List of ornithopod type specimens explained

This list of specimens is a comprehensive catalogue of all the type specimens and their scientific designations for each of the genera and species that are included in the clade ornithopoda.Ornithopoda is a clade of ornithischian dinosaurs that includes some of the most common and widespread Mesozoic animals including iguanodonts, hadrosaurs, and some animals formerly called "hypsilophodonts". The clade was named by Othniel Charles Marsh in 1881^[1] and roughly means "bird feet". This name is reflective of the tridactyl feet of most ornithopods, which are superficially similar to many birds. Ornithopods were among the first dinosaurs known to scientists. The first species to be described was Iguanodon, which was described in 1825 by Sir Richard Owen.^[2] Today, they collectively comprise the most diverse ornithischian group.

The exact origin of ornithopods is uncertain. Some authors consider very primitive ornithischians like Nanosaurus, Hypsilophodon, and Jeholosaurus to be ornithopods,^[3] while others do not.^[4] Most ornithischians were ancestrally small and bipedal, which makes the taxonomy of these species complex and uncertain.^{[5] [6]} However, they are generally agreed to have originated by the middle Jurassic and shortly thereafter, they proliferated to most of the world's continents.^[7] Ornithischians in general, but especially ornithopods, saw a large degree of diversification during the early Cretaceous.^[8] This culminated in the proliferation of the giant hadrosaurs across the Northern Hemisphere and even into Africa and South America.^[9] Ornithopod remains are known from all of the world's continents, including Antarctica.^[10]

Scope and Terminology

This list will include the type fossils of each ornithopod species. In paleontology, a type specimen is one which is definitionally a member of a biological taxon. Additional specimens can only be "referred" to these taxa if an expert deems them sufficiently similar to the type and publishes that opinion in the scientific literature.

There is no complete, canonical list of all dinosaur taxa or holotype specimens. Attempts are regularly published in the form of books, such as the Princeton Field Guide to Dinosaurs by Gregory Paul^[11] and Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages by Thomas Holtz and Luis Rey.^[12] Where appropriate, The Paleobiology Database and Fossilworks, which are both online databases of named fossil taxa, are used to supplement the entries from published encyclopedias which are missing or data-deficient.

This list will also be updated regularly as new scientific descriptions are published and new taxa are named. The most recently named ornithopod is Minqaria bata, which was described in February 2024 by Nicholas R. Longrich, Xabier Pereda-Suberbiola, Nathalie Bardet, and Nour-Eddine Jalil.^[13]

Type System

Types are also used to diagnose higher-level taxa than an individual. One individual might represent the "type specimen" of a particular species. This species would in turn represent the "type species" of a particular genus, unless it is referred to a previously described genus. Most dinosaur genera are monospecific, therefore most type specimens are also the type species of their respective genera. On this list, the type species of a genus is only noted when it belongs to a genus with multiple referred species, such as Dryosaurus or Parasaurolophus. Furthermore, when an animal is different enough from its close relatives that it is given its own family, it is conventional in dinosaur systematics to name a family after the first described, most famous, or most abundant genus assigned to it. Therefore, on this list, the type species of any type genus for a family or sub-family level taxon is also noted when appropriate.

There are several different varieties of type specimen when referring to fossil animals:

• Holotype: This is the most common and simplest form of type specimen. A holotype is the first material of a fossil taxon that is described in the scientific literature. In order to qualify as a true holotype, all of the fossils of the type must belong to the same individual animal. All type specimens on this list are holotypes, unless otherwise indicated.
• Paratype(s): These are described in the same publication as the holotype. A paratype is designated when the fossil material is diagnostic enough to belong to the same species as the holotype, but it is not from the same individual animal. In these cases, the holotype and paratype(s) are collectively called the "type series" for that taxon. On this list, paratypes are noted in the same entry as their associated holotype.
• Neotype: When a holotype specimen is lost, destroyed, or otherwise unable to be studied further by scientists, a new type specimen for that taxon is required in order to identify future material. On this list, neotypes are only given their own entries when the holotype was never formally given a specimen number, otherwise they are noted in the entry for the holotype.
• Syntype(s): This is a type series in which no single specimen is selected to serve as a holotype, nor are any designated as paratypes. This is typically done if the fossil material is believed to be from multiple animals, but none of the individual animals were well-preserved enough to provide a complete list of diagnostic characters. These are also sometimes called "cotypes" in publications, although this is discouraged by the ICZN.
• Lectotype: When a single type specimen from a series of syntypes is designated as the new primary type specimen in a subsequent publication, this is considered to be a lectotype. On this list, lectotypes are given their own entries.
• Paralectotype(s): When a lectotype is designated from a series of syntypes, the remaining syntypes become paralectotypes as part of a reorganized type series. On this list, paralectotypes are noted alongside the list entry for the lectotype of their respective series.
• Plastotype: Sometimes, if a cast of a type specimen is made and the original type specimen is lost or destroyed, the cast can be used for the purposes of diagnostic referral to a taxon. Plastotypes are only given their own entries on this list if the holotype was not given a specimen number. Otherwise, they are noted alongside the entry for the holotype.
• Topotype: When a specimen is discovered from the same locality as a holotype specimen it may be given a new specimen number. If the second specimen is later determined to belong to the same animal as the holotype after the holotype has been described, it becomes a topotype.

