Paleobiota of the Solnhofen Limestone explained

Solnhofen Limestone
Type:Geological formation
Age:Kimmeridgian-Tithonian[1]
Prilithology:Lithographic limestone[2]
Period:Jurassic
Otherlithology:Slate,Dolomite, Flint
Namedfor:Solnhofen
Region:Bavaria
Country: Germany
Underlies:Usseltal Formation
Overlies:Treuchtlingen Formation
Map:Map of the Ettling quarry - Journal.pone.0116140.g001.png

The Solnhofen Limestone or Solnhofen Plattenkalk is a collective term for multiple Late Jurassic lithographic limestones in southeastern Germany, which is famous for its well preserved fossil flora and fauna dating to the late Jurassic (Kimmeridgian-Tithonian). The paleoenvironment is also often referred to as the Solnhofen Archipelago. The Solnhofen Archipelago was located at the northern edge of the Tethys Ocean as part of a shallow epicontinental sea and is firmly a part of the Mediterranean realm.[3]

Chondrichthyes

Holocephali

!Genus!Species!Locality!Formation!Material!Notes!Images
Chimaeropsis[4] C. paradoxaSolnhofen, EichstättAltmühltal FormationFragmentary cranial remains and one complete specimen[5] Holotype destroyed in WW2 airraid
IschyodusI. egertoniEichstätt, WattendorfAltmühltal Formation, Torleite FormationMultiple complete specimens
I. quenstedtiMay be a junior synonym of I. egertoni
ElasmodectesE. avitusEichstätt, Blumenberg, WintershofAltmühltal FormationMultiple complete specimens
ChimaerothecaC. schernfeldensisEgg capsules of an unknown chimaera

Elasmobranchii

!Genus!Species!Locality!Formation!Material!Notes!Images
Hybodus (Egertonodus?)H. fraasiSolnhofenAltmühltal FormationAlmost complete but poorly preserved specimen
AsteracanthusA. ornatissimusSolnhofenAltmühltal FormationSingle almost complete specimen of a 2 meter long female
StrophodusS. spMühlheimMörnsheim FormationArticulated set of jaws and teeth

Neoselachii

!Genus!Species!Locality!Formation!Material!Notes!Images
SphenodusS. nitidusSolnhofenAltmühltal FormationIncomplete skeletal remains and isolated teeth
S. macerSolnhofenAltmühltal FormationIncomplete skeletal remains and isolated teeth
SynechodusS. ungeriSolnhofenAltmühltal FormationAlmost complete skeleton and isolated teeth
ParaorthacodusP. jurensisHaunsfeld, Blumenberg, Schernfeld, SolnhofenAltmühltal FormationSeveral complete specimens
MacrourogaleousM. hasseiEichstättAltmühltal FormationSeveral complete specimens

Batomorphii

!Genus!Species!Locality!Formation!Material!Notes!Images
AsterodermusA. platypterusSolnhofen, Kelheim, Blumenberg, BirkhofAltmühltal FormationSeveral complete specimens
SpathobatisS. bugesiacusEichstätt, SolnhofenAltmühltal FormationSeveral complete specimens
RajithecaR. spSeveral specimens in private collectionsEgg capsules of Batomorphs

Heterodontiformes

!Genus!Species!Locality!Formation!Material!Notes!Images
ParacestracionP. falciferSolnhofen? EichstättAltmühltal FormationComplete skeleton, potentially more specimens
P. viohliSchamhauptenTorleite FormationSingle incomplete skeleton
P. danieliEichstättAltmühltal FormationSingle incomplete skeleton
HeterodontusH. zitteliEichstättAltmühltal FormationSingle complete skeleton lacking teeth, several other specimens may be referrable

Orectolobiformes

!Genus!Species!Locality!Formation!Material!Notes!Images
PhorcynisP. catulinaEichstätt, Zandt, BlumenbergAltmühltal FormationSeveral skeletons
PalaeorectolobusP. agomphiusKelheimAltmühltal FormationOne incomplete specimen

Carcharhiniformes

!Genus!Species!Locality!Formation!Material!Notes!Images
BavariscylliumB. tischlingeriEichstättAltmühltal FormationSeveral complete specimens
PalaeoscylliumP. formosumSolnhofenAltmühltal FormationSeveral complete specimens
ScyliorhinothecaS. spegg capsule of a catshark

Lamniformes

!Genus!Species!Locality!Formation!Material!Notes!Images
PalaeocarchariasP. stromeriEichstätt, Kelheim, SolnhofenAltmühltal FormationSeveral complete specimens

Hexanchiformes

!Genus!Species!Locality!Formation!Material!Notes!Images
CrassodontidanusC. serranusEichstätt, DaitingAltmühltal FormationIsolated Teeth
NotidanoidesN. muensteriEichstätt, DaitingAltmühltal FormationIsolated teeth, one almost complete skeleton was presumably destroyed during WW2Another incomplete skeleton of the Genus is known from the Nusplingen limestone

Squatiniformes

!Genus!Species!Locality!Formation!Material!Notes!Images
PseudorhinaP. acanthodermaAltmühltal FormationSeveral complete specimens
P. aliferaEichstättAltmühltal FormationSeveral complete specimens

Other

!Genus!Species!Locality!Formation!Material!Notes!Images
ProtospinaxP. annectansSolnhofen, EichstättAltmühltal FormationSeveral complete specimens

Osteognathostomata

Chondostrei

GenusSpeciesLocalityFormationMaterialNotesImages
Coccolepis[6] C. bucklandiEichstätt, BlumenbergAltmühltal FormationSeveral complete Specimens
C. solnhofensis[7] SolnhofenAltmühltal Formation

