxianensis Allegro & Sciaky, 2010
Pterostichus yulongshanensis Sciaky, 1997

Subgenus Myosodus Fischer von Waldheim, 1823

Pterostichus aapsorum Belousov, 1991
Pterostichus aibgensis Starck, 1890
Pterostichus avaricus Abdurakhmanov & Kryzhanovskij, 1983
Pterostichus batesi (Tschitscherine, 1894)
Pterostichus buglaniensis Kirschenhofer, 1981
Pterostichus filipjevi (Lutshnik, 1928)
Pterostichus ingusha (Lutshnik, 1928)
Pterostichus kiritshenkoi (Lutshnik, 1928)
Pterostichus lacunosus (Chaudoir, 1844)
Pterostichus lutshnikianus Bogachev & Kurnakov, 1958
Pterostichus nivicola (Ménétriés, 1832)
Pterostichus ordinatus (Fischer von Waldheim, 1823)
Pterostichus rudestriatus (Reitter, 1883)
Pterostichus schoenherri Faldermann, 1836
Pterostichus sodalicius Heyden, 1885
Pterostichus starcki Heyden, 1885
Pterostichus stoeckleini (Straneo, 1941)
Pterostichus svanicus (Lutshnik, 1922)
Pterostichus swaneticus (Reitter, 1883)
Pterostichus variabilis (Ménétriés, 1832)
Pterostichus zamotajlovi Belousov, 1991

Subgenus Neohaptoderus Tschitscherine, 1898

Pterostichus berezowskii (Tschitscherine, 1898)
Pterostichus catei Sciaky & Wrase, 1997
Pterostichus comorus Jedlicka, 1932
Pterostichus gravis Jedlicka, 1939
Pterostichus haesitatus Fairmaire, 1889
Pterostichus ignavus (Tschitscherine, 1897)
Pterostichus jureceki (Jedlicka, 1936)
Pterostichus kleinfeldianus Sciaky & Wrase, 1997
Pterostichus komalus (Jedlicka, 1936)
Pterostichus maderi Jedlicka, 1938
Pterostichus molopsoides Jedlicka, 1934
Pterostichus montigena (Tschitscherine, 1898)
Pterostichus mundus Jedlicka, 1938
Pterostichus oreophilus (Tschitscherine, 1898)
Pterostichus orestes (Jedlicka, 1936)
Pterostichus sinicus (Tschitscherine, 1897)
Pterostichus szetschuanensis Tschitscherine, 1889
Pterostichus yunnanensis Jedlicka, 1934

Subgenus Nialoe Tanaka, 1958

Pterostichus asymmetricus Bates, 1883
Pterostichus basilobatus Sasakawa, 2005
Pterostichus biexcisus Straneo, 1955
Pterostichus bisetosus (Straneo, 1938)
Pterostichus brunneipennis Straneo, 1955
Pterostichus carsticus Kasahara & Y.Ito, 1989
Pterostichus chujoi Habu, 1959
Pterostichus cristatoides Straneo, 1955
Pterostichus daihizanus Ishida, 1968
Pterostichus daisenicus Ishida, 1958
Pterostichus dandonis Kasahara, 1989
Pterostichus enasanus Morita, 2007
Pterostichus fujimurai Habu, 1958
Pterostichus fujisanus Tanaka & Suga, 1972
Pterostichus fumikoae Kurosa, 2019
Pterostichus hakusanus Kasahara, 1989
Pterostichus hekosanensis Sasakawa, 2020
Pterostichus himifuriho Morita, 2007
Pterostichus hiraii Morita, 2015
Pterostichus hirobane Habu & Baba, 1958
Pterostichus hozumii Ishida, 1961
Pterostichus ikukoae Morita, 2007
Pterostichus ishizukai Kasahara, 1995
Pterostichus isumiensis Kasahara & Saito, 1997
Pterostichus janoi Jedlicka, 1952
Pterostichus jogaesanensis Lafer; Paik & Park, 1996
Pterostichus katashinensis Habu, 1958
Pterostichus kaya Morita, 2019
Pterostichus kongosanus Nakane, 1963
Pterostichus latistylis Tanaka, 1958
Pterostichus masahiroi Kasahara, 1988
Pterostichus masatakai Morita, 2007
Pterostichus mirificus Bates, 1883
Pterostichus mitoyamanus Tanaka, 1971
Pterostichus musashiensis Kasahara, 1993
Pterostichus nakanei Straneo, 1955
Pterostichus naokii Morita, 2003
Pterostichus napaea Kasahara, 1988
Pterostichus nishiyamai Kasahara, 1986
Pterostichus ogaensis Morita, 1995
Pterostichus ohbayashii Ishida, 1968
Pterostichus ohdaisanus Nakane, 1963
Pterostichus ohkurai Morita, 1996
Pterostichus okutamae Tanaka, 1963
Pterostichus omogoensis Nakane, 1972
Pterostichus ompoensis Jedlicka, 1932
Pterostichus opaculus N.Ito, 2010
Pterostichus ovaliphallus Sasakawa, 2005
Pterostichus parkwon Davies, 2004
Pterostichus praedo (Tschitscherine, 1901)
Pterostichus rhanis (Tschitscherine, 1902)
Pterostichus ryoheii Morita, 2012
Pterostichus ryomoensis Morita & Suda, 2007
Pterostichus satoi Ishida, 1961
Pterostichus shibatai Ishida, 1961
Pterostichus shotaroi Morita, 1987
Pterostichus sincerus Park & Kwon, 1996
Pterostichus spiculifer Bates, 1883
Pterostichus sugimurai Morita, 2007
Pterostichus tahirai Kasahara, 1992
Pterostichus takahashii Ishida, 1958
Pterostichus tanakaorum Morita & Ohkura, 1988
Pterostichus taoi Kasahara, 1993
Pterostichus tokejii Yoshida & Tanaka, 1960
Pterostichus tottoriensis Morita, 2003
Pterostichus tsurugiyamanus Habu, 1959
Pterostichus uchiyamai Morita, 1987
Pterostichus uenoi Straneo, 1955
Pterostichus watanabei Nakane, 1960
Pterostichus yamashitai Morita, 2019
Pterostichus yanoi Jedlicka, 1953
Pterostichus yatsuensis Straneo, 1955
Pterostichus yokohamae Straneo in Nakane, 1979