All name-bearing type specimens (i.e. holotypes, lectotypes, neotypes, and syntypes) have unique entries on this list, and non-name-bearing types (i.e. paratypes, paralectotypes, topotypes, and holotypes that have been subsumed by a neotype) are noted alongside their name-bearing counterpart.

Validity

Some described species are later determined to be invalid by subsequent scientific publications. However, invalid species are sometimes resurrected, such as in the case of Brontosaurus,^[14] and sometimes the validity of a species can be controversial among researchers (e.g. the case of Torosaurus and Triceratops^{[15] [16]}). For the purposes of neutrality and completeness, all described species and genera of marginocephalians are included, even those that have been considered invalid in subsequent scientific publications.

Naming conventions and terminology follow the International Code of Zoological Nomenclature (ICZN).^[17] Technical terms used include:

• Junior synonym: A name which describes the same taxon as a previously published name. If two or more taxa are formally designated and the type specimens are later assigned to the same taxon, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
• Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. T his can be an extremely controversial designation, and as such, they are only notated when their supposedly dubious status has been formally published. Furthermore, if the scientific community has yet to reach a consensus on the validity of a name or taxon, the ongoing nature of the controversy will be stated.
• Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are not included on this list.
• Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced.

Omissions

Some ornithopod taxa are not included on this list. Nomina nuda are not included because a type does not become recognized by the ICZN until it is published in a scientific journal with a full description.

Some misidentified taxa are also not included so long as there is a scientific consensus with regard to the specimen in question. If a specimen is later referred to a taxon outside ornithopoda, it is not included on this list. However, specimens that are identified as ornithopods in publications subsequent to their initial description are included under the name they are given within ornithopoda.

Referred taxa are only included on the list as separate entries when their initial description includes a unique type specimen.

In the case of specimens with uncertain taxonomic affinity, they will be excluded unless they are considered ornithopods by the most recently published nomenclatural study of ornithischian dinosaurs, published in 2021. This excludes animals like Jeholosaurus and Thescelosaurus, which are considered basal neornithischians by the authors. This is the same definition used on the article "Ornithopoda".

List of Specimens

• Binomial name: All animals species are given a unique binomial name, typically consisting of Latin or Greek words which are used to formally and scientifically identify each species.
• Catalogue number: In most museum collections, each fossil specimen will be given a unique catalogue number which is published with the description of the fossils after they are prepared. This serves as a formal name for every single described fossil so that authors are able to refer to individual fossil discoveries in the scientific literature by name.
• Institution: Most published fossils are stored in museum collections or at universities. This is also true of type specimens, many of which are on display in museums around the world. If a type specimen has been lost, the last known location of the type is listed.
• Age: The geological stage from which the specimen was recovered is listed, when it is known. The exact age of some geological formations is not known. If this is the case, a range of possible ages is given.
• Unit: Most fossils are recovered from named geologic formations (e.g. the Morrison Formation or the Hell Creek Formation). When this is not the case, a city or landmark near the locality from which the fossil was recovered is listed.
• Material: The vast majority of fossils do not preserve the complete skeleton of an animal. In these cases, the specific bones which are fossilized have been listed.
• Notes: Other general information, such as the validity status of the taxon in question, or any other material in the type series may be listed here.