Pycnodontiformes

GenusSpeciesLocalityFormationMaterialNotesImages
Arduafrons[8] A. prominorisBlumenberg, Wintershof, PaintenAltmühltal Formation, Torleite FormationSeveral complete specimens
Apomesodon[9] A. comosusEichstättAltmühltal Formation
A. spEichstätt, PaintenAltmühltal Formation, Torleite Formation
Eomesodon[10] E. gibbosusBlumenberg, Wintershof, PaintenAltmühltal Formation, Torleite FormationSeveral complete specimens
GyrodusG. circularisEichstätt, PaintenAltmühltal Formation, Torleite FormationDozens of complete specimens
G. hexagonusSolnhofen, Eichstätt, Painten, Schamhaupten, BrunnAltmühltal Formation, Torleite FormationDozens of complete specimens
MesturusM. verrucosusEichstätt, PaintenAltmühltal Formation, Torleite FormationSeveral complete specimens
MacromesodonM. heckeli
M. macropterusKelheimTorleite Formation
Piranhamesodon[11] P. pinnatomusEttlingSingular partial skeleton
ProscinetesP. elegansEichstätt. EttlingAltmühltal Formation, Torleite FormationDozens of complete specimens
P. formosus
P. bernardi[12] Schamhaupten, Ettling, PaintenTorleite FormationSeveral complete specimens
Turbomesodon[13] T. relegansEichstätt, EttlingAltmühltal Formation, Torleite Formation
TurboscinetesT. egertoniEttling
Thiollierepycnodus[14] T. wagneriWattendorfTorleite Formation

Ginglymodi

Lepisosteiformes

GenusSpeciesLocalityFormationMaterialNotesImages
"Lepidotes"[15] "L." decoratus
"L." intermedius
"L." oblongus
"L." pustulosus
"L." subovatus
"L." unguiculatus
Scheenstia[16] S. maximus
S. decoratus
S. zappi[17] SchamhauptenTorleite FormationSingle almost complete specimen

Semionotiformes

GenusSpeciesLocalityFormationMaterialNotesImages
CallipurbeckiaC. notopterus
Macrosemimimus[18] M. fegertiEttlingMultiple complete specimens

Macrosemiidae

GenusSpeciesLocalityFormationMaterialNotesImages
Histionotus[19] H. oberndorferiKelheimTorleite FormationTwo incomplete specimens
MacrosemiusM. rostratusEichstätt, Solnhofen, PaintenAltmühltal Formation, Torleite FormationDozens of complete specimens
M. fournetiBrunnTorleite FormationOne complete specimen
NotagogusN. decoratus
N. denticulatusEichstätt, Schamhaupten, Painten, Brunn, Ettling?Altmühltal Formation, Torleite FormationDozens of complete specimens
Palaeomacrosemius[20] P. thiollieriEichstätt, Solnhofen, EttlingAltmühltal FormationSeveral complete specimens
PropterusP. elongatusEichstättAltmühltal FormationSeveral complete specimens
P. microstomusEichstättAltmühltal FormationSeveral complete specimens
Eusemius?E. beatae
Voelklichthys[21] V. comitatusPaintenTorleite FormationSingular specimen