Subgenus Oreolyperus Tschitscherine, 1901

Pterostichus heptapotamicus (Lutshnik, 1927)
Pterostichus korolkowi (Tschitscherine, 1901)
Pterostichus necessarius (Tschitscherine, 1894)
Pterostichus regeli (Tschitscherine, 1894)

Subgenus Oreophilus Chaudoir, 1838

Pterostichus alberti (Jeannel, 1942)
Pterostichus cribratus (Dejean, 1828)
Pterostichus dubius Heer, 1838
Pterostichus duratii A. & G.B.Villa, 1835
Pterostichus externepunctatus (Dejean, 1828)
Pterostichus flavofemoratus (Dejean, 1828)
Pterostichus franzi Nègre, 1955
Pterostichus ienishteai Nitzu, 1988
Pterostichus impressus (Fairmaire & Laboulbène, 1854)
Pterostichus jurinei (Panzer, 1802)
Pterostichus morio (Duftschmid, 1812)
Pterostichus multipunctatus (Dejean, 1828)
Pterostichus parnassius Schaum, 1859
Pterostichus paulini (Vuillefroy, 1868)
Pterostichus planiusculus (Chaudoir, 1859)
Pterostichus rufipennis Baudi di Selve, 1889
Pterostichus spinolae (Dejean, 1828)
Pterostichus variolatus (Dejean, 1828)
Pterostichus xatartii (Dejean, 1828)
Pterostichus yvanii (Dejean, 1828)

Subgenus Oreoplatysma Jakobson, 1907

Pterostichus abishirensis Belousov, 1991
Pterostichus araraticus Kirschenhofer, 1987
Pterostichus asinubas Davies, 2004
Pterostichus belizini (Lutshnik, 1933)
Pterostichus belousovi Kryzhanovskij, 1989
Pterostichus borcka Jedlicka, 1963
Pterostichus capitolinus Kurnakov, 1962
Pterostichus casaleianus Kirschenhofer, 1991
Pterostichus cecchiniae (Jakobson, 1907)
Pterostichus chefsuricus Reitter, 1896
Pterostichus colchicus (Chaudoir, 1850)
Pterostichus cordifer Reitter, 1896
Pterostichus cunibakus Davies, 2004
Pterostichus daghestanus Reitter, 1896
Pterostichus depressidorsis Reitter, 1896
Pterostichus faunus Kurnakov, 1962
Pterostichus fencli Dvorak, 1995
Pterostichus ghilarovi Kryzhanovskij, 1988
Pterostichus jakobsonianus (Lutshnik, 1928)
Pterostichus kadleci Dvorak, 1995
Pterostichus kirschenblatti Kryzhanovskij, 1988
Pterostichus koenigi (Reitter, 1887)
Pterostichus krasnopolensis Kirschenhofer, 1987
Pterostichus kvirensis Belousov, 1991
Pterostichus lodosi Heinz, 1977
Pterostichus mediocris Dvorak, 1995
Pterostichus nikodymi Dvorak, 1995
Pterostichus odvarkai Dvorak, 1995
Pterostichus orman Jedlicka, 1963
Pterostichus pasanauricus Kirschenhofer, 1987
Pterostichus percontator Reitter, 1887
Pterostichus planaticollis Kirschenhofer, 1987
Pterostichus platyderus (Chaudoir, 1850)
Pterostichus pulchellus (Faldermann, 1836)
Pterostichus rousianus Kirschenhofer, 1987
Pterostichus rubripalpis Csiki, 1930
Pterostichus satunini (Tschitscherine, 1903)
Pterostichus satyrus Kurnakov, 1962
Pterostichus schodaicus Kirschenhofer, 1987
Pterostichus setosicus Davies, 2004
Pterostichus sojaki Kirschenhofer, 1987
Pterostichus stomoides (Chaudoir, 1868)
Pterostichus strasseri Reitter, 1898
Pterostichus tamarae Wrase & Kirschenhofer, 1991
Pterostichus validiceps Reitter, 1887
Pterostichus zolotarewi Reitter, 1911