Species	Genus	Nickname	Catalogue Number	Institution	Age	Unit	Country	Notes	Images
Acristavus gagslarsoni	Acristavus		MOR 1155^[18]	Museum of the Rockies	Campanian^[19]	Two Medicine Formation	United States
Adelolophus hutchisoni	Adelolophus		UCMP 152028^[20]	University of California Museum of Paleontology	Campanian	Wahweap Formation	United States
Adynomosaurus arcanus	Adynomosaurus		MCD 7125^[21]	Museu de la Conca Dellà	Maastrichtian	Conques Formation	Spain
Ajnabia odysseus	Ajnabia		MHNM KHG 222^[22]	Marrakech Museum of Natural History	Late Maastrichtian	Upper Couche III	Morocco
Altirhinus kurzanovi	Altirhinus		PIN 3386/8^[23]	Paleontological Institute, Russian Academy of Sciences	Late Aptian-Albian	Khuren Dukh Formation	Mongolia
Amurosaurus riabinini	Amurosaurus		AEHM 1/12^[24]	Amur Natural History Museum	Middle-Late Maastrichtian	Udurchukan Formation	Russia
Anabisetia saldiviai	Anabisetia		MCF-PVPH 74^[25]	Museo Carmen Funes	Cenomanian	Lisandro Formation	Argentina
Anasazisaurus horneri	Anasazisaurus		BYU 12950^[26]	Brigham Young University Museum of Paleontology	Early Maastrichtian	Kirtland Formation	United States
Angulomastacator daviesi	Angulomastacator		TMM 43681–1^[27]	Texas Memorial Museum	Campanian	Upper Shale Member, Aguja Formation	United States
Aquilarhinus palimentus	Aquilarhinus		TMM 42452-1^[28]	Texas Memorial Museum	Campanian	Lower Shale Member, Aguja Formation	United States
Aralosaurus tuberiferus	Aralosaurus		PIN 2229^[29]	Paleontological Institute, Russian Academy of Sciences	Late Cretaceous (Turonian?)	Beleutinskaya Svita	Kazakhstan
Arenysaurus ardevoli	Arenysaurus		MPZ 2008/1^[30]	Museo Paleontológico de la Universidad de Zaragoza	Late Maastrichtian	Tremp Formation	Spain
Arstanosaurus akkurganensis	Arstanosaurus		IZAA 1/1^[31]	Institute of Zoology of the Kazakhstan Academy of Sciences	Tentatively Santonian-Campanian	Bostobinskaya Svita	Kazakhstan
Atlascopcosaurus loadsi	Atlascopcosaurus		NMV P166409^[32]	Museums Victoria	Albian	Eumeralla Formation	Australia
Augustynolophus morrisi	Augustynolophus		LACM/CIT 2852^[33]	Natural History Museum of Los Angeles County	Lower Maastrichtian^[34]	Moreno Formation	United States
Bactrosaurus johnsoni	Bactrosaurus		AMNH 6553^[35]	American Museum of Natural History	Late Cretaceous	Iren Dabasu Formation	China
Barilium dawsoni	Barilium		NHMUK R798, R798b, R799, R800-R806, R4771, R4742^[36]	Natural History Museum, London	Valanginian	Hastings Subgroup, Wadhurst Clay Formation	United Kingdom
Barsboldia sicinskii	Barsboldia		ZPAL MgD-1/110^[37]	Institute of Paleobiology, Polish Academy of Sciences	Upper Campanian-Maastrichtian	Nemegt Formation	Mongolia
Batyrosaurus rozhdestvenskyi	Batyrosaurus		AEHM 4/1^[38]	Amur Natural History Museum	Santonian-Campanian	Bostobinskaya Svita	Kazakhstan
Bayannurosaurus perfectus	Bayannurosaurus		IMMNH PV00001^[39]	Inner Mongolia Museum of Natural History	Early Aptian	Bayingebi Formation	China
Blasisaurus canudoi	Blasisaurus		MPZ99/667^[40]	Museo Paleontológico de la Universidad de Zaragoza	Latest Maastrichtian	Arén Formation	Spain
Bolong yixianensis	Bolong		YHZ-001^[41]	Yizhou Fossil Museum	Barremian-Aptian^[42]	Yixian Formation^[43]	China
Bonapartesaurus rionegrensis	Bonapartesaurus		MCPA-Pv SM2^[44]	Museo Provincial Carlos Ameghino	Late Campanian-Early Maastrichtian	Allen Formation	Argentina
Brachylophosaurus canadensis	Brachylophosaurus		CMN 8893^[45]	Canadian Museum of Nature	>77.76 Ma(Campanian)	Oldman Formation	Canada
Brighstoneus simmondsi	Brighstoneus		MIWG 6344^[46]	Dinosaur Isle Museum	Early Barremian	Wessex Formation	United Kingdom
Burianosaurus augustai	Burianosaurus		NMP Ob 203^[47]	National Museum Prague	Late Cenomanian	Peruc-Korycany Formation	Czechia
Callovosaurus leedsi	Callovosaurus		BMNH R1993	Natural History Museum, London	Middle Callovian	Peterborough Member, Oxford Clay Formation	United Kingdom
Calvarius rapidus	Calvarius		MCD-8734^[48]	Museu de la Conca Dellà, Isona	Maastrichtian	Talarn Formation, Tremp Group	Spain
Camptosaurus dispar	Camptosaurus		YPM 1877	Peabody Museum of Natural History	Upper Jurassic^[49]	Morrison Formation	United States
Canardia garonnensis	Canardia		MDE-Ma3–16^[50]	Musée des Dinosaures d'Espéraza	Upper Maastrichtian	Marnes d'Auzas Formation	France