Dapediiformes

Notes and References

  1. Schweigert, G. 2007. Ammonite biostratigraphy as a tool for dating Upper Jurassic lithographic limestones from South Germany – first results and open questions." Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 245(1): 117-125.
  2. Rauhut, O. W., Heyng, A. M., López-Arbarello, A., & Hecker, A. (2012). A new rhynchocephalian from the Late Jurassic of Germany with a dentition that is unique amongst tetrapods. PLoS ONE, 7(10): e46839.
  3. Tyborowski . Daniel . Błażejowski . Błażej . December 2019 . New marine reptile fossils from the Late Jurassic of Poland with implications for vertebrate faunas palaeobiogeography . Proceedings of the Geologists' Association . en . 130 . 6 . 741–751 . 10.1016/j.pgeola.2019.09.004.
  4. Duffin . Christopher J. . Lauer . Bruce . Lauer . René . 2022-11-30 . Chimaeroid egg cases from the Late Jurassic of the Solnhofen area (S Germany) . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 306 . 2 . 161–175 . 10.1127/njgpa/2022/1101 . 0077-7749.
  5. Villalobos-Segura . Eduardo . Stumpf . Sebastian . Türtscher . Julia . Jambura . Patrick . Begat . Arnaud . López-Romero . Faviel . Fischer . Jan . Kriwet . Jürgen . 2023-03-08 . A Synoptic Review of the Cartilaginous Fishes (Chondrichthyes: Holocephali, Elasmobranchii) from the Upper Jurassic Konservat-Lagerstätten of Southern Germany: Taxonomy, Diversity, and Faunal Relationships . Diversity . 15 . 3 . 386 . 10.3390/d15030386 . free . 36950327 . 7614348 . 1424-2818.
  6. "Table 1; Chondrostei; Coccolepidae" in Lambers (1999) pg. 216.
  7. López-Arbarello . Adriana . Ebert . Martin . 2021 . Diversity of Coccolepis AGASSIZ, 1843 (Coccolepididae, Chondrostei) from the Upper Jurassic Solnhofen-Archipelago, Southern Germany . Acta Palaeontologica Polonica . 66 . 10.4202/app.00873.2021. free .
  8. Web site: Jürgen . Kriwet . 2016-11-15 . Success and extinction of pycnodont fishes (Neopterygii, Pycnodontomorpha) . 10.55776/p29796.
  9. Kölbl-Ebert . Martina . Ebert . Martin . Bellwood . David R. . Schulbert . Christian . November 2018 . A Piranha-like Pycnodontiform Fish from the Late Jurassic . Current Biology . 28 . 21 . 3516–3521.e2 . 10.1016/j.cub.2018.09.013 . 30344113 . 2018CBio...28E3516K . 0960-9822. free .
  10. "Table 1; Halecostomi; Pycnodontiformes" in Lambers (1999) pg. 216.
  11. Kölbl-Ebert . Ebert . Bellwood . Schulbert . Martina . Martin . David R. . Christian . October 18, 2018 . A Piranha-like Pycnodontiform Fish from the Late Jurassic . Current Biology . 28 . 21 . 3516–3521.e2 . 10.1016/j.cub.2018.09.013 . 30344113 . 53045425 . free. 2018CBio...28E3516K .
  12. Ebert . E . 2018 . The Pycnodontidae (Actinopterygii) in the late Jurassic: 2) Turboscinetes gen. nov. in the Solnhofen Archipelago (Germany) and Cerin (France) (project) . 2024-06-21 . MorphoBank datasets. 10.7934/p3271 .
  13. Poyato-Ariza . Francisco Jose . Wenz . Sylvie . 2000-03-01 . A new pycnodontiform fish from the Early Cretaceous of Las Hoyas (Cuenca, Spain) . Bulletin de la Société Géologique de France . 171 . 2 . 251–258 . 10.2113/171.2.251 . 1777-5817.
  14. Ebert . Martin . 2019-10-21 . A new genus of Pycnodontidae (Actinopterygii) from the Upper Jurassic of France and Germany, included in a phylogeny of Pycnodontiformes . Zoological Journal of the Linnean Society . 10.1093/zoolinnean/zlz087 . 0024-4082.
  15. Lambers . Paul H. . 1999 . [No title found] ]. Geologie en Mijnbouw . 78 . 2 . 215–229 . 10.1023/A:1003855831015.
  16. "Table 1; Halecostomi; Semionotidae" in Lambers (1999) pg. 216.
  17. López-Arbarello . Adriana . Sferco . Emilia . June 2011 . New semionotiform (Actinopterygii: Neopterygii) from the Late Jurassic of southern Germany . Journal of Systematic Palaeontology . 9 . 2 . 197–215 . 10.1080/14772019.2010.493751 . 2011JSPal...9..197L . 11336/53446 . 1477-2019. free .
  18. Schröder . Kerstin M. . López-Arbarello . Adriana . Ebert . Martin . May 2012 . Macrosemimimus, gen. nov. (Actinopterygii, Semionotiformes), from the Late Jurassic of Germany, England, and France . Journal of Vertebrate Paleontology . 32 . 3 . 512–529 . 10.1080/02724634.2012.649626 . 2012JVPal..32..512S . 0272-4634.
  19. "Table 1; Halecostomi; Macrosemiidae" in Lambers (1999) pg. 216.
  20. Ebert . Martin . Lane . Jennifer A. . Kölbl-Ebert . Martina . 2016-09-02 . Palaeomacrosemius thiollieri, gen. et sp. nov., a new Macrosemiidae (Neopterygii) from the Upper Jurassic of the Solnhofen Archipelago (Germany) and Cerin (France), with a revision of the genus Macrosemius . Journal of Vertebrate Paleontology . 36 . 5 . e1196081 . 10.1080/02724634.2016.1196081 . 2016JVPal..36E6081E . 0272-4634.
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  24. "Table 1; Halecomorphi; Ionoscopidae" in Lambers (1999) pg. 216.
  25. Ebert . Martin . 2019-03-04 . Zandtfuro and Schernfeldfuro, new genera of Halecomorphi (Actinopterygii) from the Upper Jurassic Solnhofen Archipelago . Journal of Vertebrate Paleontology . 39 . 2 . e1592759 . 10.1080/02724634.2019.1592759 . 2019JVPal..39E2759E . 0272-4634.
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  30. "Table 1; Halecostomi; Pachycormidae" in Lambers (1999) pg. 216.
  31. Note: Lambers abbreviated Pseudoasthenocormus as "Pseudoasth.," "Table 1; Halecostomi; Pachycormidae" in Lambers (1999) pg. 216.
  32. Maxwell . Erin . Lambers . Paul . Lopez-Arbarello . Adriana . Schweigert . Günter . 2020 . Re-evaluation of pachycormid fishes from the Late Jurassic of Southwestern Germany . Acta Palaeontologica Polonica . 65 . 10.4202/app.00749.2020.
  33. "Table 1; Teleostei; Aspidorhynchidae" in Lambers (1999) pg. 216.
  34. "Table 1; Teleostei; Pholidophoridae" in Lambers (1999) pg. 216.
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  94. Haug . C. . Haug . J. . Waloszek . D. . 2009 . Morphology and ontogeny of the Upper Jurassic mantis shrimp Spinosculda ehrlichi n. gen. n. sp. from southern Germany . Palaeodiversity . 2 . 111–118. 2601897 .
  95. Haug . Carolin . Haug . Joachim T. . 2021-04-16 . A new fossil mantis shrimp and the convergent evolution of a lobster-like morphotype . PeerJ . 9 . e11124 . 10.7717/peerj.11124 . 2167-8359 . 8054755 . 33959413 . free .
  96. "Table 1; Halecostomi; 'Halecostomi inc. sed.'" in Lambers (1999) pg. 216.| H. latus|Solnhofen|Altmühltal Formation||| rowspan="1" | |}

    Halecomorphi

    Ionoscopidae

    GenusSpeciesLocalityFormationMaterialNotesImages
    Ainia[22] A. agassiziBrunnTorleite Formation
    Altmuehlfuro[23] A. boomerang
    BrachyichthysB. radiato-puncatus
    B. typicus
    Ionoscopus[24] I. cyprinoidesEichstättAltmühltal Formation
    I. desoriZandtTorleite Formation
    I. esocinus
    I. münsteri
    I. striatissimus
    "Furo"[25] "F." angustusSolnhofen, Schamhaupten, ZandtAltmühltal Formation, Torleite Formation
    "F." longiserratusEichstätt, Solnhofen, Kelheim, ZandtAltmühltal Formation, Torleite Formation
    "F." latimanusSolnhofen, EichstättAltmühltal Formation
    Ophiopsis[26] O. muensteriKelheim, Brunn, Ettling
    OphiopsiellaO. attenuata
    O. breviceps
    O. procera
    SchernfeldfuroS. uweelleriEichstättAltmühltal Formation
    ZandtfuroZ. tischlingeriZandtTorleite Formation