Subgenus Orientostichus Sciaky & Allegro, 2013

Pterostichus chinensis (Jedlicka, 1962)
Pterostichus curtatus Fairmaire, 1886
Pterostichus davidi (Tschitscherine, 1897)
Pterostichus deceptrix (Tschitscherine, 1898)
Pterostichus distinctissimus Jedlicka, 1940
Pterostichus ferreroi Straneo, 1989
Pterostichus gallopavo Sciaky & Wrase, 1997
Pterostichus lesticoides (Straneo, 1939)
Pterostichus machulkai Jedlicka, 1931
Pterostichus perlutus Jedlicka, 1938
Pterostichus prattii Bates, 1890
Pterostichus pulcher Sciaky & Allegro, 2013
Pterostichus semirugosus (Andrewes, 1947)
Pterostichus simillimus Fairmaire, 1886
Pterostichus tachongi Jedlicka, 1936

Subgenus Orsonjohnsonus Hatch, 1933

Pterostichus johnsoni Ulke, 1889

Subgenus Paraferonia Casey, 1918

Pterostichus lubricus LeConte, 1853

Subgenus Parahaptoderus Jeanne, 1969

Pterostichus bielzii (Fuss, 1858)
Pterostichus brevis (Duftschmid, 1812)
Pterostichus vecors (Tschitscherine, 1897)

Subgenus Paralianoe Ishida, 1958

Pterostichus kiiensis Morita & Ohkura, 1988

Subgenus Parapterostichus Desbrochers des Loges, 1906

Pterostichus grajus (Dejean, 1828)
Pterostichus justusii W.Redtenbacher, 1842
Pterostichus nodicornis (Fairmaire & Laboulbène, 1854)
Pterostichus schaschli (Marseul, 1880)

Subgenus Petrophilus Chaudoir, 1838

Pterostichus abnormis (J.Sahlberg, 1880)
Pterostichus acutidens (Fairmaire, 1888)
Pterostichus alexandrovi Lafer, 1979
Pterostichus alexeji Zamotajlov & Kryzhanovskij, 1992
Pterostichus altaianus Jedlicka, 1958
Pterostichus altaicus (Germar, 1823)
Pterostichus arsenjevi Lafer, 1979
Pterostichus bungei (Tschitscherine, 1894)
Pterostichus calvitarsis Breit, 1912
Pterostichus chechcirensis Lafer, 1979
Pterostichus chinkiangensis Kirschenhofer, 1991
Pterostichus coracinus (Newman, 1838)
Pterostichus dauricus (Gebler, 1832)
Pterostichus dilutipes (Motschulsky, 1844)
Pterostichus elmbergi Poppius, 1908
Pterostichus eximius A.Morawitz, 1862
Pterostichus fallettii Sciaky, 1996
Pterostichus findelii (Dejean, 1828)
Pterostichus foveolatus (Duftschmid, 1812)
Pterostichus glaferi O. & E.Berlov, 1996
Pterostichus iberlovi O.Berlov, 1996
Pterostichus jungens (Tschitscherine, 1893)
Pterostichus jurecekianus Maran, 1940
Pterostichus kokeilii L.Miller, 1850
Pterostichus kurentzovi Lafer, 1979
Pterostichus kvanmobongus Lafer, 2011
Pterostichus lachrymosus (Newman, 1838)
Pterostichus levadensis Lafer, 1979
Pterostichus magoides (Straneo, 1937)
Pterostichus magus (Mannerheim, 1825)
Pterostichus maryseae Sun & Shi, 2018
Pterostichus melanarius (Illiger, 1798)
Pterostichus melanodes (Chaudoir, 1878)
Pterostichus mellyi (Gebler, 1843)
Pterostichus montanus (Motschulsky, 1844)
Pterostichus monticoloides Shilenkov in Kryzhanovskij et al., 1995
Pterostichus mroczkowskii Lafer, 2011
Pterostichus nigellus A.Morawitz, 1862
Pterostichus novus Straneo, 1944
Pterostichus odaesanensis Lafer, 2011
Pterostichus pawlowskii Lafer, 2011
Pterostichus pertinax (Tschitscherine, 1895)
Pterostichus petulans Jedlicka, 1938
Pterostichus poppiusi (Semenov, 1906)
Pterostichus probus Park & Kwon, 1996
Pterostichus procax A.Morawitz, 1862
Pterostichus rasilis Park & Kwon, 1996
Pterostichus relictus (Newman, 1838)
Pterostichus robustistylis Sasakawa, 2005
Pterostichus schoenmanni Kirschenhofer, 1991
Pterostichus sejunctus Bates, 1883
Pterostichus septentrionis (Chaudoir, 1868)
Pterostichus seriatus (Chaudoir, 1850)
Pterostichus seungmoi Park & Kwon, 1996
Pterostichus shingarevi Lafer, 1979
Pterostichus silvestris Sun & Shi, 2018
Pterostichus songoricus (Motschulsky, 1845)
Pterostichus stygicus (Say, 1823)
Pterostichus subaeneus (Chaudoir, 1850)
Pterostichus sungariensis Lafer, 1979
Pterostichus sutschanensis Jedlicka, 1962
Pterostichus tatianae E.Berlov, 1996
Pterostichus thunbergi A.Morawitz, 1862
Pterostichus tomensis (Gebler, 1847)
Pterostichus triseriatus (Gebler, 1847)
Pterostichus tschitscherinianus (Jakobson, 1907)
Pterostichus tuberifer Sasakawa, 2006
Pterostichus tundrae (Tschitscherine, 1894)
Pterostichus turanensis Jedlicka, 1959
Pterostichus uralensis (Motschulsky, 1850)
Pterostichus urengaicus Jurecek, 1924
Pterostichus vladivostokensis Lafer, 1979

