Paleobiota of the Yixian Formation explained

The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans about 1.6 million years during the early Cretaceous period (Barremian-Aptian stage). It is known for its fossils, listed below.

The Yixian Formation is divided into the following subunits: (ages from Zhong et al., 2021[1] unless otherwise noted)

Sauropods

GenusSpeciesCountyMemberMaterialNotesImages
DongbeititanD. dongiBeipiaoJianshangou BedPartial postcrania[2] A basal titanosauriform
cf. Euhelopuscf. E. sp.Lujiatun BedIsolated teeth[3] [4] A basal titanosauriform
LiaoningotitanL. sinensisnearly complete skeleton[5] A basal titanosauriform
RuixiniaR. zhangiBeipiaoA partial articulated skeleton[6] A somphospondylan titanosauriform
Sauropoda spp.Lujiatun & Jianshangou BedsIsolated vertebrae and teethOne or more species of sauropods, possibly titanosaurs, have been recovered from the Yixian Formation, though most remain undescribed

Ornithischians

The dinosaurs listed below were compiled in a survey by Xu and Norell in 2006, unless otherwise noted.

Ceratopsians

GenusSpeciesCountyMemberMaterialNotesImages
LiaoceratopsL. yanzigouensisBeipiaoLujiatun Bed"[Two] nearly complete skulls, juvenile to adult."[8] A ceratopsian
PsittacosaurusP. lujiatunensisBeipiaoLujiatun & Jianshangou BedsThousands of specimens in the Lujiatun Bed, somewhat less common in the Jianshangou BedA ceratopsian

Ornithopods

GenusSpeciesCountyMemberMaterialNotesImages
BolongB. yixianensisJianshangou BedA nearly complete skeleton.An iguanodontian ornithopod
Changmiania[9] C. liaoningensisLujiatun BedTwo nearly complete skeletonsA basal ornithopod that may have dug burrows
JeholosaurusJ. shangyuanensisBeipiaoLujiatun Bed"Skull with associated vertebrae and hindlimb, skull with cervical vertebrae."[10] An ornithopod
JinzhousaurusJ. yangiYixianDawangzhangzi Bed"Nearly complete skull and skeleton."[11] An ornithopod

Theropods

The dinosaurs listed below were compiled in a survey by Xu and Norell in 2006, unless otherwise noted.[12]

Avialans

GenusSpeciesCountyMemberMaterialNotesImages
Abitusavis[13] A. liiNingcheng"Nearly complete skeleton."
ArchaeorhynchusA. spathulaDawangzhangzi BedAn early, advanced short-tailed bird (ornithuran)[14]
Changchengornis[15] C. hengdaoziensisJianshangou Bed"Nearly complete skeleton."A confuciusornithid
ConfuciusornisC. chuonzhousJunior synonym of Confuciusornis sanctus
C. dui[16] Jianshangou Bed"[Two] skeletons."
C. feducciai[17] Jianshangou Bed
C. sanctusJianshangou and Dawangzhangzi Beds"Hundreds of complete skeletons."A primitive short-tailed bird (pygostylian), also present in the Jiufotang Formation
C. suniaeJunior synonym of Confuciusornis sanctus
DalingheornisD. liweiiDawangzhangzi BedAn Enantiornithine[18]
EoenantiornisE. buhleriJianshangou Bed"Nearly complete skeleton."[19] An entantiornithine
EogranivoraE. edentulataDawangzhangzi Bed"nearly complete skeleton with skull"[20] An ornithuromorph
IteravisI. huchzermeyeriAn ornithuromorph
JinzhouornisJ. yixianensisJianshangou BedA pygostylian, a junior synonym of Confuciusornis sanctus[21]
J. zhangjiyingiaJianshangou BedJunior synonym of C. sanctus
JixiangornisJ. orientalisJianshangou BedA basal avialan, also present in the Jiufotang Formation[22]
HongshanornisH. longicrestaJianshangou and Dawangzhangzi Beds[23] 4 specimensAn ornithuromorph[24]
JibeiniaJ. luaheraPreviously regarded as Yixian, horizon reassigned to the Huajiying Formation by Jin et al., 2008.
LiaoningornisL. longidigitrisLujiatun Bed"Partial skeleton."[25] An enantiornithine.[26]
LiaoxiornisL. delicatusDawangzhangzi BedConsidered a nomen dubium because the name is based on remains of a hatchling entantiornithine and cannot be confidently matched with the adult form.[27]
LingyuanornisL. parvusDawangzhangzi BedJunior objective synonym of Liaoxiornis delicatus, a nomen dubium.
LongicrusavisL. houiDawangzhangzi BedAn ornithuromorph
LongirostravisL. hanibed unknownAn enantiornithine
ProtopteryxP. fengningensisPreviously regarded as Yixian, horizon reassigned to the Huajiying Formation by Jin et al., 2008.[28]
MirusavisM. parvusPartial skeleton.[29] An immature female enantiornithine preserved with medullary bone tissue.
ShanweiniaoS. cooperorumDawangzhangzi BedAn enantiornithine.[30]
VescornisV. hebeiensisPreviously regarded as Yixian, horizon reassigned to the Huajiying Formation by Jin et al., 2008.
YixianosaurusY. longimanusDawangzhangzi BedAn anchiornithid
ZhongornisZ. haoaeDawangzhangzi BedA primitive avialan with a short but non-pygostylian tail[31]