    Amiiformes

    !Genus!Species!Locality!Formation!Material!Notes!Images
    Amblysemius[27] A. bellicianusEichstättAltmühltal Formation
    A. pachyurus
    "A." granulatusKelheimAltmühltal Formation
    CaturusC. furcatusKelheim, Solnhofen, Ettling, Wattendorf?Altmühltal Formation, Torleite Formation
    "C." macrurusKelheimAltmühltal Formation
    "C." latusSolnhofenAltmühltal Formation
    "C." granulatusKelheimAltmühltal Formation
    "C." feroxSolnhofenAltmühltal Formation
    "Furo"[28] "F." microlepidotusEichstättAltmühltal Formation
    StrobilodusS. giganteusPaintenTorleite Formation
    LiodesmusL. gracilisSolnhofenAltmühltal Formation
    L. sprattiformisSolnhofen, EichstättAltmühltal Formation
    Lophiurus?L. minutus
    AmiopsisA. lepidotaEichstätt, Solnhofen, EttlingAltmühltal Formation
    SolnhofenamiaS. elongataEichstättAltmühltal Formation

    Pleuropholidae

    GenusSpeciesLocalityFormationMaterialNotesImages
    Pleuropholis[29] P. laevissimaDaiting, PaintenMörnsheim Formation, Torleite Formation
    P. pompecki
    P. wagneri

    Pachycormidae

    GenusSpeciesLocalityFormationMaterialNotesImages
    Asthenocormus[30] A. titaniusEichstätt, PaintenAltmühltal Formation, Torleite Formation
    HypsocormusH. insignisObereichstätt, SolnhofenAltmühltal Formation
    H. macrodon
    Leedsichthys?L. sp
    OrthocormusO. cornutusBirkhof, Blumenberg, SchernfeldAltmühltal Formation
    O. roeperiBrunnTorleite Formation
    O. teyleri?SchamhauptenTorleite Formation
    Pseudoasthenocormus[31] P. retrodorsalisLangenaltheimAltmühltal Formation
    PachycormidaeIndeterminateWattendorfTorleite FormationGiant skull fragment of a presumably 4 meter long pachycormid
    SauropsisS. longimanusEichstätt, BlumenbergAltmühltal Formation
    S. depressusBlumenberg, Schernfeld, EichstättAltmühltal Formation
    Simocormus[32] S. macrolepidotusSolnhofen, EichstättAltmühltal Formation

    Aspidorhynchidae

    GenusSpeciesLocalityFormationMaterialNotesImages
    Aspidorhynchus[33] A. acutirostrisEichstätt, Zandt, PaintenAltmühltal Formation, Torleite Formation
    A. ornatissimus
    A. sanzenbacheriEttling, Painten?Torleite Formation
    BelonostomusB. kochiKelheim, EttlingAltmühltal Formation, Torleite Formation?
    B. muensteriDaiting, Mühlheim, Solnhofen, Langenaltheim, Zandt, SchamhauptenMörnsheim Formation, Altmühltal Formation, Torleite Formation
    B. tenuirostrisSolnhofen, Zandt, LangenaltheimAltmühltal Formation, Torleite Formation
    B. speciosusSolnhofenAltmühltal Formation
    B. spyraenoidesEichstätt, Schernfeld, WintershofAltmühltal Formation

    "Pholidophoridae"

    GenusSpeciesLocalityFormationMaterialNotesImages
    Ankylophorus[34] Indeterminate.
    EurycormusE. speciosusEichstätt, Painten, ZandtAltmühltal Formation, Torleite Formation
    PholidophoristionP. micronyx
    P. ovatusEttling
    "Pholidophorus""P." armatus
    "P." diagonalis
    "P." falcifer
    "P." microps
    "P." sculptus
    "P." latimanus
    "P." latus
    "P." longiserratus
    "P." smilis
    Siemensichthys[35] S. macrocephalusKapfelbergAltmühltal Formation
    S. siemensiSchamhauptenTorleite Formation

    Crossognathiformes

    !Genus!Species!Locality!Formation!Material!Notes!Images
    BavarichthysB. incognitusEttlingTorleite Formation?Single complete skeletonThe oldest record of a crossognathiform in Europe

    Ichthyodectiformes

    GenusSpeciesLocalityFormationMaterialNotesImages
    Allothrissops[36] A. mesogaster
    A. salmoneusEichstätt, Painten, brunnAltmühltal Formation, Torleite Formation
    Ascalabothrissops[37] A. voelkliSchamhauptenTorleite Formation
    PachythrissopsP. propterusKelheim, Zandt, EttlingAltmühltal Formation, Torleite Formation
    ThrissopsT. formosusEichstätt, Kelheim, Mühlheim, EttlingAltmühltal Formation, Torleite Formation, Mörnsheim Formation
    T. subovatusEichstätt, Kelheim, EttlingAltmühltal Formation, Torleite Formation?