Subgenus Phaenoraphis Tschitscherine, 1901

Pterostichus acuspina (Tschitscherine, 1901)
Pterostichus peninsularis Park & Kwon, 1996
Pterostichus sagittus Park, 2013

Subgenus Phonias Gozis, 1886

Pterostichus apfelbecki Csiki, 1908
Pterostichus burkhan Berlov & Anichtchenko, 2005
Pterostichus corrusculus LeConte, 1873
Pterostichus datshenkoae Sundukov, 2013
Pterostichus defossus Bates, 1883
Pterostichus diligens (Sturm, 1824)
Pterostichus dostali Kirschenhofer, 1981
Pterostichus eobius (Tschitscherine, 1899)
Pterostichus femoralis (Kirby, 1837)
Pterostichus gyrosus (Motschulsky, 1866)
Pterostichus innshanensis Jedlicka, 1960
Pterostichus jankowskyi (Tschitscherine, 1897)
Pterostichus kutensis Poppius, 1905
Pterostichus liodactylus (Tschitscherine, 1898)
Pterostichus longinquus Bates, 1873
Pterostichus longipennis Straneo, 1942
Pterostichus monostigma (Tschitscherine, 1898)
Pterostichus morawitzianus (Lutshnik, 1922)
Pterostichus neglectus A.Morawitz, 1862
Pterostichus ovoideus (Sturm, 1824)
Pterostichus patruelis (Dejean, 1831)
Pterostichus perisi Novoa, 1979
Pterostichus ripensis (Motschulsky, 1866)
Pterostichus sasajii Morita, 2007
Pterostichus setipes (Tschitscherine, 1898)
Pterostichus sotkaensis Jedlicka, 1958
Pterostichus strenuus (Panzer, 1796)
Pterostichus stricticollis (Solsky, 1874)
Pterostichus subitus Csiki, 1930
Pterostichus taksonyis Csiki, 1930
Pterostichus ussuriensis (Tschitscherine, 1897)

Subgenus Platypterus Chaudoir, 1838

Pterostichus dilatatus A. & G.B.Villa, 1835
Pterostichus lineatopunctatus L.Miller, 1850
Pterostichus lombardus K.Daniel, 1900
Pterostichus ottomanus Apfelbeck, 1908
Pterostichus panzeri (Panzer, 1802)
Pterostichus truncatus (Dejean, 1828)
Pterostichus ziegleri (Duftschmid, 1812)

Subgenus Platysma Bonelli, 1810

Pterostichus chotanensis (Tschitscherine, 1893)
Pterostichus cordaticollis Heyden, 1884
Pterostichus crenulatopunctatus (R.F.Sahlberg, 1844)
Pterostichus eschscholtzii (Germar, 1823)
Pterostichus galinae Kabak, 1992
Pterostichus insignicollis (Tschitscherine, 1893)
Pterostichus leptis Bates, 1883
Pterostichus niger (Schaller, 1783)
Pterostichus planicola (Tschitscherine, 1899)
Pterostichus sachtlebeni Jedlicka, 1962
Pterostichus turcomannicus Motschulsky, 1850

Subgenus Plectes Fischer von Waldheim, 1822

Pterostichus consors (Tschitscherine, 1893)
Pterostichus drescheri (Fischer von Waldheim, 1817)

Subgenus Pledarus Motschulsky, 1866

Pterostichus gibbicollis (Motschulsky, 1844)
Pterostichus larisae Sundukov, 2013
Pterostichus lutschniki Jedlicka, 1962
Pterostichus nigellatus Kirschenhofer, 1991