Compsognathids

GenusSpeciesCountyMemberMaterialNotesImages
HuaxiagnathusH. orientalisBeipiaoJianshangou BedA compsognathid
SinocalliopteryxS. gigasBeipiaoJianshangou BedA large compsognathid
SinosauropteryxS. primaBeipiao & LingyuanJianshangou & Dawangzhangzi BedsFour nearly complete specimens[32] [33] A compsognathid. The first non-avian dinosaur found with feather impressions.
S.? sp.BeipiaoJianshangou BedOne specimen (GMV 2124) with partial skull, postcrania, and preserved feathersAn unnamed coelurosaur originally described as a specimen of Sinosauropteryx, later found to be a new genus and species.[34]

Dromaeosaurs

GenusSpeciesCountyMemberMaterialNotesImages
Changyuraptor[35] C. yangiJianchangA complete articulated specimen with extensive feathering.A dromaeosaurid
GraciliraptorG. lujiatunensisBeipiaoLujiatun BedAn incomplete specimen.A dromaeosaurid
SinornithosaurusS. haoianaYixianDawangzhangzi BedMight be a junior synonym of Sinornithosaurus millenii.
S. milleniiBeipiaoJianshangou BedThree specimens with skulls and postcrania, two with preserved feathers.[36] Feathered integument is also preserved.[37] A dromaeosaurid, the first ever found with feather impressions.
TianyuraptorT. ostromiLingyuanDawangzhangzi BedA dromaeosaurid
Zhenyuanlong[38] Z. suniJianchangOne nearly complete skeleton with preserved feathers, including long tail feathers and large wing feathers.A dromaeosaurid
Zhongjianosaurus[39] Z. yangiLingyuanA partial skeleton includes an arm, legs, vertebrae, ribs, and most of the tail. A microraptorian dromaeosaurid with very long legs.
Unnamed dromaeosauridSpecimen NGMC 91LingyuanDawangzhangzi BedComplete specimen with feather imprintsA juvenile dromaeosaurid that may be a specimen of Sinornithosaurus millenii.[40] [41] Informally nicknamed "Dave".

Ornithomimosaurs

GenusSpeciesCountyMemberMaterialNotesImages
HexingH. qingyiBeipiaoLujiatunA primitive ornithomimosaur[42]
ShenzhousaurusS. orientalisBeipiaoLujiatun Bed"Articulated skull and incomplete skeleton."[43] A primitive ornithomimosaur

