Okenia Explained

Okenia is a genus of colorful sea slugs, specifically of dorid nudibranchs, marine gastropod mollusks in the family Goniodorididae.

Species

Species within the genus Okenia include:

• Okenia academica Camacho-Garcia & Gosliner, 2004
• Okenia africana Edmunds, 2009^[1]
• Okenia ameliae Ortea, Moro & Caballer, 2014
• Okenia amoenula (Bergh, 1907)^[2]
• Okenia angelensis Lance, 1966
• Okenia angelica Gosliner & Bertsch, 2004
• Okenia ascidicola M.P. Morse, 1972
• Okenia aspersa (Alder & Hancock, 1845)
• Okenia atkinsonorum Rudman, 2007
• Okenia aurorapapillata Paz-Sedano & Pola in Paz-Sedano, Wilson, Carmona, Gosliner & Pola, 2021
• Okenia barnardi Baba, 1937
• Okenia brunneomaculata Gosliner, 2004^[3]
• Okenia cochimi Gosliner & Bertsch, 2004
• Okenia cupella (Vogel & Schultz, 1970)
• Okenia digitata (Edmunds, 1966)
• Okenia distincta Baba, 1940
• Okenia echinata Baba, 1949
• Okenia elegans (Leuckart, 1828) - type species^[3]
• Okenia elisae Paz-Sedano & Pola in Paz-Sedano, Wilson, Carmona, Gosliner & Pola, 2021
• Okenia eolida (Quoy & Gaimard, 1832)
• Okenia evelinae Er. Marcus, 1957
• Okenia felis Gosliner, 2010^[4]
• Okenia ghanensis Edmunds, 2009
• Okenia hallucigenia Rudman, 2004
• Okenia harastii Pola, Roldán & Padilla, 2014
• Okenia hiroi (Baba, 1938)
• Okenia hispanica Valdes & Ortea, 1995
• Okenia impexa Er. Marcus, 1957
• Okenia japonica Baba, 1949^[3]
• Okenia kendi Gosliner, 2004^[3]
• Okenia kondoi (Hamatani, 2001)^[3] - originally described as Sakishimaia kondoi Hamatani, 2001
• Okenia lambat Gosliner, 2004^[3]
• Okenia leachii (Alder & Hancock, 1854)
• Okenia liklik Gosliner, 2004^[3]
• Okenia longiductis Pola, Paz-Sedano, Macali, Minchin, Marchini, Vitale, Licchelli & Crocetta, 2019
• Okenia luna Millen, Schrödl, Vargas, & Indacochea, 1994
• Okenia mediterranea (Ihering, 1886)
• Okenia mellita Rudman, 2004
• Okenia mexicorum Gosliner & Bertsch, 2004
• Okenia mica Ortea & Moro, 2014
• Okenia mija Burn, 1967
• Okenia miramarae Ortea & Espinosa, 2000
• Okenia nakamotoensis (Hamatani, 2001)^[3]
• Okenia nakanoae Paz-Sedano & Pola in Paz-Sedano, Wilson, Carmona, Gosliner & Pola, 2021
• Okenia opuntia Baba, 1960
• Okenia pellucida Burn, 1967^[3]
• Okenia picoensis Paz-Sedano, Ortigosa & Pola, 2017
• Okenia pilosa (Bouchet & Ortea, 1983)^[3]
• Okenia plana Baba, 1960^[3]
• Okenia polycerelloides (Ortea & Bouchet, 1983)
• Okenia problematica Pola, Paz-Sedano, Macali, Minchin, Marchini, Vitale, Licchelli & Crocetta, 2019
• Okenia pulchella Alder & Hancock, 1854
• Okenia purpurata Rudman, 2004
• Okenia purpureolineata Gosliner, 2004^[3]
• Okenia rhinorma Rudman, 2007
• Okenia rosacea (MacFarland, 1905)^[3]
• Okenia sapelona Ev. Marcus & Er. Marcus, 1967
• Okenia siderata Paz-Sedano & Pola in Paz-Sedano, Wilson, Carmona, Gosliner & Pola, 2021
• Okenia stellata Rudman, 2004
• Okenia tenuifibrata Paz-Sedano & Pola in Paz-Sedano, Wilson, Carmona, Gosliner & Pola, 2021
• Okenia vancouverensis (O'Donoghue, 1921)
• Okenia vena Rudman, 2004
• Okenia virginiae Gosliner, 2004^[3]
• Okenia zoobotryon (Smallwood, 1910)

Nomen dubium:

• Okenia pusilla Sordi, 1974

Species names considered to be synonyms:

• Okenia dautzenbergi Vayssière, 1919: synonym of Okenia aspersa (Alder & Hancock, 1845)
• Okenia modesta (A. E. Verrill, 1875): synonym of Okenia aspersa (Alder & Hancock, 1845)
• Okenia quadricornis (Montagu, 1815): synonym of Okenia aspersa (Alder & Hancock, 1845)

References

• Leuckart F. S. (1828). Breves animalium quorundum maxima ex parte marinorum descriptiones. Heidelberg pp. 24:
• Bergh, L. S. R. 1881. Ueber die Gattung Idalia, Leuckart. Archiv für Naturgeschichte 47(1):140-181, pls. 6-8
• Odhner N.J. 1941. New polycerid nudibranchiate mollusca and remarks on this family. Göteborgs Kungl. Vetenskaps- och Vitterhets-Samhälles Handlingar, Ser. B, Matematiska och Naturvetenskapliga Skrifter, 1(11): 1-20
• Edmunds, M. (1966). Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana. Journal of the malacological Society of London. 37: 69-72
• Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca. in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels. 50: pp. 180–213.
• Hamatani I. (2001) Two new species of Goniodorididae (Opisthobranchia; Nudibranchia) with a new genus from Kuroshima Island, Okinawa, Japan. Venus, 60(3): 151-156

Further reading

• Behrens David W., 1980, ‘’Pacific Coast Nudibranchs: a guide to the opisthobranchs of the northeastern Pacific’’, Sea Challenger Books, California
• http://www.catalogueoflife.org accessed 25 August 2009

Notes and References

1. Edmunds M. (2009) "Opisthobranchiate Mollusca from Ghana: Goniodorididae". Journal of Conchology 40(1): 37-54. abstract .
2. Gosliner T.M. 1987. Nudibranchs of Southern Africa
3. Gosliner T. M. (2004). "Phylogenetic Systematics of Okenia, Sakishmaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific". Proceedings of the California Academy of Sciences 55(5): 125-161. PDF.
4. Gosliner T. M. (2010). "Two New Species of Nudibranch Mollusks From the Coast of California". Proceedings of the California Academy of Sciences 66: 623-631. PDF