    Elopiformes

    !Genus!Species!Locality!Formation!Material!Notes!Images
    Anaethalion[38] "A." angustissimusEichstätt, PaintenAltmühltal Formation, Torleite Formation
    A. angustusLangenaltheim, Eichstätt, SchernfeldAltmühltal Formation
    A. knorri
    A. subovatus
    A. zapporumSchamhauptenTorleite Formation
    DaitingichthysD. tischlingeriDaitingMörnsheim Formation
    EichstaettiaE. mayriEichstättAltmühltal Formation
    ElopsomolosE. frickhingeriBlumenbergAltmühltal Formation
    E. sp 1
    E. sp 2
    E. sp 3

    Osteoglossomorpha

    GenusSpeciesLocalityFormationMaterialNotesImages
    Ascalabos[39] A. voithiiEichstätt, SchernfeldAltmühltal Formation
    Tischlingerichthys[40] T. viohli
    Tharsis[41] T. dubiusEichstätt, PaintenAltmühltal Formation, Torleite Formation
    T. elleriSchernfeldAltmühltal Formation
    T. spWattendorfTorleite Formation
    EbertichthysE. ettlingensisEttlingTorleite Formation?
    Leptolepides[42] L. haerteisiZandt, SchamhauptenTorleite Formation
    L. sprattiformisSolnhofen, Eichstätt, Kelheim, Langenaltheim, Schernfeld, BlumenbergAltmühltal FormationMultiple Mass mortality plates of this species are known.It represents the most common vertebrate of the Solnhofen limestone.
    OrthogonikleithrusO. hoelliEttlingTorleite Formation?
    O. leichiEttlingTorleite Formation?

    Coelacanths

    GenusSpeciesLocalityFormationMaterialNotesImages
    Coccoderma[43] C. bavaricumKelheimAltmühltal Formation
    C. gigasKelheimAltmühltal Formation
    C. suevicum
    C. nudumKelheimAltmühltal Formation
    LibysL. polypterusKelheimAltmühltal Formation
    L. superbusZandtTorleite Formation
    UndinaU. acutidensKelheimAltmühltal Formation
    U. penicillataKelheim, Painten, WattendorfAltmühltal Formation, Torleite Formation
    MacropomaM. willemoesiEichstättAltmühltal Formation

    Reptiles

    Lizards

    NameSpeciesLocalityFormationMaterialNotesImages
    ArdeosaurusA. brevipesEichstättAltmühltal FormationSmall Lepidosaur thought to be close to the base of Geckota[44]
    A. digitatellusSolnhofen, EichstättAltmühltal Formation
    BavarisaurusB. macrodactylusKelheimAltmühltal Formation
    PalaeolacertaP. bavarica
    EichstaettisaurusE. schroederiEichstättAltmühltal Formation
    SchoenesmahlS. dyspepsiaKelheimAltmühltal FormationBavariasaurid lizard that was found in the stomach of the Compsognathus Holotype

    Rhynchocephalians

    GenusSpeciesLocalityFormationMaterialNotesImages
    HomoeosaurusH. maximillioniEichstätt, Painten, Schamhaupten, KelheimAltmühltal Formation, Torleite Formation
    SapheosaurusS. laticepsEichstätt, PaintenAltmühltal Formation, Torleite FormationLarge sapheosaurid reaching 70 cm in length, it may be semi-aquatic
    SphenofontisS. velseraegenBrunnTorleite Formation
    KallimodonK. pulchellusEichstättAltmühltal FormationSapheosauridae
    OenosaurusO. muehlheimensisMühlheimMörnsheim FormationRhynchocephalian with dentition unique among Tetrapods[45]
    VadasaurusV. herzogiPaintenTorleite FormationRhynchocephalian close to the base of Pleurosauridae[46]
    PiocormusP. laticepsPainten?Torleite Formation
    PleurosaurusP. goldfussiEichstätt, Langenaltheim, Painten, Mühlheim, WattendorfAltmühltal Formation, Torleite Formation, Mörnsheim FormationAcrosaurus fischmanni is a synonym.[47]

    Ichthyosaurs

    GenusSpeciesLocalityFormationMaterialNotesImages
    Aegirosaurus[48] A. leptospondylusEichstätt, Solnhofen[49] Altmühltal Formation
    A. sp[50] EichstättAltmühltal Formation
    IchthyosauriaIndeterminatePaintenTorleite Formation
    IchthyosauriaIndeterminatePaintenTorleite Formation
    "Ichthyosaurus""I." posthumusEichstätt, Kelheim, SolnhofenAltmühltal FormationNomen dubium. Known only from isolated teeth.
    Nannopterygius?N. spDaitingMörnsheim Formation

    Turtles

    GenusSpeciesLocalityFormationMaterialNotesImages
    Achelonia[51] A. formosaWattendorfTorleite Formation
    CyrturaC. temnospondylaSolnhofenAltmühltal Formation
    Eurysternum[52] E. wagleriSolnhofen, Kelheim, Painten, WattendorfAltmühltal Formation, Torleite Formation
    IdiochelysI. fitzingeriEichstättAltmühltal Formation
    PalaeomedusaP. testaKelheim, Eichstätt, SolnhofenAltmühltal FormationEurysternum crassipes and Thalassemys marina are synonyms.
    PlatychelysP. oberndorferiKelheimAltmühltal FormationPlatychelyid sea turtle
    Solnhofia[53] S. parsonsiEichstätt, PaintenAltmühltal Formation, Torleite Formation
    TropidemysT. langiiWattendorfTorleite Formation
    Thalassemys[54] T. bruntrutanaWattendorfTorleite Formation

    Crocodylomorphs

    GenusSpeciesLocalityFormationMaterialNotesImages
    Aeolodon[55] A. priscusDaitingMörnsheim FormationTwo partial skeletonsA teleosaur
    Alligatorellus[56] A. bavaricusEichstätt, Kelheim, PaintenAltmühltal Formation, Mörnsheim FormationPartial skeleton[57] An atoposaurid
    AtoposaurusA. oberndorferiEichstättAltmühltal FormationAn atoposaurid
    Cricosaurus[58] C. albersdoerferiPaintenTorleite FormationComplete skeleton preserving soft tissue
    C. bambergensisWattendorfTorleite FormationOne almost complete skeleton
    C. elegansEichstätt, Solnhofen, DaitingAltmühltal Formation, Mörnsheim FormationSkull and vertebrae (NHM 43005)[59]

    Partial skull (NHM 37006)