Oviraptorosaurs

GenusSpeciesCountyMemberMaterialNotesImages
CaudipteryxC. dongiBeipiaoJianshangou Bed"[One] partial postcranium with feather imprints."[44] Oviraptorosaurs
C. zouiBeipiaoJianshangou BedCommon, only over a small region
C. sp.[45]
IncisivosaurusI. gauthieriBeipiaoLujiatun Bed"[One] skull [and a] partial cervical."An oviraptorosaur
NingyuansaurusN. wangiComplete skeleton with feather impressions.[46] An oviraptorosaur
ProtarchaeopteryxP. robustaBeipiaoJianshangou Bed"Nearly complete skull and skeleton, at least [two] individuals."[47] A possible relative of Incisivosaurus
SimilicaudipteryxS. yixianensisTwo specimensAn oviraptorosaur, also present in the Jiufotang Formation[48]
XingtianosaurusX. ganqiLiaoningDakangpu BedOne nearly complete specimen (IVPP V13390), minus the headAn oviraptorosaur[49]

Therizinosaurs

GenusSpeciesCountyMemberMaterialNotesImages
BeipiaosaurusB. inexpectusBeipiaoJianshangou Bed"Incomplete skeleton with integument."[50] A therizinosaur
JianchangosaurusJ. yixianensisJianchangA nearly complete juvenile specimen with feather impressions.[51] A therizinosaur

Troodontids

GenusSpeciesCountyMemberMaterialNotesImages
DaliansaurusD. liaoningensisBeipiaoLujiatun BedA very complete specimen.[52] A troodontid
JianianhualongJ. tengiBeipiaoDawangzhangzi BedA very complete specimen with feather impressions.[53] A troodontid
LiaoningvenatorL. currieiBeipiaoA very complete specimen.A troodontid
MeiM. longBeipiaoLujiatun BedA small troodontid fossilized in a sleeping position
SinovenatorS. changiiBeipiaoLujiatun BedThree specimens with partial skulls and postcrania.[54] [55] Hundreds of additional specimens known but undescribed.A troodontid
SinusonasusS. magnodensBeipiaoLujiatun BedA troodontid
Undescribed deinonychosaur/possible troodontidDawangzhangzi BedOne specimen with feather imprintsA possible troodontid.[56]
Undescribed troodontidSpecimen CAGS-IG01-004Lujiatun BedSkull and partial skeletonA troodontid similar to Mei.[57]

Tyrannosaurs and other theropods

GenusSpeciesCountyMemberMaterialNotesImages
DilongD. paradoxusBeipiaoLujiatun & Jianshangou BedRareA tyrannosauroid
Grallatorlower Jianshangou BedFootprints belonging to the Grallator form taxon, belonging to a small theropod. Of known Yixian theropods, the footprints most closely match Caudipteryx.[58]
RaptorexR. kriegsteiniA tyrannosaurid, probably from higher Mongolian strata.
ScansoriopteryxS. heilmanniInitially reported from the Dawangzhangzi Bed, probably from the Daohugou Beds instead.[59]
YutyrannusY. hualiUnknownThree nearly complete skeletons of various agesA large tyrannosauroid[60]
Unnamed theropodNot yet describedYixianLujiatun BedPartial post-cranial skeleton[61] A possible carnosaur[62]

Pterosaurs

The pterosaurs listed below were compiled in a survey by Wang and Zhou in 2006, unless otherwise noted.[63]

GenusSpeciesCountyMemberMaterialNotesImages
BeipiaopterusB. chenianusJianshangou BedA ctenochasmatid
BoreopterusB. cuiaeJianshangou BedA boreopterid
CathayopterusC. grabauiDawangzhangzi BedA ctenochasmatid
Elanodactylus[64] E. prolatusJianshangou BedA ctenochasmatid
EopteranodonE. lii[65] BeipiaoA tapejarid
E. yixianensis[66] BeipiaoJianshangou BedA tapejarid
EosipterusE. yangiJianshangou BedA ctenochasmatid
Gegepterus[67] G. changaeJianshangou BedA ctenochasmatid
GladocephaloideusG. jingangshanensisJianshangou BedA ctenochasmatid
HaopterusH. gracilisJianshangou BedAn ornithocheirid
Luchibang[68] L. xingzheOne nearly complete articulated skeleton (ELDM 1000)An istiodactylid, the most complete known
MoganopterusM. zhuianaLujiatun bed?A ctenochasmatid
NingchengopterusN. liuaeJianshangou or Dawangzhangzi bedA pterodactyloid[69]
NurhachiusN. sp.Jingangshan Member[70] An incomplete specimenAn istiodactylid
PterofiltrusP. qiuiJianshangou BedA ctenochasmatid[71]
YixianopterusY. jingangshanensisAn istiodactyliform
ZhenyuanopterusZ. longirostrisJianshangou BedA boreopterid.[72] May be adult form of Boreopterus.[73]