Cedrorestes crichtoni	Cedrorestes		DMNH 47994	Denver Museum of Nature and Science	Lower Cretaceous (Barremian?)	Cedar Mountain Formation	United States
Changmiania liaoningensis	Changmiania		PMOL AD00114^[51]	Paleontological Museum of Liaoning	Barremian	Yixian Formation	China
Charonosaurus jiayinensis	Charonosaurus		CUST J-V1251-57^[52]	Changchun University of Sciences and Technology	Late Maastrichtian	Yuliangze Formation	China
Choyrodon barsboldi	Choyrodon		MPC-D 100/801^[53]	Institute of Paleontology and Geology, Mongolian Academy of Sciences	Middle-Late Albian	Khuren Dukh Formation	Mongolia
Cionodon arctatus	Cionodon		AMNH 3951^[54]	American Museum of Natural History	Late Cretaceous	Laramie Formation	United States	Nomen dubium
Claosaurus agilis	Claosaurus		YPM 1190^[55]	Peabody Museum of Natural History	Late Coniacian	Smoky Hill Chalk Member, Niobrara Chalk	United States
Convolosaurus marri	Convolosaurus		SMU 72834^[56]	Shuler Museum of Paleontology	Aptian	Twin Mountains Formation	United States
Corythosaurus casuarius	Corythosaurus		AMNH 5240^[57]	American Museum of Natural History	Cretaceous	Belly River Group	Canada
Cumnoria prestwichii	Cumnoria		OUM J.3303^[58]	Oxford University Museum of Natural History	Upper Jurassic	Kimmeridge Clay^[59]	United Kingdom
Dakotadon lakotaensis	Dakotadon		SDSM 8656^[60]	South Dakota School of Mines and Technology	Early Cretaceous (Barremian?)	Lakota Formation	United States
Darwinsaurus evolutionis	Darwinsaurus		NHMUK R1831, NHMUK R1833, NHMUK R1835 and NHMUK R1836^[61]	Natural History Museum, London	Valanginian	Wadhurst Clay Formation	United Kingdom	Nomen dubium, type specimens can be referred to Hypselospinus fittoni and Mantellisaurus atherfieldensis
Datonglong tianzhensis	Datonglong		SXMG V 00005^[62]	Shanxi Museum of Geology	Upper Cretaceous	Huiquanpu Formation	China
Delapparentia turolensis	Delapparentia		MPT/I.G^[63]	Museo de Teruel	Lower Barremian	Camarillas Formation	Spain
Diclonius pentagonus	Diclonius		AMNH 3972^[64]	American Museum of Natural History	Campanian^[65]	Judith River Formation	United States^[66]	Nomen dubium
Diluvicursor pickeringi	Diluvicursor		NMV P221080^[67]	Museums Victoria	Lower Albian	Eumeralla Formation	Australia
Draconyx loureiroi	Draconyx		ML 357^[68]	Museum of Lourinhã	Tithonian	Praia Azul Member, Lourinhã Formation^[69]	Portugal
Dryosaurus altus	Dryosaurus		YPM 1876^[70]	Peabody Museum of Natural History	Upper Kimmeridgian?	Morrison Formation	United States
Dryosaurus elderae	Dryosaurus		CM 3392^[71]	Carnegie Museum of Natural History	Upper Jurassic	Morrison Formation	United States
Dysalotosaurus lettowvorbecki	Dysalotosaurus				Kimmeridgian^[72]	Tendaguru Beds	Tanzania
Edmontosaurus annectens	Edmontosaurus		USNM 2414	National Museum of Natural History	Maastrichtian	Lance Formation^[73]	United States
Edmontosaurus regalis	Edmontosaurus		Holotype: CMN 2288^[74] Paratype: CMN 2289	Canadian Museum of Nature	Campanian	Horsethief Member, Horseshoe Canyon Formation	Canada
Elrhazosaurus nigeriensis	Elrhazosaurus		MNHN GDF 332^[75]	Muséum National d'Histoire Naturelle	Aptian	Elrhaz Formation	Niger
Eolambia caroljonesa	Eolambia		CEUM 9758^[76]	USU Eastern Prehistoric Museum	Albian-Cenomanian	Mussentuchit Member, Cedar Mountain Formation	United States
Eotrachodon orientalis	Eotrachodon		MSC 7949^[77]	McWane Science Center	Latest Santonian	Mooreville Chalk	United States
Eousdryosaurus nanohallucis	Eousdryosaurus		SHN(JJS)-170^[78]	Sociedade de Historia Natural	Upper Kimmeridgian	Alcobaça Formation	Portugal
Equijubus normani	Equijubus		IVPP V12534^[79]	Institute of Vertebrate Paleontology and Paleoanthropology	Early Cretaceous	Xinminbao Group	China
Eucercosaurus tanyspondylus	Eucercosaurus		Syntypes: CAMSM B55610–29^[80]	Sedgwick Museum of Earth Sciences	Upper Albian	Gault Formation, reworked into Cambridge Greensand	United Kingdom	Nomen dubium
Fostoria dhimbangunmal	Fostoria		LRF 3050.A^[81]	Australian Opal Centre	Cenomanian	Griman Creek Formation	Australia
Fukuisaurus tetoriensis	Fukuisaurus		FPDM-V-40-1 and FPDM-V-40-2^[82]	Fukui Prefectural Dinosaur Museum	Late Hauterivian-Barremian	Kitadani Formation	Japan
