Mollia (plant) explained

Mollia is a genus of flowering plants belonging to the family Malvaceae.[1] It is within the Grewioideae subfamily,[2] and the Grewieae tribe.[3]

Description

Mollia are large trees,[4] or tall shrubs,[5] growing up to tall.[4]

They have 3-veined from the base leaves which have serrated, toothed (dentate) or smooth margins.[4] [5] The indumentum (surface covering) has stellate hairs and pelate scales. They have rudimentary (or small) stipules (a small appendage at the bases of leaves),[5] which are caducous (fall off early).[4]

It has inflorescences in few- to many flowered units, sometimes on a common peduncle (stalk), or solitary flowers. They are showy and white. The sepals are narrow with the petals being truncate and glandless. The stamens are numerous, in 2 whorls each of 5 phalanges with basally fused filaments (stamen stalks), the outer whorl is antesepalous and longer than the inner.[5]

The anthers are introrse (with opening toward the centre of flower), subasifixed, diamorphic (having two different forms), in outer whorl they are cordate at base and in the inner whorl, they are sagittate (arrow-head shaped).[5]

The ovary is 2 locular, or incompletely so. The ovules are 2 seriate (arranged in rows) in each locule and numerous. The style is filiform (needle-like) and the stigma is punctiform (dot-like).[5] The fruit capsule is 2 locular (or bivalved),[4] [5] brown when mature and 1cm (00inches) in diameter.[4] The capsule is also loculicidal (splits along an edge), globose or compressed contrary to the septum and sometimes winged along the line of dehiscence (splitting edge). The loculi, each with two rows of 8 - 20 flattened seeds and fitting between narrow membranous partitions.[5]

The seeds are short winged,[5] various per fruit, plants pubescent.[4] The embryo (inside the seed) is flat, endosperm copious and oily.[5]

Taxonomy

The genus name of Mollia is in honour of Karl von Moll (1760–1838), an Austrian naturalist and statesman.[6] The genus was first described and published in Nov. Gen. Sp. Pl. Bras. Vol.1 on page 96 in 1826.

The genus is recognized by the United States Department of Agriculture and the Agricultural Research Service, but they do not list any known species.[7]

In the 1800s the genus was once placed within the family Tiliaceae,[8] It was returned in the Malvales family in 1935.[9]

It was placed within Grewioidaee subfamily due to molecular data (phospholipase gene) Blatter (unpulished source), but another author, Meijer thought it to be related to the Trichospermum genus due to similar capsule shapes.[5] [10]

The genus has been studied in 2021 and the various synonyms have been determined.[11]

Range and habitat

Its native range is southern Tropical America,[4] [12] and it is found in Bolivia, (northern, north-eastern and west central) Brazil, Colombia, Ecuador, Guyana,[5] Peru and Venezuela.

It is found in the savannah forests of Cerrado, Brazil.[13]

Known species

about 18 species known,[5] According to Kew:

Mollia glabrescens and Mollia nitida are listed by IUCN as 'Vulnerable'.[14]

Notes and References

  1. Web site: Mollia Mart. Plants of the World Online Kew Science . Plants of the World Online . 7 October 2021 . en.
  2. Mambrín . Mónica V. . Avanza . María M. . Ferrucci . María S. . Morphological and morphometrical pollen analy- sis of Corchorus, Heliocarpus, Luehea, Mollia and Triumfetta (Malvaceae, Grewioideae) in southern South America . Darwiniana . Nueva Serie . 2010 . 48 . 1.
  3. Muellner-Riehl . Alexandra . Brunken . Ulrike . A new tribal classification of Grewioideae (Malvaceae) based on morphology and molecular phylogenetic evidence . Systematic Botany . July 2012 . 37 . 3.
  4. Fernando Cornejo and John Janovec
  5. Klaus Kubitzki and Clemens Bayer (editors)
  6. Book: Burkhardt, Lotte . Verzeichnis eponymischer Pflanzennamen – Erweiterte Edition . Index of Eponymic Plant Names – Extended Edition . Botanic Garden and Botanical Museum, Freie Universität Berlin . 2018 . 978-3-946292-26-5 . pdf . German . Berlin . 10.3372/epolist2018 . 1 January 2021.
  7. Web site: Genus Mollia Mart. . npgsweb.ars-grin.gov . 27 November 2021.
  8. [John Lindley]
  9. Edlin . L. . A Critical Revision of Certain Taxonomic Groups of the Malvales. Part II H. . The New Phytologist . 15 May 1935 . 34 . 2 . 122–143.
  10. Meijer, Mem. N.Y. Bot Gard. 29, 68-82, 1978
  11. Costa . Maria Tereza Rodrigues . Bovini . Massimo G. . Guimarães . Elsie Franklin . Nomenclatural synopsis and typifications in Mollia (Malvaceae: Grewioideae) . Brittonia . September 2021 . 10.1007/s12228-021-09681-w.
  12. M.M. Grandtner and Julien Chevrette
  13. Maciel . Everton A. . Oliveira-Filho . Ary T. . Eisenloh . Pedro V. . Prioritizing rare tree species of the Cerrado-Amazon ecotone: warnings and insights emerging from a comprehensive transitional zone of South America Author links open overlay panel . Natureza & Conservação . July–December 2016 . 14 . 2 . 74–82.
  14. Web site: The IUCN Red List of Threatened Species . IUCN Red List of Threatened Species . 6 December 2021.