Misc. reptiles

Choristoderans

GenusSpeciesCountyMemberMaterialNotesImages
HyphalosaurusH. lingyuanensisDawangzhangzi Bed
H. baitaigouensisJingangshan BedThousands of specimens
MonjurosuchusM. splendensDawangzhangzi Bed
SinohydrosaurusS. lingyuanensisDawangzhangzi BedJunior synonym of Hyphalosaurus lingyuanensis

Lizards

GenusSpeciesCountyMemberMaterialNotesImages
DalinghosaurusD. longidigitusJianshangou BedA lizard related to shinisaurs
LiushusaurusL. acanthocaudata[74] Jianshangou or Dawangzhangzi bedA lizard related to geckos and skinks
XianglongX. zhaoiJianshangou BedA gliding lizard related to chameleons and dragon lizards
YabeinosaurusY. tenuisDawangzhangzi & Jingangshan BedsA lizard related to geckos and skinks

Turtles

GenusSpeciesCountyMemberMaterialNotesImages
ManchurochelysM. liaoxiensisJianshangou BedReclassified as Ordosemys liaoxiensis[75]
M. manchoukuoensis[76] Jingangshan Bed
OrdosemysO. liaoxiensisJianshangou Bed

Mammals

The mammals listed below were compiled in a survey by Meng and colleagues in 2006, unless otherwise noted.[77]

GenusSpeciesCountyMemberMaterialNotesImages
AcristatheriumA. yanensis[78] BeipiaoLujiatun BedA eutherian
AkidolestesA. cifelliiLingyuanDawangzhangzi BedA spalacotheriid "symmetrodont"
Ambolestes[79] A. zhouiNinchengJianshangou BedA eutherian
Chaoyangodens[80] C. liiLingyuanDawangzhangzi BedA eutriconodont
EomaiaE. scansoriaLingyuanDawangzhangzi BedA tribosphenidan, possibly a eutherian or stem-therian
GobiconodonG. zofiaeBeipiaoLujiatun BedA gobiconodont
G. luoianus[81] Lujiatun BedA gobiconodont; junior synonym of G. zofiae
JeholodensJ. jenkinsiChaoyangJianshangou BedA eutriconodont
Juchilestes[82] J. liaoningensisBeipiaoLujiatun BedAn amphidontid
Jueconodon[83] J. cheniA eutriconodont
MaotheriumM. sinensisChaoyangJianshangou BedA zhangeotheriid "symmetrodont"
M. asiaticus[84] BeipiaoLujiatun BedA zhangeotheriid "symmetrodont"
MeemannodonM. lujiatunensisBeipiaoLujiatun BedA gobiconodont
OrigolestesO. liiA zhangeotheriid "symmetrodont"
RepenomamusR. giganticusBeipiaoLujiatun BedA gobiconodont
R. robustusBeipiaoLujiatun BedA large gobiconodont, which ate small dinosaurs
SinobaatarS. lingyuanensisLingyuanJianshangou and Dawangzhangzi BedsA multituberculate
SinodelphysS. szalayiLingyuanDawangzhangzi BedA therian, initially interpreted as a metatherian and later reinterpreted as a eutherian
Yanoconodon[85] Y. alliniFengningA eutricondodont
ZhangheotheriumZ. quinquecuspidensChaoyangJianshangou BedA zhangeotheriid "symmetrodont"

Amphibians

GenusSpeciesCountyMemberMaterialNotesImages
CallobatrachusC. sanyanensisSihetunJianshangou BedJunior synonym of Liaobatrachus
LaccotritonL. subsolanusPaozhanggouFengshan BedA urodelan
LiaobatrachusL. grabauiSihetunJianshangou BedA spadefoot toad
L. zhaoiBeipiao & HeitizigouLujiatun Bed & Jianshangou BedA spadefoot toad
MesophryneM. beipiaoensisBeipiao & HeitizigouLujiatun Bed & Jianshangou BedJunior synonym of Liaobatrachus
SinerpetonS. fengshanensisPaozhanggouFengshan BedA urodelan

