Nemastomatinae Explained

Nemastomatinae is a subfamily of harvestmen with over 125 described species in 19 (extant) genera. They are usually found in temperate regions.

Species

For complete breakdown, see Nemastomatidae, or:

See main article: List of Dyspnoi species.

The genera are as follows (as of 2023):

• Acromitostoma Roewer, 1951 (2 species)Spain, Morocco
• Burnia Prieto, 2021 (2 species)Spain
• Carinostoma Kratochvíl, 1958 (2 species) Southern Europe
• Caucnemastoma Martens, 2006 (2 species) - Russia
• Centetostoma Kratochvíl, 1958 (2 species)Western Europe
• Giljarovia Kratochvíl, 1958 (11 species) Eastern Europe, Russia, etc. (See Tchemeris & Kovblyuk, 2012)^[1]
• Hadzinia Šilhavý, 1966 (See Novak & Kozel, 2014)^[2] (2 species)European Balkans
• Histricostoma Kratochvíl, 1958 (8 species)Southern Europe
• Mediostoma Kratochvíl, 1958Southeastern Europe, Anatolia, Central Asia^[3]
  • Mitostoma Roewer, 1951Europe (widespread), Caucasus Nemaspela Šilhavý, 1966Caucasus, Southeastern Europe^{[4] [5]}
• Nemastoma C.L. Koch, 1836Europe (Widespread), Algeria (?)
• Nemastomella Mello-Leitão, 1936Western Europe, Algeria (?)
• Paranemastoma Redikorzev, 1936Europe (Widespread), Caucasus, Anatolia, (plus India?)
• Pyza Staręga, 1976Southeastern Europe, Anatolia
• Saccarella Schönhofer & Martens, 2012Italy
• Sinostoma Martens, 2016China^[6]
• Starengovia Snegovaya, 2010Central Asia, China^{[7] [8]}
• Vestiferum Martens, 2006Caucasus, Anatolia

Fossil species

There are currently 5 described fossil harvestmen that have been assigned to Nemastomatinae, 4 of which are assigned to modern genera:^[9]

• Histricostoma Kratochvíl, 1958

• † Histricostoma tuberculatum (C.L. Koch & Berendt, 1854) - Paleogene Baltic & Bitterfield amber
• Mitostoma Roewer, 1951

• † Mitostoma denticulatum (C.L. Koch & Berendt, 1854) - Paleogene Baltic amber

• † Mitostoma gruberi Dunlop & Mitov, 2009 - Paleogene Baltic & Bitterfield amber
• Nemastoma (C.L. Koch, 1836)

• † Nemastoma incertum C.L. Koch & Berendt, 1854 - Paleogene Baltic amber
• † Paragilarovia Elsaka, Mitov, & Dunlop, 2019^[10]

• † Paragilarovia hochae Elsaka, Mitov, & Dunlop, 2019 - Paleogene Baltic amber

Notes and References

1. Web site: A new species of the genus Giljarovia Kratochvíl, 1959 (Arachnida:Opiliones: Phalangiidae) from the Crimea. ResearchGate. en. 2019-05-23.
2. Web site: Hadzinia ferrani, sp. n. (Opiliones: Nemastomatidae), a highly specialized troglobiotic harvestman from Slovenia. ResearchGate. en. 2019-05-23.
3. Snegovaya. N.. Kurt. K.. Yağmur. E.A.. 2016. Mediostoma izmirica sp. nov.: A new harvestman species (Arachnida, Opiliones, Nemastomatidae) from Turkeyjournal=Turkish Journal of Zoology. 40. 4. 489-498. 10.3906/zoo-1412-10. 1300-0179. en. free. 2023-01-04.
4. Two new harvestman species (Arachnida: Opiliones) from the collection of Siberian Zoological Museum . ResearchGate. 10.15298/arthsel.22.1.03 . 2013 . Tchemeris . A. N. . Arthropoda Selecta . 22 . 41046–0 . free .
5. Nemaspela ladae sp. N., a new troglobitic nemastomatid (Opiliones, Dyspnoi, Nemastomatidae) from a Dinaric cave. . ResearchGate. 10.11646/zootaxa.3694.3.4 . 2013 . Karaman . IVO M. . Zootaxa . 3694 . 3 . 240–248 . 26312286 .
6. Sinostoma yunnanicum, the first nemastomatine harvestman in China (Arachnida: Opiliones: Nemastomatidae). . 10.11646/zootaxa.4126.3.9 . ResearchGate. 2016 . Martens . Jochen . Zootaxa . 4126 . 3 . 444–450 . 27395599 .
7. Martens . Jochen. 2017. On Starengovia Snegovaya, a genus of Asian nemastomatines (Arachnida: Opiliones: Nemastomatidae). Revue Suisse de Zoologie. 124 . 2. 187–201. ResearchGate. 10.5281/zenodo.893462. 52200517.
8. Zhang. Chao. Martens. Jochen. 2018. Ancient home or in exile? The easternmost species of genus Starengovia Snegovaya, 2010 found in China (Opiliones, Nemastomatidae, Nemastomatinae). ZooKeys. 770. 105–115. 10.3897/zookeys.770.25491. 1313-2970. 6041357. 30002591. free.
9. Web site: A summary list of fossil spiders and their relatives. wsc.nmbe.ch. en. 2019-05-23.
10. Web site: New fossil harvestmen (Arachnida: Opiliones) in the HOFFEINS ambe...: Ingenta Connect . www.ingentaconnect.com . dead . https://web.archive.org/web/20200917202937/https://www.ingentaconnect.com/content/schweiz/njbgeol/2019/00000292/00000002/art00003 . 2020-09-17.