Subgenus Pseudethira Sciaky, 1996

Pterostichus angoarnigi Morvan, 1994
Pterostichus atrox (Andrewes, 1937)
Pterostichus balachowskyi Morvan, 1972
Pterostichus balu J.Schmidt, 2009
Pterostichus bhutanensis Morvan, 1978
Pterostichus brancuccii Straneo, 1982
Pterostichus brevilama Straneo, 1989
Pterostichus bureli Morvan, 1994
Pterostichus chainapaani J.Schmidt, 2006
Pterostichus championi Andrewes, 1926
Pterostichus conaensis J.Schmidt & Tian, 2011
Pterostichus deuvei Lassalle, 1985
Pterostichus dhorpatanicus Straneo, 1977
Pterostichus dolens (Tschitscherine, 1900)
Pterostichus dorjulensis Morvan, 1978
Pterostichus exochus Andrewes, 1930
Pterostichus fritzhiekei J.Schmidt, 1994
Pterostichus gagates (Hope, 1831)
Pterostichus ganesh J.Schmidt, 2006
Pterostichus ganja J.Schmidt, 1995
Pterostichus gerdi J.Schmidt, 2009
Pterostichus gobettii Straneo, 1983
Pterostichus gompanus Straneo, 1983
Pterostichus harmandi (Tschitscherine, 1900)
Pterostichus hartmanni J.Schmidt, 1995
Pterostichus hedkeinekus Morvan, 1995
Pterostichus hoelli J.Schmidt, 2009
Pterostichus immarginatus Straneo, 1977
Pterostichus jaljaleensis J.Schmidt, 2009
Pterostichus janbritoi Morvan, 1994
Pterostichus juga J.Schmidt, 2006
Pterostichus kadoudali Morvan, 1982
Pterostichus kalo J.Schmidt, 2012
Pterostichus kangchenjunga J.Schmidt, 2012
Pterostichus keltiekus Morvan, 1981
Pterostichus kleinfeldi Straneo, 1982
Pterostichus kopetzi J.Schmidt, 2006
Pterostichus leica J.Schmidt, 2012
Pterostichus letensis Habu, 1973
Pterostichus lugi Morvan, 1982
Pterostichus makalu J.Schmidt, 2012
Pterostichus matsumurai Habu, 1973
Pterostichus mewakholensis J.Schmidt, 2012
Pterostichus miehei J.Schmidt, 2006
Pterostichus nepalensis Straneo, 1977
Pterostichus nowitzkii (Tschitscherine, 1899)
Pterostichus olafi J.Schmidt, 2006
Pterostichus perlamatus Straneo, 1989
Pterostichus pseudoharmandi Morvan, 1981
Pterostichus pseudoplatysma Straneo, 1982
Pterostichus raraensis Morvan, 1980
Pterostichus rugosiceps J.Schmidt, 2012
Pterostichus sagarmatha J.Schmidt, 2012
Pterostichus santostamangi J.Schmidt, 2006
Pterostichus schrettenbrunneri J.Schmidt, 2012
Pterostichus thanglaensis J.Schmidt, 2012
Pterostichus tulobalu J.Schmidt, 2009
Pterostichus weigeli J.Schmidt, 2006
Pterostichus weiperti J.Schmidt, 2006
Pterostichus wittmeri Morvan, 1978

Subgenus Pseudoferonina Ball, 1965

Pterostichus amadeus Bousquet, 2012
Pterostichus bousqueti Bergdahl in Bergdahl & Kavanaugh, 2011
Pterostichus campbelli Bousquet, 1985
Pterostichus humidulus (Van Dyke, 1943)
Pterostichus lanei Van Dyke, 1926
Pterostichus lolo Bergdahl in Bergdahl & Kavanaugh, 2011
Pterostichus shulli (Hatch, 1949)
Pterostichus smetanai Bousquet, 1985
Pterostichus spathifer Bousquet, 1992

Subgenus Pseudohaptoderus Tschitscherine, 1888

Pterostichus aemiliae Facchini & Sciaky, 2000
Pterostichus jugivagus (Tschitscherine, 1898)
Pterostichus ssemenovi Tschitscherine, 1888

Subgenus Pseudomaseus Chaudoir, 1838

Pterostichus aimaki Jedlicka, 1968
Pterostichus ambigenus Bates, 1883
Pterostichus anthracinus (Illiger, 1798)
Pterostichus basipunctatus Straneo, 1955
Pterostichus carri Angus; Galian; Wrase & Chaladze, 2009
Pterostichus chujoiellus Jedlicka, 1962
Pterostichus fuscicornis (Reiche & Saulcy, 1855)
Pterostichus gracilis (Dejean, 1828)
Pterostichus hejkali B.Gueorguiev & Skoupy, 2011
Pterostichus koslovi Solodovnikov, 2001
Pterostichus luctuosus (Dejean, 1828)
Pterostichus michailovi Wrase, 1992
Pterostichus minor (Gyllenhal, 1827)
Pterostichus mukdenensis Breit, 1933
Pterostichus nigrita (Paykull, 1790)
Pterostichus oenotrius Ravizza, 1975
Pterostichus piceolus (Chaudoir, 1850)
Pterostichus rhaeticus Heer, 1837
Pterostichus rotundangulus A.Morawitz, 1862
Pterostichus tenuis (Casey, 1924)
Pterostichus tsukubasanus Kasahara, 1988

Subgenus Pseudorambousekiella Schweiger, 1967

Pterostichus montisdeorum Schweiger, 1967

Subgenus Pseudorites Ganglbauer, 1891

Pterostichus nicaeensis (A. & G.B.Villa, 1835)

Subgenus Pseudosteropus Chaudoir, 1838

Pterostichus illigeri (Panzer, 1802)
Pterostichus schmidtii (Chaudoir, 1838)