Fish

GenusSpeciesCountyMemberAbundanceNotesImages
Hybodontoidea indet.Family, genus and species indeterminateAn elasmobranch known from a single dorsal fin spine[86]
LycopteraL. davidiJianshangou & Dawangzhangzi BedsAlso present in the Jiufotang Formation & Huajiying FormationAn osteoglossiform
L. muroiiJingangshan Bed
L. sinensisJianshangou Bed
MesomyzonM. mengaeA lamprey
PeipiaosteusP. paniJianshangou & Dawangzhangzi BedAlso present in the Jiufotang FormationA peipiaosteid fish related to sturgeons
ProtopsephurusP. liuiDawangzhangzi BedAlso present in the Jiufotang & Huajiying FormationsA paddlefish
SinamiaS. liaoningensisJianshangou BedA bowfin
YanosteusY. longidorsalisAlso present in the Huajiying FormationA peipiaosteid

Invertebrate fauna of the Yixian Formation

The invertebrate species listed below follow a survey by Chen and colleagues published in 2005, unless otherwise noted.[87]

Arachnids

The arachnid species listed below follow a catalogue compiled by Dunlop, Penney and Jekel in 2010, unless otherwise noted.[88]

GenusSpeciesCountyMemberAbundanceNotesImages
?AraneusA.? aethusOrb-weaver spiders
A.? beipiaoensis
A.? liaoxiensis
A.? reheensis
CretaraneusC. liaoningensisA nephiline spider
?GnaphosaG.? liaoningensisA ground spider
?TheridionT.? atalusA tangle web spider

Beetles (Coleoptera)

GenusSpeciesCountyMemberAbundanceNotesImages
AnoixisA. complanusA desmatin click beetle[89]
ApoclionA. antennatusDesmatin click beetles
A. clavatus
A. dolini
CoptoclavaC. longipodaA beetle
DesmatinusD. cognatusA desmatin click beetle
FortiseodeF. pervalimandAn eodromeine trachypachid beetle[90]
GlaresisG. orthochilusA laresid scarab beetle[91]
HydrophilopsiaH. gracilisHydrophiloid water scavenger beetles[92]
H. hydraenoides
H. shatrovskiyi
LasiosyneL. integeraHuangbanjigou Village, Liaoning Province, northeastern China.[93] A lasiosynid beetle.
L. parvaHuangbanjigou Village, Liaoning Province, northeastern China.A lasiosynid beetle.
LiaoximordellaL. hongiA mordellid beetle
MirimordellaM. gracilicruralisA mordellid beetle[94]
OxyporusO. yixianusAn oxyporine rove beetle[95]
ParadesmatusP. dilatatusA desmati click beetle
ParageotrupesP. incanusA Geotrupine scarab beetle[96]
SinocupesS. validusA beetle
SinosperchopsisS. silinaeA hydrophiloid water scavenger beetle
SinoxytelusS. breviventerOxyteline rove beetles[97]
S. euglypheus
S. longisetosus