Fulgurotherium australe	Fulgurotherium		BMNH R 3719^[83]	Natural History Museum, London	Albian	Wallangalla Sandstone Member, Griman Creek Formation	Australia	Nomen dubium
Fylax thyrakolasus	Fylax		IPS-36338^[84]	Museu de l'InstitutCatalà de Paleontologia Miquel Crusafont	Maastrichtian	Figuerola Formation	Spain
Galleonosaurus dorisae	Galleonosaurus		NMV P229196	Museums Victoria	Upper Barremian	Wonthaggi Formation	Australia
Gasparinisaura cincosaltensis	Gasparinisaura		MUCPv-208^[85]	Museum of the Universidad Nacional del Comahue	Coniacian-Santonian	Anacleto Member, Río Colorado Formation	Argentina
Gideonmantellia amosanjuanae	Gideonmantellia		MPG-PBCH^[86]	Museo Paleontológico de Galve	Barremian	Camarillas Formation	Spain
Gilmoreosaurus mongoliensis	Gilmoreosaurus		AMNH 6551^[87]	American Museum of Natural History	Cenomanian?	Iren Dabasu Formation	China^[88]
Glishades ericksoni	Glishades		AMNH 27414^[89]	American Museum of Natural History	Upper Cretaceous	Two Medicine Formation	United States
Gobihadros mongoliensis	Gobihadros		MPC-D100/746^[90]	Mongolian Paleontological Center	Cenomanian-Santonian	Baynshire Formation	Mongolia
Gongpoquansaurus mazongshanensis	Gongpoquansaurus		IVPP V 11333^[91]	Institute of VertebratePaleontology and Paleoanthropology	Albian	Zhonggou Formation	China
Gonkoken nanoi	Gonkoken		CPAP 3054^[92]	Paleontological Collection of Antarctica and Patagonia	Lower Maastrichtian	Dorotea Formation	Chile
Gryposaurus latidens	Gryposaurus		AMNH FARB 5465^[93]	American Museum of Natural History	Campanian	Two Medicine Formation	United States
Gryposaurus monumentensis	Gryposaurus		RAM 6797	Raymond M. Alf Museum of Palaeontology	Late Campanian	Kaiparowits Formation	United States
Gryposaurus notabilis	Gryposaurus		CMN 2278^[94]	Canadian Museum of Nature	Campanian	Dinosaur Park Formation	Canada
Hadrosaurus foulkii	Hadrosaurus		ANSP 9201-9204, 10005^[95]	Academy of Natural Sciences of Philadelphia	Campanian	Woodbury Formation	United States
Hippodraco scutodens	Hippodraco		UMNH VP 20208^[96]	Utah Museum of Natural History	Upper Barremian—Lowermost Aptian	Yellow Cat Member, Cedar Mountain Formation	United States
Huallasaurus australis	Huallasaurus		MACN-PV 2^[97]	Museo Argentino de Ciencias Naturales Bernardino Rivadavia	Campanian-Maastrichtian	Los Alamitos Formation	Argentina
Huehuecanauhtlus tiquichensis	Huehuecanauhtlus		IGM 6253^[98]	Instituto de Geología, Universidad Nacional Autónoma de México	Santonian		Mexico
Huxleysaurus hollingtoniensis	Huxleysaurus		NHMUK R1148	Natural History Museum, London		Wadhurst Clay Formation	United Kingdom	Nomen dubium, referred to Hypselospinus fittoni
Hypacrosaurus altispinus	Hypacrosaurus		AMNH 5204^[99]	American Museum of Natural History	Upper Cretaceous	Horseshoe Canyon Formation	Canada
Hypacrosaurus stebingeri	Hypacrosaurus		MOR 549^[100]	Museum of the Rockies	Late Cretaceous^[101]		United States
Hypselospinus fittoni	Hypselospinus		NHMUK R1635	Natural History Museum, London	Valanginian	Wadhurst Clay Formation	United Kingdom
Hypsibema crassicauda	Hypsibema		USNM 7189^[102]	National Museum of Natural History	Cretaceous	Black Creek Formation	United States^[103]
Hypsibema missouriensis	Hypsibema		USNM 16733^[104]	National Museum of Natural History	Early Maastrichtian?	Ripley Formation	United States
Iani smithi	Iani		NCSM 29373^[105]	North Carolina Museum of Natural Sciences	Cenomanian	Mussentuchit Member, Cedar Mountain Formation	United States
Iguanacolossus fortis	Iguanacolossus		UMNH VP 20205	Utah Museum of Natural History	Lower Barremian	Yellow Cat Member, Cedar Mountain Formation	United States
Iguanodon bernissartensis	Iguanodon		IRSNB 1534^[106]	Royal Belgian Institute of Natural Sciences	Upper Barremian-Lowermost Aptian	Sainte-Barbe Clays Formation	Belgium
Iguanodon galvensis	Iguanodon		MAP-4787	Museo Aragonés de Paleontología	Lower Barremian	Camarillas Formation	Spain
Isasicursor santacrucensis	Isasicursor		MPM 21525^[107]	Museo Padre Molina	Upper Campanian-Lower Maastrichtian	Chorillo Formation	Argentina
Iyuku raathi	Iyuku		AM 6150^[108]	Albany Museum	Valanginian	Kirkwood Formation	South Africa