Subgenus Pterostichus Bonelli, 1810

Pterostichus andreinii Dodero, 1922
Pterostichus auratus Heer, 1837
Pterostichus bischoffianus Jedlicka, 1936
Pterostichus bruckii Schaum, 1859
Pterostichus cantalicus (Chaudoir, 1868)
Pterostichus cristatus (L.Dufour, 1820)
Pterostichus devillei Puel, 1924
Pterostichus dux L.Schaufuss, 1862
Pterostichus epiroticus Csiki, 1930
Pterostichus fasciatopunctatus (Creutzer, 1799)
Pterostichus focarilei Casale & Giachino, 1985
Pterostichus funestes Csiki, 1930
Pterostichus hagenbachii Sturm, 1824
Pterostichus honnoratii (Dejean, 1828)
Pterostichus impressicollis (Fairmaire & Laboulbène, 1854)
Pterostichus lanista (Tschitscherine, 1898)
Pterostichus lasserrei (Dejean, 1828)
Pterostichus latifianus Apfelbeck, 1906
Pterostichus lumensis Apfelbeck, 1905
Pterostichus malissorum Apfelbeck, 1905
Pterostichus meisteri Reitter, 1885
Pterostichus merklii (J.Frivaldszky, 1879)
Pterostichus micans Heer, 1838
Pterostichus pedemontanus Ganglbauer, 1891
Pterostichus pentheri Apfelbeck, 1918
Pterostichus phaeopus (Sainte-Claire Deville, 1903)
Pterostichus reiseri Ganglbauer, 1889
Pterostichus rhilensis Rottenberg, 1874
Pterostichus ruffoi Sciaky, 1984
Pterostichus rufipes (Dejean, 1828)
Pterostichus rutilans (Dejean, 1828)
Pterostichus sacheri I.Frivaldszky von Frivald, 1865
Pterostichus vagepunctatus Heer, 1837
Pterostichus walteri Reitter, 1883

Subgenus Rambousekiella Knirsch, 1925

Pterostichus ledenikensis (Knirsch, 1925)

Subgenus Rhagadus Motschulsky, 1866

Pterostichus brittoni Habu, 1958
Pterostichus glabripennis Jedlicka, 1962
Pterostichus harponifer Tanaka, 1987
Pterostichus ishiii Morita; Kurosa & Mori, 2009
Pterostichus kalhys (Motschulsky, 1866)
Pterostichus kimurai Morita, 1994
Pterostichus komiyai Morita, 2007
Pterostichus laevipunctatus Tschitscherine, 1889
Pterostichus latemarginatus (Straneo, 1936)
Pterostichus microcephalus (Motschulsky, 1861)
Pterostichus modicellus (Tschitscherine, 1897)
Pterostichus mundatus Jedlicka, 1940
Pterostichus nimbatidius (Chaudoir, 1878)
Pterostichus polygenus Bates, 1883
Pterostichus satsumanus Habu, 1958
Pterostichus solskyi (Chaudoir, 1878)
Pterostichus takaosanus Habu, 1958
Pterostichus thorectes Bates, 1873
Pterostichus thorectoides Jedlicka, 1958

Subgenus Sinoreophilus Sciaky, 1996

Pterostichus potanini Tschitscherine, 1889
Pterostichus scalptus Sciaky & Wrase, 1997
Pterostichus strigosus Sciaky & Wrase, 1997
Pterostichus validior Tschitscherine, 1889

Subgenus Sinosteropus Sciaky, 1994

Pterostichus barbarae Sciaky & Facchini, 2003
Pterostichus carinatus Sciaky & Facchini, 2003
Pterostichus cathaicus Sciaky, 1994
Pterostichus curvatus Sciaky & Facchini, 2003
Pterostichus horstweiperti J.Schmidt & Guorguiev, 2013
Pterostichus jaechianus Sciaky & Facchini, 2003
Pterostichus jani Sciaky & Facchini, 2003
Pterostichus latitemporis Sciaky & Wrase, 1997
Pterostichus microbus Sciaky & Facchini, 2003
Pterostichus perhoplites J.Schmidt & Tian, 2011
Pterostichus pseudobarbarae Sciaky & Facchini, 2003
Pterostichus pseudojugivagus J.Schmidt & Tian, 2011
Pterostichus pseudorotundus Sciaky & Facchini, 2003
Pterostichus pseudosinensis Sciaky & Facchini, 2003
Pterostichus rotundus Sciaky, 1994
Pterostichus schneideri Sciaky & Facchini, 2003
Pterostichus scuticollis (Fairmaire, 1889)
Pterostichus septemtrionalis Sciaky & Facchini, 2003
Pterostichus sinensis Jedlicka, 1962
Pterostichus triangularis Sciaky & Facchini, 2003
Pterostichus wrasei Sciaky & Facchini, 2003

