Philine Explained

Philine is a genus of sea slugs or sea snails, marine gastropod molluscs in the subfamily Philininae of the family Philinidae, the headshield slugs or paper bubbles.^[1]

Species

Species within the genus Philine include:

• Philine abyssicola Valdès, 2008
• Philine aethiopica Thiele, 1925
• Philine alboides Price, Gosliner & Valdés, 2011
• Philine amabilis Verrill, 1880
• Philine angasi Crosse and Fischer, 1865
• Philine angulata Jeffreys, 1867 - angled paper-bubble, angled paperbubble
• Philine aperta (Linnaeus, 1767)
• Philine approximans Dautzenberg & Fischer H., 1896
• Philine araneosa van der Linden, 1995
• Philine argentina Carcelles, 1947
• Philine auriformis Suter, 1909
• Philine azorica Bouchet, 1975
• Philine babai Valdés, 2008
• Philine bakeri Dall, 1919
• Philine baxteri Valdés, Cadien & Gosliner, 2016
• Philine beachportensis Verco, 1909
• Philine berghi E. A. Smith, 1910
• Philine buchensis Caballer & Ortea, 2015
• Philine burrowsi Burn, 1961
• Philine caballeri Ortea, Espinosa & Moro, 2001
• Philine caledonica Risbec, 1951
• Philine candeana (d'Orbigny, 1841)
• Philine catena (Montagu, 1803)
• Philine cerebralis Malaquias, Ohnheiser, Oskars & Willassen, 2016
• Philine columnaria Hedley & May, 1908
• Philine complanata Watson, 1897
• Philine constricta Murdoch and Suter, 1906
• Philine cumingii (Adams 1862)
• Philine denticulata (J. Adams, 1800)
• Philine dentiphallus Gonzales & Gosliner, 2014
• Philine elegans Bergh, 1905
• Philine exigua Challis, 1969
• Philine fenestra Price, Gosliner & Valdés, 2011
• Philine gelida van der Linden, 1995
• Philine guineensis Ev. Marcus & Er. Marcus, 1966
• Philine habei Valdés, 2008
• Philine harrisae Valdés, Cadien & Gosliner, 2016
• Philine hemphilli Dall, 1919
• Philine infortunata Pilsbry, 1895
• Philine infundibulum Dall, 1889
• Philine intricata Monterosato, 1875
• Philine iris Tringali, 2001
• Philine kinglipini Tchang, 1934
• Philine kurodai Habe, 1946
• Philine lucida Dall, 1927
• Philine malaquiasi Valdés, Cadien & Gosliner, 2016
• Philine mcleani Valdés, Cadien & Gosliner, 2016
• Philine mera Ev. Marcus & Er. Marcus, 1969
• Philine monilifera Bouchet, 1975
• Philine monterosati Monterosato, 1874
• Philine multipapillata Gonzales & Gosliner, 2014
• Philine orca Gosliner, 1988
• Philine orientalis A. Adams, 1854
• Philine multipapillata Gonzales & Gosliner, 2014
• Philine paucipapillata Price, Gosliner & Valdés, 2011
• Philine pittmani Gonzales & Gosliner, 2014
• Philine planata Dall, 1889
• Philine polystrigma (Dall, 1908)
• Philine powelli Rudman, 1870
• Philine puka Price, Gosliner & Valdés, 2011
• Philine punctata (J. Adams, 1800)
• Philine quadripartita Ascanius, 1772
• Philine rubra Bergh, 1905
• Philine rubrata Gosliner, 1988
• Philine rugosula Dautzenberg & Fischer H., 1896
• Philine sagra (d'Orbigny, 1841) - crenulate paper-bubble, crenulate paperbubble
• Philine sarcophaga Price, Gosliner & Valdés, 2011
• Philine scalpta Adams, 1862
• Philine schrammi Malaquias, Ohnheiser, Oskars & Willassen, 2016
• Philine sinuata Stimpson, 1851
• Philine striatula Monterosato, 1874
• Philine striolata A. Adams, 1862
• Philine talismani Sykes, 1905
• † Philine tepikia Rudman, 1970
• Philine teres Hedley, 1903
• Philine tincta A. E. Verrill, 1882 - tinted paper-bubble, tinted paperbubble
• Philine trapezia Hedley, 1902
• Philine umbilicata Murdoch and Suter, 1906
• Philine vaillanti Issel, 1869
• Philine verdensis Gonzales & Gosliner, 2014
• Philine vestita (Philippi, 1840)
• Philine wareni Valdés, Cadien & Gosliner, 2016

Taxa inquirenda

• Philine acutangula A. Adams, 1862
• Philine desmotis Watson, 1897
• Philine minuta Thiele, 1925
• Philine sykesii Melvill, 1904
• Philine truncatissima G. B. Sowerby II, 1870
• Philine vitrea Gould, 1859

Species brought into synonymy:

• Philine alata Thiele, 1912: synonym of Antarctophiline alata (Thiele, 1912) (original combination)
• Philine alba Mattox, 1958 - white paper-bubble, white paperbubble: synonym of Philinorbis albus (Mattox, 1958) (original combination)
• Philine alternans van der Linden, 1995: synonym of Laona alternans (van der Linden, 1995) (original combination)
• Philine antarctica Smith, 1902: synonym of Waegelea antarctica (E. A. Smith, 1902) (original combination)
• Philine apertissima Smith, 1902: synonym of Antarctophiline apertissima (E. A. Smith, 1902) (original combination)
• Philine apertissima de Folin, 1893: synonym of Philine quadripartita Ascanius, 1772
• Philine argentata Gould, 1859: synonym of Philine orientalis A. Adams, 1854
• Philine californica Willett, 1944 - California paper-bubble, California paperbubble : synonym of Laona californica (Willett, 1944) (original combination)
• Philine calva van der Linden, 1995: synonym of Philine striatula Monterosato, 1874
• Philine chilla Er. Marcus & Ev. Marcus, 1969: synonym of Laona chilla (Er. Marcus & Ev. Marcus, 1969) (original combination)
• Philine cingulata G. O. Sars, 1878 - girdled paper-bubble, girdled paperbubble : synonym of Philine finmarchica M. Sars, 1858
• Philine condensa van der Linden, 1995: synonym of Laona condensa (van der Linden, 1995) (original combination)
• Philine confusa Ohnheiser & Malaquias, 2013: synonym of Laona confusa (Ohnheiser & Malaquias, 2013)
• Philine falklandica A. W. B. Powell, 1951: synonym of Antarctophiline falklandica (Powell, 1951) (original combination)
• Philine fragilis G. O. Sars, 1878 - fragile paper-bubble, fragile paperbubble : synonym of Philine finmarchica M. Sars, 1858: synonym of Praephiline finmarchica (M. Sars, 1859)
• Philine gibba (Strebel, 1908): synonym of Antarctophiline gibba (Strebel, 1908) (original combination)
• Philine gouldi Doello-Jurado, 1918: synonym of Antarctophiline gibba (Strebel, 1908)
• Philine grandioculi Ohnheiser & Malaquias, 2013: synonym of Laona grandioculi (Ohnheiser & Malaquias, 2013)
• Philine indistincta Ohnheiser & Malaquias, 2013: synonym of Hermania indistincta (Ohnheiser & Malaquias, 2013) (original combination)
• Philine japonica Lischke, 1872: synonym of Philine orientalis A. Adams, 1854
• Philine kawamurai (Habe, 1958): synonym of Globophiline kawamurai Habe, 1958
• Philine kerguelensis Thiele, 1925: synonym of Spiraphiline kerguelensis (Thiele, 1925) (original combination)
• Philine laevissima M. Sars, 1859: synonym of Diaphana hiemalis (Couthouy, 1839)
• Philine lima (T. Brown, 1825) - file paper-bubble, file paperbubble: synonym of Retusophiline lima (T. Brown, 1827)
• Philine lineolata (Couthouy, 1839): synonym of Philine lima (T. Brown, 1825): synonym of Retusophiline lima (T. Brown, 1827)
• Philine loveni Malm, 1858: synonym of Philine scabra (O. F. Müller, 1784)
• Philine milneedwardsi Locard, 1897: synonym of Philine quadripartita Ascanius, 1772
• Philine monterosatoi Sykes, 1905: synonym of Philine monterosati Monterosato, 1874 (unjustified emendation)
• Philine ossiansarsi Friele, 1877: synonym of Philine finmarchica M. Sars, 1858
• Philine polaris Aurivillius, 1885 - axial paper-bubble, axial paperbubble: synonym of Philine quadrata (S. Wood, 1839)
• Philine pruinosa (Clark, 1827): synonym of Laona pruinosa (W. Clark, 1827)
• Philine polystrigma (Dall, 1908): synonym of Woodbridgea polystrigma (Dall, 1908)
• Philine quadrata (S. V. Wood, 1839) - quadrate paper-bubble, quadrate paperbubble: synonym of Laona quadrata (S. Wood, 1839)
• Philine retifera (Forbes, 1844): synonym of Philine vestita (Philippi, 1840)
• Philine rugulosa (Dautzenberg & Fischer H., 1896): synonym of Philine rugosula Dautzenberg & Fischer H., 1896
• Philine scabra Muller, 1784: synonym of Hermania scabra (O. F. Müller, 1784)
• Philine sinuata Stimpson, 1851 - sinuate paper-bubble, sinuate paperbubble : synonym of Philine denticulata (Adams J., 1800)
• Philine striatella Tapparone-Canefri, 1874 : synonym of Philine orientalis A. Adams, 1854
• Philine trachyostraca Watson, 1897: synonym of Philine retifera (Forbes, 1844)
• Philine thurmanni Ev. Marcus & Er. Marcus, 1969: synonym of Laona thurmanni (Ev. Marcus & Er. Marcus, 1969)
• Philine ventricosa (Jeffreys, 1865): synonym of Laona ventricosa (Jeffreys, 1865)

References

• Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). . XII, 195 pp.
• Chaban E.M. & Soldatenko E.V. (2009) Description of a new genus Praephiline gen. nov. (Gastropoda: Opisthobranchia: Philinidae). Zoosystematica Rossica 18(2): 205-211.
• Price R.M., Gosliner T.M. & Valdés A. (2011) Systematics and phylogeny of Philine (Gastropoda: Opisthobranchia), with emphasis on the Philine aperta species complex. The Veliger 51(2): 1-58.

External links

• • • Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979

Notes and References

1. Gofas, S. (2011). Philine Ascanius, 1772. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138339 on 2011-09-21