Pleosporomycetidae Explained

Pleosporomycetidae is a subclass of Dothideomycetes consisting of four orders: Pleosporales, Hysteriales, Mytilinidiales, and Jahnulales.^{[1] [2]} One of its defining features is the presence of pseudoparaphyses.^[3] These are sterile cells extending down from the upper portion of the cavity inside sexual structures containing the sac-like asci with sexually produced spores (ascospores). Pseudoparaphyses are initially attached at both their ends, but sometimes the upper part may become free. Some orders and families where these cells are present remain outside the subclass since DNA-based phylogenies cannot place them with confidence. However, they could conceivably be included within Pleosporomycetidae in future.

Notes and References

1. Schoch CL, Shoemaker RA, Seifert KA, Hambleton S, Spatafora JW, Crous PW, 2006. A multigene phylogeny of the Dothideomycetes using four nuclear loci. Mycologia 98, 1041–1052.
2. Schoch CL, Crous PW, Groenewald JZ, Boehm EW, Burgess TI, de Gruyter J, de Hoog GS, Dixon LJ, Grube M, Gueidan C, Harada Y, Hatakeyama S, Hirayama K, Hosoya T, Huhndorf SM, Hyde KD, Jones EB, Kohlmeyer J, Kruys A, Li YM, Lucking R, Lumbsch HT, Marvanova L, Mbatchou JS, McVay AH, Miller AN, Mugambi GK, Muggia L, Nelsen MP, Nelson P, Owensby CA, Phillips AJ, Phongpaichit S, Pointing SB, Pujade-Renaud V, Raja HA, Plata ER, Robbertse B, Ruibal C, Sakayaroj J, Sano T, Selbmann L, Shearer CA, Shirouzu T, Slippers B, Suetrong S, Tanaka K, Volkmann-Kohlmeyer B, Wingfield MJ, Wood AR, Woudenberg JH, Yonezawa H, Zhang Y, Spatafora JW, 2009. A class-wide phylogenetic assessment of Dothideomycetes. Stud Mycol 64, 1-15S10. http://www.studiesinmycology.org/cgi/content/abstract/64/1/1
3. Eriksson OE, 1981. The families of bitunicate ascomycetes. Opera Botanica 60, 1-220.