Jaxartosaurus aralensis	Jaxartosaurus		PIN 1/5009^[109]	Paleontological Institute, Russian Academy of Sciences	Lower Santonian	Dabrazinskaya Svita	Kazakhstan
Jeyawati rugoculus	Jeyawati		MSM P4166^[110]	Arizona Museum of Natural History	Turonian	Moreno Hill Formation	United States
Jintasaurus meniscus	Jintasaurus		FDRC: GJ 06-2-52^[111]	Fossil Research and Development Center, Third Geology and Mineral Resources Exploration Academy of the Gansu Provincial Bureau of Geo-Exploration and Mineral Development	Early Cretaceous (Albian?)	Xinminpu Group	China
Jinzhousaurus yangi	Jinzhousaurus		IVPP V12691^[112]	Institute of Vertebrate Paleontology and Paleoanthropology	Hauterivian-Barremian	Yixian Formation	China
Kamuysaurus japonicus	Kamuysaurus		HMG-1219^[113]	Hobetsu Museum	Early Maastrichtian	Hakobuchi Formation, Yezo Group	Japan
Kangnasaurus coetzeei	Kangnasaurus		SAM 2732^[114]	Iziko South African Museum			South Africa
Kazaklambia convincens	Kazaklambia		PIN 2230/1	Paleontological Institute, Russian Academy of Sciences	Santonian	Dabrazinskaya Svita	Kazakhstan
Kelumapusaura machi	Kelumapusaura		MPCN-PV-807	Museo Patagónico de Ciencias Naturales Juan Carlos Salgado	Maastrichtian	Allen Formation	Argentina
Kerberosaurus manakini	Kerberosaurus		AENM 1/319^[115]	Amur Natural History Museum	Maastrichtian	Tsagayan Formation	Russia
Koshisaurus katsuyama	Koshisaurus				Early Cretaceous^[116]	Kitadani Formation	Japan
Koutalisaurus kohlerorum	Koutalisaurus		IPS SRA 27	Institut de Paleontologia Miquel Crusafont	Maastrichtian	Tremp Formation	Spain	Likely synonymous with Pararhabdodon isonensis
Kritosaurus navajovius	Kritosaurus		AMNH 5799^[117]	American Museum of Natural History	Cretaceous	Ojo Alamo Formation	United States
Kukufeldia tilgatensis	Kukufeldia		NHMUK 28660^[118]	Natural History Museum, London	Middle-Upper Valanginian	Grinstead Clay Member, Tunbridge Wells Sand Formation	United Kingdom
Kundurosaurus nagornyi	Kundurosaurus		AENM 2/921^[119]	Amur Natural History Museum	Maastrichtian	Udurchukan Formation	Russia
Laiyangosaurus youngi	Laiyangosaurus		IVPP V 23401^[120]	Institute of Vertebrate Paleontology and Paleoanthropology	Upper Cretaceous	Jingangkou Formation, Wangshi Group	China
Lambeosaurus clavinitialis	Lambeosaurus				Campanian^[121]	Oldman Formation	Canada
Lambeosaurus lambei	Lambeosaurus		CMN 2869^[122]	Canadian Museum of Nature	Upper Cretaceous^[123]	Belly River Formation	Canada
Lambeosaurus magnicristatus	Lambeosaurus		CMN 8705^[124]	Canadian Museum of Nature	Late Cretaceous	Dinosaur Park Formation	Canada
Lanzhousaurus magnidens	Lanzhousaurus		GSLTZP01-001^[125]	Fossil Research and Development Center of the Third Geology and Mineral Resources Exploration Academy of Gansu Province	Early Cretaceous	Hekou Group	China
Lapampasaurus cholinoi	Lapampasaurus		MPHN-Pv-01^[126]	Museo Provincial de Historia Natural	Late Campanian-Early Maastrichtian	Allen Formation	Argentina
Latirhinus uitstlani	Latirhinus		IGM 6583^[127]	Instituto de Geología, Universidad Nacional Autónoma de México	Upper Campanian	Cerro del Pueblo Formation	Mexico
Levnesovia transoxiana	Levnesovia		USNM 538191^[128]	National Museum of Natural History	Middle–Late Turonian	Bissekty Formation	Uzbekistan
Loncosaurus argentinus	Loncosaurus		MACN-1629^[129]	Museo Argentino de Ciencias Naturales Bernardino Rivadavia	Upper Cretaceous?		Argentina	Nomen vanum
Lophorhothon atopus	Lophorhothon		FMNH P 27383^[130]	Field Museum of Natural History	Late Cretaceous	Mooreville Chalk Formation	United States
Lurdusaurus arenatus	Lurdusaurus		MNHN GDF 1700^[131]	Muséum National d'Histoire Naturelle	Aptian	Elrhaz Formation	Niger
Macrogryphosaurus gondwanicus	Macrogryphosaurus		MUCPv-321^[132]	Museo Universidad Nacional del Comahue	Coniacian	Portezuelo Formation	Argentina
Magnamanus soriaensis	Magnamanus		MNS 200/132, 2001/122, 2002/95, 2003/69, 2004/54^[133]	Museo Numantino de Soria	Upper Hauterivian-Lower Barremian	Golmayo Formation	Spain
Magnapaulia laticaudus	Magnapaulia		LACM 17715^[134]	Natural History Museum of Los Angeles County	Campanian	El Gallo Formation	Mexico
Mahuidacursor lipanglef	Mahuidacursor		MAU-Pv-CO-596^[135]	Museo Municipal Argentino Urquiza	Santonian	Bajo de la Carpa Formation	Argentina