Subgenus Sphodroferonia Kasahara & Y.Ito, 1987

Pterostichus abaciformis Straneo, 1955
Pterostichus arcuaticarinatus Kasahara, 1986
Pterostichus audax (Tschitscherine, 1895)
Pterostichus bilobatus N.Ito, 2010
Pterostichus bisulcitarsis N.Ito, 2012
Pterostichus confusianus N.Ito, 2010
Pterostichus constricticollis N.Ito, 2010
Pterostichus esakii Ishida, 1959
Pterostichus fenestratus Kasahara & Ohtani, 1989
Pterostichus geojensis Sasakawa; Kim & Kubota, 2006
Pterostichus gotoensis Kasahara & Matsumoto, 1990
Pterostichus gracilitarsis N.Ito, 2010
Pterostichus hikosanus Kasahara, 1994
Pterostichus hisamatsui Ishida & Shibata, 1961
Pterostichus imasakai Kasahara & Ohtani, 1988
Pterostichus imitatorius N.Ito, 2010
Pterostichus ishizuchiensis Kasahara, 1985
Pterostichus itoi Kasahara, 1986
Pterostichus kosakai Morita, 1998
Pterostichus koujitanakai N.Ito, 2010
Pterostichus kyushuensis Habu, 1955
Pterostichus laeviceps N.Ito, 2010
Pterostichus macrocephalus Habu, 1955
Pterostichus marginellus Sasakawa; Kim & Kim, 2006
Pterostichus masatakayoshidai N.Ito, 2010
Pterostichus masidai Ishida, 1959
Pterostichus masumotoi Tanaka; Morita & Suga, 1987
Pterostichus miyamai Kasahara & Y.Ito, 1987
Pterostichus mosaicus Sasakawa, 2005
Pterostichus mucronatus Straneo, 1955
Pterostichus multinodosus Sasakawa; Kim & Kim, 2006
Pterostichus namedai Morita, 2019
Pterostichus nasui Kasahara, 1993
Pterostichus nishidai Nakane, 1989
Pterostichus obscurimpressus N.Ito, 2010
Pterostichus okiensis Nakane, 1989
Pterostichus orionis Jedlicka, 1962
Pterostichus pachinus Bates, 1883
Pterostichus platyocularis N.Ito, 2010
Pterostichus platypennis N.Ito, 2010
Pterostichus plesiomorphus Nemoto, 1989
Pterostichus prominens N.Ito, 2010
Pterostichus pseudopachinus Nakane, 1963
Pterostichus punctatulus N.Ito, 2010
Pterostichus punctibasalis N.Ito, 2012
Pterostichus raptor (Tschitscherine, 1901)
Pterostichus sakuragii N.Ito, 2010
Pterostichus sculpturalis N.Ito, 2012
Pterostichus scurra (Tschitscherine, 1901)
Pterostichus scurroides Jedlicka, 1958
Pterostichus sediorcrus Sasakawa & Kim, 2006
Pterostichus seunglaki Park & Kwon, 1996
Pterostichus shiibanus Habu, 1958
Pterostichus sphodriformis Bates, 1873
Pterostichus strigipennis N.Ito, 2012
Pterostichus subabacipennis N.Ito, 2012
Pterostichus sulcatus N.Ito, 2010
Pterostichus suruganus Straneo in Nakane, 1979
Pterostichus symmetricus Straneo, 1955
Pterostichus taradakensis Kasahara & Ohtani, 1988
Pterostichus tosanus Kasahara & Y.Ito, 1997
Pterostichus touzalinioides Sasakawa; Kim & Kim, 2007
Pterostichus tsurugaensis Ishida, 1959
Pterostichus variicornis Sasakawa; Kim & Kim, 2007
Pterostichus yakushimanus Nakane & Ishida, 1961
Pterostichus yamajii Kasahara, 1993
Pterostichus yoshidai Kasahara, 1985
Pterostichus yoshikawai Ishida, 1959
Pterostichus yoshiyukiitoi N.Ito, 2010

Subgenus Steropanus Fairmaire, 1889

Pterostichus aequus (Andrewes, 1937)
Pterostichus alveolatus Fedorenko, 2018
Pterostichus asulcatus Fedorenko, 2018
Pterostichus boriskataevi Fedorenko, 2020
Pterostichus cavifrons Fedorenko, 2018
Pterostichus deliciatus (Andrewes, 1937)
Pterostichus felix (Andrewes, 1937)
Pterostichus forticornis (Fairmaire, 1889)
Pterostichus fossifrons Fedorenko, 2020
Pterostichus glymmiger (Andrewes, 1937)
Pterostichus infissus (Andrewes, 1937)
Pterostichus mengtzei Jedlicka, 1931
Pterostichus obliteratus Fedorenko, 2018
Pterostichus pseudoglymmiger Fedorenko, 2020
Pterostichus pseudoviolaceus Fedorenko, 2018
Pterostichus securipenis Fedorenko, 2020
Pterostichus sulcatipennis Fedorenko, 2018
Pterostichus thailandensis (Morvan, 1992)
Pterostichus thailandicus Straneo, 1989
Pterostichus violaceus (Straneo, 1949)

Subgenus Steropus Dejean, 1821

Pterostichus catalonicus J.Daniel in K. & J.Daniel, 1906
Pterostichus ebenus (Quensel, 1806)
Pterostichus ferreri (Español & Mateu, 1942)
Pterostichus galaecianus Lauffer, 1909
Pterostichus gallega (Fairmaire, 1859)
Pterostichus ghilianii (Putzeys, 1845)
Pterostichus insidatrix (Piochard de la Brûlerie, 1872)
Pterostichus madidus (Fabricius, 1775)
Pterostichus rifensis Antoine, 1933
Pterostichus validus (Dejean, 1828)

