Ntawere Formation Explained

Ntawere Formation
Coordinates:-10.8°N 33.1°W
Paleocoordinates:53.6°N 20.6°W
Extent:Luangwa Basin
Overlies:Escarpment Grit
Underlies:Red Marls
Subunits:Lower Ntawere, Upper Ntawere
Region:Eastern Province
Otherlithology:Sandstone, marl
Prilithology:Mudstone
Period:Anisian
Age:Anisian
~
Type:Geological formation

The Ntawere Formation is a Middle Triassic (Anisian) geological formation in Zambia, preserving fossils of synapsids, archosaurs, and temnospondyls.[1]

Geology

Several different facies are present in the Ntawere Formation, reconstructing a floodplain environment. The coarsest facies are trough cross-bedded conglomeratic sandstone full of mineral concretions. These coarse deposits formed in ancient channels such as riverbeds. Another type of facies involves thick beds of mudstone interbedding with thinner layers of fine-grained sandstone, indicating alternating low- and high-energy water flow within the channels. Graded sandstone to mudstone overbank deposits (complete with ripple marks) occur near channel deposits. Extensive successions of laminated or massive mudstone are common, often containing slickensides, calcareous nodules or layers (marls), and/or hematite nodules. These types of thick mud/marl layers likely formed in more quiet parts of the floodplain isolated from turbulent channels.

The floodplain was seemingly more active during the deposition of the Lower Ntaware Formation, as coarser sandstone channel fills are prevalent in that section while extensive mudstone layers are more common in the Upper Ntaware Formation. Fossils are typically found preserved in calcareous nodules in the mudstone of the Upper Ntaware, although they occur in some parts of the Lower Ntaware as well. The Ntaware Formation is an example of red beds, which are typically deposited in warm environments with seasonal ponds and rivers tied to wet and dry seasons.

Paleobiota

Fish

Fish of the Ntawere Formation
TaxonMemberMaterialNotes
Hybodontoidei indet.Upper NtawareFin spinesAn indeterminate non-hybodontid hybodontoid shark, similar to Lissodus and Lonchidion
Ptycoceratodontidae indet.Lower Ntaware, Upper NtawereTooth platesAn indeterminate lungfish, similar to Ptychoceratodus

Amphibians

Amphibians of the Ntawere Formation
TaxonMemberMaterialNotesImages
Batrachosuchus concordiUpper NtawareA skull and forelimbsA brachyopid stereospondyl
Cherninia megarhinaUpper NtawareA partial skullA mastodonsaurid stereospondyl
"Stanocephalosaurus" pronusUpper NtawareA skullA new species of capitosauroid, likely not part of the genus Stanocephalosaurus
Stereospondyli indet.Upper NtawareVertebrae and part of a jawIndeterminate stereospondyl material. The jaw fragment may belong to a rhinesuchid, lydekkerinid, or rhytidosteid

Synapsids

Synapsids of the Ntawere Formation
TaxonMemberMaterialNotesImages
Cricodon metabolusUpper NtawareA partial skeletonA trirachodontid cynodont
Cynognathus crateronotusLower NtawareTeeth, vertebrae, partial humerus, iliumA cynognathian cynodont
Diademodon tetragonusLower NtawareA skullA diademodontid cynodont
"Kannemeyeria" latirostrisLower NtawareA skullA kannemeyeriiform dicynodont, possibly a species of Kannemeyeria or Dolichuranus
Kannemeyeria lophorhinusLower NtawareA skullA kannemeyeriid dicynodont
Luangwa drysdalliUpper NtawareSkulls and postcranial materialA traversodontid cynodont
Sangusaurus edentatusUpper NtawareA partial skullA stahleckeriid dicynodont
Traversodontidae sp.Upper NtawareSkulls and postcranial materialAn unnamed species of gomphodont-like traversodontid cynodont
Zambiasaurus submersusUpper NtawareA skull fragmentA stahleckeriid dicynodont

Archosauromorphs

Archosauromorphs of the Ntawere Formation
TaxonMemberMaterialNotesImages
Archosauriformes indet.Upper NtawareTeethVarious indeterminate large carnivorous archosauriforms, likely loricatans
Archosauromorpha indet.Upper NtawareA neck vertebraAn indeterminate archosauromorph, possibly an allokotosaur
Dinosauriformes indet.Upper NtawareA partial fibulaA large dracohortian, likely a silesaurid
Loricata indet.Upper NtawareVertebraeAn indeterminate large archosaur, similar to Stagonosuchus
Lutungutali sitwensisUpper NtawareAn iliumA basal silesaurid
Shuvosauridae indet.Upper NtawarePart of a femurAn indeterminate shuvosaurid archosaur similar to Shuvosaurus and the indeterminate "Moenkopi poposauroid"
Silesauridae indet.Upper NtawareA maxilla, femurs, and material from the hip, ankle, and footIndeterminate basal silesaurid material comparable to Asilisaurus and Lutungutali

See also

Further reading

Notes and References

  1. Peecook. Brandon R.. Steyer. J. Sébastien. Tabor. Neil J.. Smith. Roger M. H.. 2017-11-29. Updated geology and vertebrate paleontology of the Triassic Ntawere Formation of northeastern Zambia, with special emphasis on the archosauromorphs. Journal of Vertebrate Paleontology. 37. sup1. 8–38. 10.1080/02724634.2017.1410484. 2017JVPal..37S...8P . 133878741. 0272-4634.