Maiasaura peeblesorum	Maiasaura		YPM-PU 22405^[136]	Peabody Museum of Natural History	Campanian^[137]	Two Medicine Formation	United States
Malefica deckerti	Malefica		TxVP 41917-1^[138]	Jackson School Museum of Earth History	Late Campanian	Aguja Formation	United States
Mandschurosaurus amurense	Mandschurosaurus				Cretaceous^[139]	Yuliangze Formation	China	Nomen dubium
Mantellisaurus atherfieldensis	Mantellisaurus		NHMUK R5764^[140]	Natural History Museum, London	Barremian-Aptian	Vectis Formation	United Kingdom
Mantellodon carpenteri	Mantellodon		NHMUK R3741	Natural History Museum, London	Lower Aptian	Lower Greensand Formation	United Kingdom	Nomen dubium, can be referred to Mantellisaurus atherfieldensis
Matheronodon provincialis	Matheronodon		MMS/VBN-02-102^[141]	Musée du Moulin Seigneurial	Late Campanian		France
Microhadrosaurus nanshiungensis	Microhadrosaurus		IVPP V4732^[142]	Institute of Vertebrate Paleontology and Paleoanthropology	Late Cretaceous	Nanxiong Formation	China
Minqaria bata	Minqaria		MHNM.KHG.1395	Museum d'Histoire Naturelle de Marrakech, Cadi Ayyad University	Upper Maastrichtian	Ouled Abdoun Basin	Morocco
Mochlodon suessi	Mochlodon		PIUW 2349^[143]	Paläontologisches Institut, University of Vienna	Lower Campanian	Grünbach Formation	Austria
Mochlodon vorosi	Mochlodon		MTM V 2010.105.1	Hungarian Natural History Museum	Santonian	Csehbánya Formation	Hungary
Morelladon beltrani	Morelladon		CMP-MS-03^[144]	Museo de la Valltorta	Barremian	Arcillas de Morella Formation	Spain
Morrosaurus antarcticus	Morrosaurus		MACN Pv 197^[145]	Bernardino Rivadavia Natural Sciences Argentine Museum	Maastrichtian	Cape Lamb Member, López de Bertodano Formation	Antarctica
Muttaburrasaurus langdoni	Muttaburrasaurus		QM F6140^[146]	Queensland Museum	Albian	Mackunda Formation	Australia
Naashoibitosaurus ostromi	Naashoibitosaurus		NMMNH P-16106^[147]	New Mexico Museum of Natural History and Science	Late Campanian	Kirtland Formation	United States
Nanningosaurus dashiensis	Nanningosaurus		NHMG8142^[148]	Natural History Museum of Guangxi	Upper Cretaceous	Nadu Formation	China
Nanyangosaurus zhugeii	Nanyangosaurus		IVPP V 11821^[149]	Institute of Vertebrate Paleontology and Paleoanthropology	Early Cretaceous	Xiaguan Formation	China
Napaisaurus guangxiensis	Napaisaurus		FS-20-007, FS-20-007^[150]		Early Cretaceous	Xinlong Basin	China
Nipponosaurus sachalinensis	Nipponosaurus		UHR 6590^[151]	University of Hokkaido	Late Santonian-Early Campanian	Upper Yezo Group	Russia
Notohypsilophodon comodorensis	Notohypsilophodon		UNPSJB — PV 942^[152]	National University of the Patagonia San Juan Bosco	Cenomanian?	Bajo Barreal Formation	Argentina
Oblitosaurus bunnueli	Oblitosaurus		CPT-1440, CPT-1444, and MAP-8290 to MAP-8299^[153]	Museo Aragonés de Paleontología	Late Jurassic	Villar del Arzobispo Formation	Spain
Oligosaurus adelus	Oligosaurus		PIUW 3518^[154] ^[155]	Paläontologisches Institut der Universität Wien	Late Cretaceous	Gosau Group	Austria
Olorotitan arharensis	Olorotitan		AEHM 2/845^[156]	Amur Natural History Museum	Middle-Late Maastrichtian	Tsagayan Formation	Russia
Ornatops incantatus	Ornatops		WSC 10058^[157]	Western Science Center	Middle Campanian	Allison Member, Menefee Formation	United States
Ornithomerus gracilis	Ornithomerus		PIUW 2349/3	Paläontologisches Institut der Universität Wien	Late Cretaceous	Gosau Group	Austria
Orthomerus dolloi	Orthomerus		NHMUK 42954-57^[158]	Natural History Museum, London	Late Maastrichtian	Likely Maastricht Formation	The Netherlands or Belgium
Osmakasaurus depressus	Osmakasaurus		USNM 4753^[159]	National Museum of Natural History	Barremian-Aptian	Lakota Formation	United States
Ouranosaurus nigeriensis	Ouranosaurus		MNHN GDF 300^[160]	Muséum National d'Histoire Naturelle	Late Aptian	Upper Elrhaz Formation	Niger
Owenodon hoggii	Owenodon		NHM R2998^[161]	Natural History Museum, London	Mid-Berriasian	Purbeck Limestone Formation	United Kingdom
Pararhabdodon isonensis	Pararhabdodon		Holotype: IPS SRA 1^[162] Paratype: IPS SRA 15-18	Institut de Paleontologia Miquel Crusafont	Maastrichtian	Tremp Formation	Spain
Parasaurolophus cyrtocristatus	Parasaurolophus		FMNH P-27393^[163]	Field Museum of Natural History	Campanian(75.02-76.14 Ma)	Fossil Forest Member, Fruitland Formation	United States