Crustaceans

GenusSpeciesCountyMemberAbundanceNotesImages
Chenops[98] C. oblongusNotostracansframeless
C. yixianensis
CricoidoscelosusC. aethusJunior synonym of Palaeocambarus[99]
CyprideaC. (Cypridea) liaoningensisJianshangou BedOstracods
C. (C.) sihetunensisJianshangou Bed
C. (C.) veridica argutaJingangshan Bed
C. (C.) sp.Jianshangou Bed
C. (Ulwellia) beipiaoensisBeipiaoJianshangou Bed
DamonellaD. formosaJianshangou BedAn ostracod
DarwinulaD. leguminellaJianshangou BedAn ostracod
DiestheriaD. jeholensisJianshangou BedClam shrimps
D. yixianensisDawangzhangzi Bed
D.? sp.Jianshangou Bed
EosestheriaE. jingangshanensisJingangshan BedClam shrimps
E. lingyuanensisDawangzhangzi Bed
E. aff. middendorfiiJingangshan Bed
E. ovataJianshangou Bed
E. sihetunensisJianshangou Bed
EosestheriopsisE. gujialingensisJianshangou BedA clam shrimp
JeholopsJ. hongiA notostracanframeless
LiaoningogriphusL. quadripartitusJianshangou and Dawangzhangzi bedsA shrimp
LycopterocyprisL. infantilisJianshangou BedAn ostracod
MantellianaM. sp.Jianshangou BedAn ostracod
"Mongolarachne""M." chaoyangensisDawangzhangzi BedsJunior synonym of Palaeocambarus[100]
MongolianellaM. cf. palmosaJianshangou BedAn ostracod
PalaeocambarusP. licentiA crayfish[101]
RhinocyrpisR. jurassicaJianshangou BedAn ostracod
TimiriaseviaT. jianshangouensisJianshangou BedAn ostracod
?YanjiestheriaY.? beipiaoensisBeipiaoJianshangou BedA clam shrimp
YanshaniaY. dabeigouensisJianshangou BedAn ostracod

Dragonflies (Odonata)

The dragonfly species listed below follow a review compiled by Zhang, Ren and Panf in 2008, unless otherwise noted.[102]

GenusSpeciesCountyMemberAbundanceNotesImages
AbrohemeroscopusA. mengiA dragonfly
AeschnidiumA. heishankowenseReclassified as Sinaeschnidia heishankowense
A. sp.
ArchaeogomphusA. labiusNymphsReclassified as Yixiangomphus labius,[103] which is a junior synonym of Sinaeschnidia cancellosa
BellabrunetiaB. catherinaeA dragonfly
ChrysogomphusC. beipiaoensisA dragonfly
DissurusD. liaoyuanensisPoorly preserved nymphReclassified as Sinaeschnidia cf. cancellosa
DracontaeschnidiumD. orientaleA dragonfly
HebeiaeschnidiaH. fengningensisA dragonfly
HuaxiagomphusH. taushanensisA dragonfly
LiogomphusL. yixianensisA dragonfly
MesocorduliaM. borealaA corduliid dragonfly
PalaeogomphusP. labiusA dragonfly, reclassified as of Yixiangomphus labius, which is a junior synonym of Sinaeschnidia cancellosa
ParapetalaP. liaoningensisA dragonfly
P. yixianensisJianshangou Unit[104] An aeshnopteran dragonfly
ProtoliupanshaniaP. wangiJianshangou UnitA liupanshaniid dragonfly
PseudocymatophlebiaP. bisectaNingcheng County[105] A well-preserved specimen.An aktassiid dragonfly
PseudoliupanshaniaP. magnanicellulaJianshangou Unit
PseudosamaruraP. larginaOne specimen, nymphReclassified as Sinaeschnidia cf. cancellosa
RudiaeschnaR. limnobiaAn aeshnid dragonfly[106]
SinaeschnidiaS. heishankowenseJianshangou & Dawangzhangzi BedsAdults and nymphs
S. cancellosaJunior synonym of S. heishankowense[107]
S. sp.Indeterminate nymph specimensSome nymph specimens have been given their own genera and species though probably belong so S. heishankowense but too poorly preserved to be certain
SinogomphusS. taoshanensisA dragonfly
SinojagoriaS. imperfectaA dragonfly
SopholibellulaS. amoenaDragonflies
S. eleganti
StylaeschnidiumS. rarumA dragonfly
TelmaeshnaT. paradoxicaBeipiaoA dragonfly
TuranophlebiaT. liaoningensisBeipiao[108] Jianshangou UnitA tarsophlebiid dragonfly.
YixiangomphusY. labiusNymphsJunior synonym of Sinaeschnidia cancellosa[109]

Flies (Diptera)

