Trochoidea (superfamily) explained

Trochoidea is a superfamily of small to very large vetigastropod sea snails with gills and an operculum.^[1] Species within this superfamily have nacre as the inner shell layer. The families within this superfamily include the Trochidae, the top snails. This superfamily is the largest vetigastropodan superfamily, containing more than 2,000 species.^[2]

This taxon is not the same as a pulmonate land snail genus which is spelled the same way: Trochoidea (genus).

Taxonomy

2005 taxonomy

This superfamily consisted of nine following families (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

• Trochidae Rafinesque, 1815
• Calliostomatidae Thiele, 1924 (1847)
• † Elasmonematidae Knight, 1956
• † Eucochlidae Bandel, 2002
• † Microdomatidae Wenz, 1938
• † Proconulidae Cox, 1960
• Solariellidae Powell, 1951 - synonym: Minoliinae Kuroda, Habe & Oyama, 1971
• † Tychobraheidae Horný, 1992
• † Velainellidae Vasseur, 1880

2017 taxonomy

Williams et al. (2008)^[3] have made further refinements to the taxonomy of Trochoidea. The revised taxonomy as defined by Bouchet et al. (2017) resulted in the following families.^[4] The fossil families, tentatively included in this list, are based on Tracey et al. (1993) ^[5]

• Angariidae Gray, 1857
• † Anomphalidae Wenz, 1938
• † Araeonematidae Nützel, 2012
• Areneidae McLean, 2012
• Calliostomatidae Thiele, 1924 (1847)
• Colloniidae Cossmann, 1917
• Conradiidae Golikov & Starobogatov, 1987
• † Elasmonematidae Knight, 1956
• † Epulotrochidae Gründel, Keupp & Lang, 2017
• † Eucochlidae Bandel, 2002
• Liotiidae Gray, 1850
• Margaritidae Thiele, 1924
• †Metriomphalidae Gründel, Keupp & Lang, 2017
• † Microdomatidae Wenz, 1938
• †Nododelphinulidae Cox, 1960
• Phasianellidae Swainson, 1840
• † Proconulidae Cox, 1960
• † Sclarotrardidae Gründel, Keupp & Lang, 2017
• Skeneidae Clark W., 1851
• Solariellidae Powell, 1951
• Tegulidae Kuroda, Habe & Oyama, 1971
• Trochidae Rafinesque, 1815
• Turbinidae Rafinesque, 1815
• † Tychobraheidae Horný, 1992
• † Velainellidae Vasseur, 1880
• Trochoidea (unassigned)

Unassigned to a family:

• Margarella Thiele, 1893 - Margarella rosea belongs to Cantharidinae, but Margarella antarctica belongs to Calliostomatidae/Thysanodontinae as Carinastele antarctica.^[6]

Nomen dubium:

• Ataphridae Cossmann, 1915

Families brought into synonymy:

• Crosseolidae Hickman, 2013: synonym of Conradiidae Golikov & Starobogatov, 1987
• Cyclostrematidae P. Fischer, 1865: synonym of Liotiidae Gray, 1850
• Delphinulidae Stoliczka, 1868: synonym of Angariidae Gray, 1857
• Gazidae Hickman & McLean, 1990: synonym of Margaritidae Thiele, 1924
• Minoliinae Kuroda, Habe & Oyama, 1971: synonym of Solariellidae Powell, 1951
• † Parataphrinae Calzada, 1989: synonym of † Proconulidae Cox, 1960
• Stomatellidae Gray, 1840: synonym of Stomatellinae Gray, 1840
• Stomatiidae Carpenter, 1861: synonym of Stomatellinae Gray, 1840
• Tricoliidae Woodring, 1928: synonym of Tricoliinae Woodring, 1928 (rank change)

References

• Hickman C. S. & McLean J. H. (1990). Systematic revision and suprageneric classification of trochacean gastropods. Science Series of Natural History Museum of Los Angeles County 35: 1-169
• Hickman C.S. (1992). Reproduction and development of Trochacean gastropods. The Veliger. 35(4): 245-272
• Geiger. Daniel L.. Christine E. Thacker . Molecular phylogeny of Vetigastropoda reveals non-monophyletic Scissurellidae, Trochoidea, and Fissurelloidea. Molluscan Research. 2005 . 25. 1. 47–55. 10.11646/mr.25.1.6 .
• Suzanne T. Williams (2012), Advances in molecular systematics of the vetigastropod superfamily Trochoidea; Zoologica Scripta Volume 41, Issue 6, pages 571–595, November 2012

Notes and References

1. Gofas, S. (2013). Trochoidea. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=156489 on 2013-06-29
2. Advances in molecular systematics of the vetigastropod superfamily Trochoidea . 10.1111/j.1463-6409.2012.00552.x . 2012 . Williams . Suzanne T. . Zoologica Scripta . 41 . 6 . 571–595 .
3. Williams S. T., Karube S. & Ozawa T. (September 2008) "Molecular systematics of Vetigastropoda: Trochidae, Turbinidae and Trochoidea redefined". Zoologica Scripta 37(5): 483-506.
4. "Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families" by Philippe Bouchet & Jean-Pierre Rocroi, Bernhard Hausdorf, Andrzej Kaim, Yasunori Kano, Alexander Nützel, Pavel Parkhaev, Michael Schrödl and Ellen E. Strong in Malacologia, 2017, 61(1–2): 1–526
5. Tracey S., J.A. Todd & D.H. Erwin, 1993, Mollusca: Gastropoda. Pp. 131–167, in: M. J. Benton, ed., The Fossil Record, volume 2. chapman & hall, london. 845 pp.
6. Williams S.T., Donald K.M., Spencer H.G. & Nakano T. (2010) Molecular systematics of the marine gastropod families Trochidae and Calliostomatidae (Mollusca: Superfamily Trochoidea). Molecular Phylogenetics and Evolution 54:783-809.