    A metriorhynchid
    C. rauhutiMühlheimMörnsheim FormationPartial 3D skull
    CrocodilaemusC. spBrunnTorleite FormationComplete skeleton of a juvenile
    Dakosaurus[60] D. maximusEichstätt, Schernfeld, Schamhaupten, Painten, KelheimAltmühltal Formation, Torleite FormationIsolated teeth (JME-SOS4577, JME-SOS2535)[61] A metriorhynchid
    GeosaurusG. giganteusSolnhofen, DaitingAltmühltal Formation, Mörnsheim FormationSkull and mandible (NHM 37020)

    Isolated tooth (NHM 37016)[62]

    A metriorhynchid
    G. grandisDaitingMörnsheim Formation
    RhacheosaurusR. gracilisEichstätt, Zandt, DaitingAltmühltal Formation, Mörnsheim Formation, Torleite FormationSkull and skeleton (NHM R.3948)A metriorhynchid
    MachimosaurusM. spMühlheimMörnsheim FormationIsolated teeth
    "Steneosaurus""S." jugleri

    Dinosaurs

    GenusSpeciesLocalityFormationMaterialNotesImage
    Archaeopteryx[63] A. albersdoerferi[64] DaitingMörnsheim FormationParts of the skull and wings
    A. siemensii[65] Langenaltheim, WorkerszellAltmühltal Formation
    A. lithographicaLangenaltheim, EichstättAltmühltal Formation
    Alcmonavis[66] A. poeschliMühlheimMörnsheim FormationPartial wing
    CompsognathusC. longipesKelheim (Goldberg)Altmühltal FormationComplete skeletonStomach contents are preserved in the form of the lizard Schoenesmahl.A second specimen of the genus is known from the French Canjuers Lagerstätte
    Juravenator[67] J. starkiSchamhauptenTorleite FormationSingle complete skeleton with soft tissue
    Ostromia[68] O. crassipesJachenhausenTorleite FormationPartial limb bones
    Sciurumimus[69] S. albersdoerferiPaintenTorleite FormationComplete skeleton of a juvenile preserving soft tissue

    Pterosaurs

    GenusSpeciesLocalityFormationMaterialNotesImages
    AerodactylusA. scolopacicepsFormerly classified as a species of Pterodactylus, closely related to Ardeadactylus, Aurorazhdarcho, and Cycnorhamphus.[70]
    AnurognathusA. ammoniThe holotype consists of a partial skeleton, while the second specimen is represented by a perfectly preserved juvenile including soft tissue.
    ArdeadactylusA. longicollumThe holotype was destroyed in a WW2 air raid, but another more complete specimen is known from the Nusplingen limestone.Formerly considered a species of Pterodactylus, it is more closely related to Gnathosaurus and Germanodactylus.[71]
    Altmuehlopterus[72] A. rhamphastinus
    Aurorazhdarcho[73] A. micronyx
    Bellubrunnus[74] B. rothgaengeriBrunnTorleite FormationSingle complete skeleton of a juvenile
    Balaenognathus[75] B. maeuseriWattendorfTorleite Formation
    CtenochasmaC. elegansOriginally classified as Pterodactylus elegans
    C. gracile
    C. spMühlheimMörnsheim FormationSkull and two cervical vertebrae
    CycnorhamphusC. suevicusOriginally classified as Pterodactylus suevicus.

    This genus originally hails from the Nusplingen Limestone

    DiopecephalusD. kochi
    GermanodactylusG. cristatusOriginally classified as Pterodactylus cristatus
    GnathosaurusG. subulatus
    GnathosaurinaeIndeterminateBrunnTorleite Formation
    PterodactylusP. antiquusOnly one species is currently supported as valid, the rest represent growth stages, often of other species, synonyms, or nomina dubia.[76]
    Petrodactyle[77] P. wellnhoferiMühlheimMörnsheim Formation
    Propterodactylus[78] P. frankerlaeUlmensePainten Formation[79] A complete skeletonNotable for its transitional features between pterodactyloids and more basal monofenestratans
    Rhamphorhynchus[80] R. muensteriEichstätt, Solnhofen, Painten, KelheimAltmühltal Formation, Torleite FormationOnly one species is currently supported as valid, the rest represent growth stages, often of other species, synonyms, or nomina dubia.
    "Rhamphodactylus"IndeterminateMühlheimMörnsheim Formation
    Scaphognathus[81] S. crassirostrisEichstätt, MühlheimAltmühltal Formation, Mörnsheim FormationA Solnhofen genus erected in 1861 by J. A. Wagner when he recognized the rhamphorhynchoid nature of "Pterodactylus" crassirostris after the discovery of a specimen with a preserved tail. The Solnhofen Limestone is the only known source of Scaphognathus fossils.

    S. crassirostris was originally named P. crassirostris in 1831 by August Goldfuss who mistook the tailless specimen for a new Pterodactylus species.

    Sauropterygia

    !Genus!Species!Locality!Formation!Material!Notes!Images
    Pliosauridae[82] IndeterminateKelheim, PaintenAltmühltal Formation, Torleite FormationIsolated teeth of great size

    Echinodermata

    Crinoids

    GenusSpeciesLocalityFormationNotesImages
    Liliocrinus[83] L. sp
    MillericrinusM. mespiliformis
    M. nobilis
    BalanocrinusB. sigmaringensis
    B. subteres
    PentacrinusP. cf. pentagonalis
    ComaturellaC. pennataSolnhofen, Zandt, Langenaltheimer HaardtAlmtühltal Formation, Painten Formation
    C. formosaZandtPainten Formation
    SolanocrinitesS. costatus
    S. gracilis
    S. imperialis
    Saccocoma[84] S. pectinataSaccocoma remains are the most common macroscopic fossils in the Solnhofen limestone.
    S. tenellaEichstätt, Solnhofen, BlumenbergAltmühltal Formation