GenusSpeciesCountyMemberAbundanceNotesImages
AlleremonomusA. xingiJianshangou BedA brachyceran fly
AtalosciophilaA. yanensisA mycetophilid gnat[110]
ChironomapteraC. gregariaA phantom midge
GigantoberisG. liaoningensisA soldier fly[111]
LichnopleciaL. kovaleviA protopleciid fly[112]
PalaepangoniusP. eupterusA horse-fly
ProtapioceraP. megistaAn asiloid fly
ProtempisP. minutaA brachyceran fly
ProtonemestriusP. jurassicusA tangle-veined fly

Wasps (Hymenoptera)

GenusSpeciesCountyMemberAbundanceNotesImages
AbropelecinusA. annulatusBeipiaoAn iscopinine pelecinid[113]
AngaridyelaA. robustaJianshangou BedA sawfly
AzygopelecinusA. clavatusBeipiaoAn iscopinine pelecinid
ChengdeserphusC. petidatusA proctotrupid wasp
CretevaniaC. exquisitaBeipiaoHatchet wasps[114]
C. pristineJianshangou bed
C. vesca
HoplitolydaH. duolunicaThe largest known sawfly ever[115]
LiaotomaL. linearisA xyelotomid woodwasp
ManlayaM. flexuosusAn aulacid woodwasp
ProcretevaniaP. exquisitaReclassified as Cretevania spp.[116]
P. pristine
P. vesca
RudisiriciusR. belliPraesiricid sawflies[117]
R. celsus
R. crassinodus
ScolichneumonS. rectiveniusAn ichneumonoid wasp
SinopelecinusS. viriosusA pelecinid
TanychoraT. exquisitaJianshangou BedAn ichneumon wasp

Other insects

GenusSpeciesCountyMemberAbundanceNotesImages
AbrigrammaA. calophlebaA kalligrammatid lacewing[118]
AlloraphidiaA. bliquivenaticaSnakeflies
A. longistigmasaJianshangou Bed
ArcheraphidiaA. shangyuanensisA snakefly
BaissopterusB. liaoningensisA snakefly
CaenoephemeraC. shangyuanensisJianshangou BedA mayfly[119]
CaloraphidiaC. glossophyllaA snakefly
ChoromyrmeleonC. othneiusAn antlion
CirculaboilusC. aureusA grig[120]
CretophasmomimaC. melanogramma A stick insect[121]
?Ephemeropsis?E. trisetalisJianshangou and Jingangshan bedsA mayfly. Chinese specimens probably represent species of Epicharmeropsis
EpicharmeropsisE. hexavenulosus[122] PingquanMayflies
E. quadrivenulosusBeipiaoJianshangou Bed
HagiphasmaH. paradoxaJianshangou BedA stick insect
IlerdocossusI. beipiaoensisPalaeontinid hemipterans[123] [124]
I. exiguus
I. exquisitus
I. fengningensis
I. hui
I. pingquanensis
IthigrammaI. multinerviaKalligrammatid lacewings
KalligrammaK. liaoningensisJianshangou Bed
KaratavoblattaK. formosaA cockroach
LiaoaphidiaL. fornicataA snakefly
LiaonemobiusL. tanaeA sword-tail cricket
LongicellochrysaL. yixianaA mesochrysopid lacewing[125]
MesascalaphusM. yangiAn owlfly
MesolygaeusM. laiyangensisA water bug
MetabittacusM. colosseusJianshangou BedA scorpionfly[126]
MiracossusM. gongiPalaeontinid hemipterans
M. ingentius
NipponoblattaN. acerbaJianshangou BedA cockroach
OloberothaO. sinicaA beaded lacewing
OregrammaO. aureolosaKalligrammatid lacewings
O. illecebrosa
PseudopulexP. magnusStem-fleas[127] [128]
P. tanlan
RenphasmaR. sinicaA stick insect[129]
SaurophthirusS. exquisitusStem-fleas[130]
S. laevigatus
SinochresmodaS. magnicorniaA chresmodid water-skiing insect[131]
SinosharaperlaS. zhaoiBeipiaoA stonefly[132]
SinoviparosiphumS. liniAn aphid
SophogrammaS. liiJianshangou BedKalligrammatid lacewings[133]
S. pingquanica
TetraphelerusT. laetusA moth
TyrannopsyllaT. beipiaoensisA stem-flea[134]
VaricapitatusV. sinuolatus"Huangbanjigou, Chaomidian Village, Beipiao City."[135] A hemipteran
Unnamed VitimotauliidaeUnnamedA caddisfly pupa, likely belonging to the Vitimotauliidae[136]