Subgenus Tausternus Fedorenko, 2020

Pterostichus hoii Fedorenko, 2020

Subgenus Tinautius Mateu, 1997

Pterostichus exilis (Mateu, 2001)
Pterostichus troglophilus (Mateu, 1997)

Subgenus Tschitscherinea Berg, 1898

Pterostichus farkaci Sciaky, 1997
Pterostichus filum (Tschitscherine, 1897)
Pterostichus krali Sciaky, 1997
Pterostichus mulensis Sciaky, 1997

Subgenus Tubuliphallus Sciaky & Allegro, 2013

Pterostichus megacephalus Sciaky & Allegro, 2013

Subgenus Unitrichus Sciaky, 1997

Pterostichus platyops Sciaky, 1997

Subgenus Vietosteropus Fedorenko, 2017

Pterostichus abramovi Fedorenko, 2017
Pterostichus anichkini Fedorenko, 2017
Pterostichus annamita (Straneo, 1939)
Pterostichus bidoupensis Fedorenko, 2017
Pterostichus cavicollis Straneo, 1984
Pterostichus chupanphan Fedorenko, 2017
Pterostichus dalatensis Fedorenko, 2017
Pterostichus gialaiensis Fedorenko, 2017
Pterostichus honbaensis Fedorenko, 2017
Pterostichus konchurang Fedorenko, 2017
Pterostichus konplongensis Fedorenko, 2017
Pterostichus kontumensis Fedorenko, 2017
Pterostichus kuznetsovi Fedorenko, 2020
Pterostichus ngoclinhensis Fedorenko, 2017
Pterostichus ngokboci Fedorenko, 2017
Pterostichus semiopacus Fedorenko, 2017
Pterostichus sulcicollis Fedorenko, 2017

Subgenus Wraseiellus Shi & Sciaky, 2013

Pterostichus andrewesi Jedlicka, 1931
Pterostichus comatus Shi & Sciaky, 2013
Pterostichus crassiapex Shi & Sciaky, 2013
Pterostichus diversus (Fairmaire, 1886)
Pterostichus kambaiti (Andrewes, 1947)
Pterostichus meyeri Jedlicka, 1934
Pterostichus pseudodiversus Shi & Sciaky, 2013
Pterostichus stictopleurus (Fairmaire, 1889)

Unassigned Subgenus

Pterostichus analis Jedlicka, 1963
Pterostichus antepunctatus Straneo, 1949
Pterostichus aristochroides Deuve, 2006
Pterostichus chenpengi Li, 1992
Pterostichus clepsydra Sciaky & Wrase, 1997
Pterostichus deuvesianus Morvan, 1995
Pterostichus georgi Straneo, 1963
Pterostichus glabricollis Jedlicka, 1962
Pterostichus gwervaenus Morvan, 1995
Pterostichus heilongjiangensis Li, 1992
Pterostichus imitatus Morvan, 1978
Pterostichus jelepus Andrewes, 1932
Pterostichus kiangsu Jedlicka, 1965
Pterostichus kolosovi (Lutshnik, 1928)
Pterostichus leviculus Andrewes, 1933
Pterostichus lidarus (Andrewes, 1937)
Pterostichus lukjanovitshi (Lutshnik, 1927)
Pterostichus malaisei Jedlicka, 1965
Pterostichus migliaccioi Straneo, 1982
Pterostichus orobius Fairmaire, 1861
Pterostichus peilingi Jedlicka, 1937
Pterostichus punctisternus Straneo, 1989
Pterostichus quadriimpressus Straneo, 1983
Pterostichus reuteri Wrase & J.Schmidt, 2006
Pterostichus rolex Morvan, 1995
Pterostichus sintanus Andrewes, 1927
Pterostichus styx (Andrewes, 1937)
Pterostichus szekessyianus Sciaky, 1996
Pterostichus tantillus (Fairmaire, 1889)
Pterostichus tonkinensis Straneo, 1980
Pterostichus zhejiangensis Kirschenhofer, 1997
†Pterostichus abrogatus Scudder, 1890
†Pterostichus depletus Scudder, 1900
†Pterostichus destitutus Scudder, 1890
†Pterostichus destructus Scudder, 1890
†Pterostichus dormitans Scudder, 1890
†Pterostichus fernquisti Wickham, 1931
†Pterostichus fractus Scudder, 1890
†Pterostichus laevigatus Scudder, 1890
†Pterostichus minax (Oustalet, 1874)
†Pterostichus minutulus Heer, 1862
†Pterostichus provincialis (Oustalet, 1874)
†Pterostichus pumpellyi Scudder, 1900
†Pterostichus walcotti Scudder, 1900

Notes and References

Bervoets . L . Habitat preference and activity period of some carabid species (Coleoptera, Carabidae) in a heathland ecosystem based on a four day sampling period . Bull. Ann. Soc. R. Ent. Belg. . 130 . 115–118 . 1994 . Int Panis . L . Verheyen . R.F..