    Ophiuroidea

    !Genus!Species!Locality!Formation!Notes!Images
    GeocomaG. carinataSolnhofen, ZandtAltmühltal Formation, Painten Formation
    G. planata
    OphiopetraO. lithographicaKelheim, HienheimPainten Formation
    OphiurellaO. speciosaSolnhofen, PaintenAltmühltal Formation, Painten Formation
    SinosuraS. kelheimenseHienheim, Ried, KelheimPainten Formation

    Asteroidea

    !Genus!Species!Locality!Formation!Notes!Images
    ArchasteropectenA. elegans
    LithasterL. jurassicus
    PentasteriaP. tithonicaHienheimPainten Formation
    P. lithographicaZandtPainten Formation
    RiedasterR. reicheliRiedPainten Formation
    TerminasterT. cancriformisBöhmfeldAltmühltal Formation

    Echinoidea

    Cidaroidea

    !Genus!Species!Locality!Formation!Notes!Images
    DiplocidarisD. spEichstättAltmühltal Formation
    RhabdocidarisR. boehmi
    R. mayri
    R. orbignyanaBrunn, PaintenTorleite Formation, Painten Formation
    R. trispinata
    R. spPaintenPainten Formation
    PlegiocidarisP. blumenbachii
    P. sp
    ParacidarisP. sp

    Euechinoidea

    !Genus!Species!Locality!Formation!Notes!Images
    DiademataceaIndeterminate
    EchinothurioideaIndeterminate
    PolydiademaP. sp
    DiademopsisD. sp
    HemipedinaH. sp
    PedinaP. lithographica
    P. spPaintenPainten Formation
    PhymopedinaP. marchamensisPaintenPainten Formation
    P. sp
    HemicidarisH. intermediaEichstätt, Zandt, PaintenAltmühltal Formation, Painten Formation
    AcrocidarisA. sp
    DiplopodiaD. subangularis
    PseudodiademaP. lithographicaRied, BreitenhillPainten Formation
    P. pseudodiadema
    TetragrammaT. planissimaPaintenPainten Formation

    Other Echinoids

    !Genus!Species!Locality!Formation!Notes!Images
    PhymosomaP. spMörnsheim, BreitenhillMörnsheim Formation, Painten Formation
    PseudosaleniaP. asperaBrunn, SchamhauptenPainten Formation, Torleite Formation
    PygasterP. sp
    HolectypusH. corallinus
    H. sp
    KieripygurusK. jurensis
    PygurusP. spEichstätt, Kelheim, Schamhaupten, PaintenAltmühltal Formation, Painten Formation
    NucleolitesN. spBrunn, KelheimPainten Formation, Torleite Formation
    CollyropsisC. carinata
    C. sp
    StomechinusS. perlatusKapfelbergAltmühltal Formation?

    Holothuroidea

    !Genus!Species!Locality!Formation!Notes!Images
    JumarainaJ. sp
    AchistrumA. sp
    HemisphaerantosH. sp
    PriscopedatusP. sp
    SynallactidaeIndeterminate

    Molluscs

    Cephalopods

    Nautiloidea

    !Genus!Species!Locality!Formation!Notes!Images
    PseudaganidesP. franconicusSolnhofen?Altmühltal Formation
    P. spMühlheimMörnsheim Formation
    PseudonautilusP. spMühlheimMörnsheim Formation

    Ammonoidea

    !Genus!Species!Locality!Formation!Notes!Images
    PhyllocerasP. serumDaitingMörnsheim Formation
    GlochicerasG. cf. lens
    StreblitesS. spEichstätt, BlumenbergAltmühltal Formation
    FontannesiellaF. prolithographicaSolnhofen, Daiting, MörnsheimAltmühltal Formation, Mörnsheim Formation
    F. cf. franciscanumEichstättAltmühltal Formation
    F. cf. euglyptumLangenaltheimer HaardtAltmühltal Formation
    LingulaticerasL. planulatum
    L. solenoidesWegscheidAltmühltal Formation
    L. sp
    MetahaplocerasM. acallopistum
    M. rebouletianum
    M. subnudatum
    NeochetocerasN. bousBlumenberg, BirkhofAltmühltal Formation
    N. mucronatum
    N. praecursor
    N. steraspisSolnhofen, Eichstätt, HummelbergAtlmühltal Formation, Mörnsheim Formation
    ParalingulaticerasP. haeberleini
    P. lithographicumSolnhofenAltmühltal Formation
    P. nodosum
    P. percevali
    TaramellicerasT. hemipleurum
    AspidocerasA. hoplisumSolnhofen, WegscheidAtlmühltal Formation
    A .spBrunnTorleite Formation
    PhysodocerasP. episoides
    P. hermanniBrunnTorleite Formation
    P. nattheimense
    P. pipini
    SutneriaS. aporaBirkhof, WintershofAltmühltal Formation
    S. bracheri
    S. rebholzi
    S. subeumelaBrunnTorleite Formation
    S. sp
    HybonotellaH. mundulaLangenaltheimer HaardtAltmühltal Formation
    HybonoticerasH. hybonotumSolnhofen, BlumenbergAltmühltal Formation
    H. pseudohybonotum
    H. sp
    GravesiaG. gigasHienheim, Ried, WegscheidPainten Formation
    G. sp
    BerckhemeriaB. cf. scherzingeri
    VirgataxiocerasV. setatum
    V. supinum
    V. sp
    EuvirgalithacocerasE. supremumSolnhofen, EichstättAtlmühltal Formation
    E. eystettenseEichstättAltmühltal Formation
    HoelderiaH. spWintershofAltmühltal Formation
    LithacocerasL. eigeltingenseZandtPainten Formation
    L. eystettense
    L. cf. liptingenseLangenaltheimer HaardtAltmühltal Formation
    L. cf. riedense
    L. cf. ulmense
    L. spSchernfeld, EichstättAltmühltal Formation
    PseudodiscosphinctesP. ardescicus
    SilisphinctesS. cf. russiZandtPainten Formation
    S. spWegscheidAltmühltal Formation
    SubplanitesS. moernsheimensisDaitingMörnsheim Formation
    S. postrueppellianusSolnhofen, Blumenberg, Wintershof, EichstättAltmühltal Formation
    S. rueppellianusSolnhofen, WegscheidAltmühltal Formation
    S. viohliEichstättAltmühltal Formation
    S. laisackerensisBirkhofAltmühltal Formation
    "Torquatisphinctes""S". filiplex
    "S". isolatus
    AulacostephanusA. cf. subundoraePaintenTorleite Formation?, Painten Formation