Molluscs

GenusSpeciesCountyMemberAbundanceNotesImages
AmplovalvataA. sp.Jianshangou BedA gastropod
ArguniellaA. lingyuanensisJianshangou BedA bivalve, also present in the Jiufotang and Fuxin Formations
A. yanshanensisJianshangou BedA bivalve, also present in the Jiufotang Formation
BithyniaB. haizhouensisJingangshan BedA gastropod
CorbiculaC. anderssoniJianshangou BedReclassified as Sphaerium anderssoni
C. jeholenseReclassified as Sphaerium jeholense
GyraulusG. sp.Jianshangou BedA ramshorn snail
ProbaicaliaP. vitimensisJianshangou BedA gastropod
PtychastylusP. harpaeformisJianshangou BedGastropods
P. cf. phillipiJianshangou Bed
SphaeriumS. anderssoniJianshangou BedA fingernail clam, also present in the Jiufotang Formation
S. jeholenseJunior synonym of S. anderssoni
S. selenginenseJunior synonym of S. anderssoni
?ViviparusV.? cf. matsumotoiJingangshan BedA snail

Flora of the Yixian Formation

Conifers

The coniferous trees and shrubs listed below were compiled in a survey by Wang and colleagues in 2006, unless otherwise noted.[137]

GenusSpeciesCountyMemberAbundanceNotesImages
BrachyphyllumA cypress-like conifer
CephalotaxopsisA yew
CunninghamiaA cypress
ElatidesA cypress
ElatocladusE. leptophyllusConifers of uncertain classification
E. manchurica
LiaoningocladusL. boiiJingangshan BedA conifer of uncertain classification
PagiophyllumA cypress-like conifer
PityocladusA pine
PityolepisP. sp.A pine
PityophyllumP. cf. lindstroemiA pine
PityostrobusA pine
PodozamitesP. lanceolatusA conifer of uncertain classification
SchizolepisS. jeholensis
?SequoiaS.? jeholensis
TaxusA yew
TorreyaA yew
YanliaoiaY. sinensis

Ferns

GenusSpeciesCountyMemberAbundanceNotesImages
CladophlebisC. spp.
ConiopterisC. sp.
OnychiopsisO. elongataJunior synonym of O. psilotoides
O. psilotoides
?RuffordiaR.? sp.

Flowering plants

GenusSpeciesCountyMemberAbundanceNotesImages
ArchaefructusA. eofloraJianshangou BedA primitive, aquatic angiosperm
A. liaoningensisJianshangou Bed
A. sinensisisDawangzhangzi Bed
CallianthusC. dilae[138] LiaoningA plant with true flowers
?DavalliaD.? niehhutzuensis
EreniaE. stenopteraLiaoningActually a specimen of Callianthus
LeefructusL. mirus[139] A eudicot
LiaoxiaL. chenii
?PotamogetonP.? jeholensis
RanunculusR. jeholensisJunior synonym of Potamogeton jeholensis
SinocarpusS. decussatusA primitive eudicot

Ginkgos

GenusSpeciesCountyMemberAbundanceNotesImages
BaieraB. cf. gracilis
B. orientalis
CzekanowskiaC. rigida
GinkgoG. apodes[140]
GinkgoitesG. sibiricus
LeptostrobusL. sp.
?NageiopsisN.? sp.
PhoenicopsisP. cf. augustifolia
SolenitesS. sp.
SphenobaieraS. longifola

Other plants

GenusSpeciesCountyMemberAbundanceNotesImages
ChaoyangiaC. liangii[141] A gnetale gymnosperm
ChaoyangicarpusC. liangiiJunior synonym of Chaoyangia liangii
EphedraE. sp.A gymnosperm
EquisetumE. sp.A horsetail
EragrositesE. changiiA grass-like gnetale gymnosperm
SiphonospermumS. simplex[142] BeipiaoJianshangou BedA gnetale gymnosperm
TyrmiaT. sp.A bennettite

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