    Belemnoidea

    !Genus!Species!Locality!Formation!Notes!Images
    AcanthoteuthisA. leichi
    A. speciosaSolnhofen, Eichstätt, BirkhofAltmühltal Formation
    BelemnoteuthisB. mayriWintershofAltmühltal Formation
    HibolithesH. hasatusSolnhofen, Breitenhill, PaintenAltmühltal Formation, Painten Formation, Torleite Formation
    H. semisulcatus
    PavloteuthisP. kapitzkeiWegscheidAltmühltal Formation
    RhaphibelusR. aciculaEichstätt. Blumenberg, WegscheidAltmühltal Formation
    WinkleriteuthisW. problematicusSchernfeldAltmühltal Formation

    Vampyromorpha

    !Genus!Species!Locality!Formation!Notes!Images
    BoreopeltisB. sagittataSolnhofenAltmühltal Formation
    Engeseriteuthis[85] E. arcuatusSchamhauptenPainten Formation
    PlesioteuthisP. priscaEichstätt, Solnhofen, Wegscheid, SchamhauptenAltmühltal Formation, Painten Formation
    P. subovata
    SenefelderiteuthisS. tricarinataEichstätt, DaitingAltmühltal Formation, Mörnsheim Formation
    Trachyteuthis[86] T. hastiformisDaiting, Painten, Eichstätt, BrunnMörnsheim Formation, Altmühltal Formation, Painten Formation, Torleite Formation
    T. teudopsiformisBlumenberg, SchamhauptenAltmühltal Formation, Painten Formation
    Teudopsinia[87] T. grossheideiEichstättAltmühltal Formation
    TyrionellaT. fauseriSchernfeldAltmühltal Formation
    DonovaniteuthisD. schoepfeli
    LeptoteuthisL. gigasSolnhofen, Eichstätt, BlumenbergAltmühltal Formation
    ListroteuthisL. conicaDaitingMörnsheim FormationHolotype specimen destroyed during WW2
    BavaripeltisB. bavaricaSolnhofenAltmühltal Formation
    DoryanthesD. munsteriiEichstätt, Blumenberg, MörnsheimMörnsheim Formation, Altmühltal Formation
    Palaeololigo[88] P. albersdoerferiEichstättAltmühltal Formation
    P. oblongaSchernfeldAltmühltal Formation
    CelaenoteuthisC. incertaSolnhofen, Eichstätt, BirkscheidAltmühltal Formation
    MuensterellaM. scutellarisEichstätt, Solnhofen, Wegscheid, Wintershof, Painten, MörnsheimMörnsheim Formation, Altmühltal Formation, Painten Formation
    M. spinosaDaitingMörnsheim Formation

    Arthropods

    Crustacea

    Crustaceans of the Solnhofen Formation
    NameSpeciesPresenceNotesImages
    Cancrinos[89] C. clavigerSolnhofenA cancrinid
    CycleryonC. elongatusSolnhofenAn eryonid
    C. orbiculatusAn eryonid
    C. propinquusAn eryonid
    ErymaE. modestiformeAn erymid
    E. veltheimiiAn erymid
    EryonE. cuvieri[90] An eryonid
    EtalloniaE. longimanaSolnhofenAn axiid
    GigantosculdaG. ehrlichfeckei[91] EichstättA mantis shrimp
    GlypheaG. pseudoscyllarusA glypheid
    G. tenuisEichstättA glypheid
    KnebeliaK. bilobataSolnhofenAn eryonid
    MagilaM. desmarestiiSolnhofenAn axiid
    M. latimanaEichstättAn axiid
    MecochirusM. dubiusA mecochirid
    M. longimanatusA mecochirid
    OrhomalusO. deformisSolnhofenA pagurid
    PalaeastacusP. fuciformisAn erymid
    Palaeobresilia[92] P. kurthetriegeriA bresiliid shrimp
    PalaeopentachelesP. roettenbacheriA polychelid
    PalinurinaP. longipesA spiny lobster
    P. teneraEichstättA spiny lobster
    PustulinaP. minutaAn erymid
    PseudastacusP. pustulosusA stenochirid
    RosenfeldiaR. oppeliEichstättAn eryonid
    SculdaS. pennata[93] A mantis shrimp
    S. spinosaA mantis shrimp
    Spinosculda[94] S. ehrlichiA mantis shrimp
    StenochirusS. angustusFlonheimA stenochirid
    S. mayeriSolnhofenA stenochirid
    SquamosoglypheaS. squamosaKelheimA glypheid
    Tyrannosculda[95] T. lauraeEichstättA mantis shrimp

    Thylacocephalans

    Thylacocephalans of the Solnhofen Formation
    TaxaPresenceNotesImages
    Clausocaris C. lithographica
    DollocarisD. michelorum
    MayrocarisM. bucculata
    FalcatacarisF. bastelbergeri

    Insects

    References

    General

    • Lambers, P. H. (1999). The actinopterygian fish fauna of the Late Kimmeridgian and Early Tithonian 'Plattenkalke' near Solnhofen (Bavaria, Germany): state of the art. Geologie en Mijnbouw 78:215-229.

    ]