Parasaurolophus tubicen	Parasaurolophus		PMU.R.1250^[164]	Paleontologiska Museet, Uppsala	Campanian(73.5-73.6 Ma)	Kirtland Formation	United States
Parasaurolophus walkeri	Parasaurolophus		ROM 768^[165]	Royal Ontario Museum	Campanian(76.8-76.9 Ma)	Dinosaur Park Formation	Canada
Pareisactus evrostos	Pareisactus		MCD 5371^[166]	Museu de la Conca Dellà	Lower Maastrichtian	Conques Formation	Spain
Parksosaurus warreni	Parksosaurus		ROM 804^[167]	Royal Ontario Museum	Campanian-Maastrichtian	Horseshoe Canyon Formation	Canada
Penelopognathus weishampeli	Penelopognathus		IMM 2002-BYGB-1^[168]	Inner Mongolia Museum	Albian	Bayan Gobi Formation	China
Planicoxa venenica	Planicoxa		DMNH 42504^[169]	Denver Museum of Natural History	Lower Cretaceous	Poison Strip Member, Cedar Mountain Formation	United States
Plesiohadros djadokhtaensis	Plesiohadros		MPC-D100/745^[170]	Mongolian Paleontology Center	Upper Campanian?	Djadokhta Formation	Mongolia
Portellsaurus sosbaynati	Portellsaurus		MQ98-II-1^[171]	Colección Museográfica de Cinctorres	Barremian	Margas de Mirambell Formation	Spain
Proa valdearinnoensis	Proa		AR-1/19^[172]	Museo Aragonés de Paleontología	Lower Albian	Escucha Formation	Spain
Probactrosaurus gobiensis	Probactrosaurus		PIN 2232/1^[173]	Paleontological Institute, Russian Academy of Sciences	Barremian-Albian		China
Probrachylophosaurus bergei	Probrachylophosaurus		MOR 2919	Museum of the Rockies	Campanian(78.5-78.2 Ma)	Judith River Formation	United States
Proplanicoxa galtoni	Proplanicoxa		NHMUK R8649	Natural History Museum, London	Early Cretaceous		United Kingdom	Nomen dubium, likely referrable to Mantellisaurus atherfieldensis
Prosaurolophus maximus	Prosaurolophus		AMNH 5386^[174]	American Museum of Natural History	Campanian^[175]	Dinosaur Park Formation	Canada
Protohadros byrdi	Protohadros		SMU 74582^[176]	Shuler Museum of Paleontology	Mid-Cenomanian	Woodbine Formation	United States
Pteropelyx grallipes	Pteropelyx		AMNH 3791	American Museum of Natural History				Bones indistinguishable from Corythosaurus
Qantassaurus intrepidus	Qantassaurus		NMV P199075	Museums Victoria	Upper Barremian	Wonthaggi Formation	Australia
Ratchasimasaurus suranareae	Ratchasimasaurus		NRRU-A2064^[177]	Northeastern Research Institute of Petrified Wood and Mineral Resources, Nakhon Ratchasima Rajabhat University	Aptian	Khok Kruat Formation	Thailand
Rhabdodon priscus	Rhabdodon				Early Maastrichtian^[178]	Rognacian Formation	France
Rhabdodon septimanicus	Rhabdodon				Late Campanian-Early Maastrichtian	Grès à Reptiles	France
Rhinorex condrupus	Rhinorex		BYU 13258^[179]	Brigham Young University Museum of Paleontology	Campanian	Neslen Formation	United States
Riabininohadros weberae	Riabininohadros		ZGTM 5751^[180]	ChernyshevCentral Research Museum of Geology and Exploration	Upper Maastrichtian		Ukraine
Sahaliyania elunchunorum	Sahaliyania		GMH W453^[181]	Geological Museum of Heilongjiang	Maastrichtian	Yuliangze Formation	China
Saurolophus angustirostris	Saurolophus		PIN 551/8^[182]	Paleontological Institute, Russian Academy of Sciences	Upper Campanian-Lower Maastrichtian	Nemegt Formation	Mongolia
Saurolophus osborni	Saurolophus		AMNH 5220^[183]	American Museum of Natural History	Early Maastrichtian	Horseshoe Canyon Formation	Canada
Secernosaurus koerneri	Secernosaurus		FMNH P13423^[184]	Field Museum of Natural History	Late Campanian-Early Maastrichtian	Bajo Barreal Formation	Argentina
Sektensaurus sanjuanboscoi	Sektensaurus		UNPSJB-PV 1054^[185]	National University of Patagonia San Juan Bosco	Coniacian-Maastrichtian	Lago Colhué Huapi Formation	Argentina
Sellacoxa pauli	Sellacoxa		BMNH R 3788^[186]	Natural History Museum, London	Lower Valanginian	Lower Wadhurst Clay Formation	United Kingdom
Shantungosaurus giganteus	Shantungosaurus		GMV 1780-1^[187]	Geological Museum of China	Middle-Late Campanian	Wangshi Group	China
Shuangmiaosaurus gilmorei	Shuangmiaosaurus		LPM 0165^[188]	Liaoning Paleontological Museum	Mid-Cretaceous	Sunjiawan Formation	China
Siamodon nimngami	Siamodon		PRC-4^[189]	Paleontological Research and Education Centre, Mahasarakham University	Aptian	Khok Kruat Formation	Thailand
Siluosaurus zhanggiani	Siluosaurus		IVPP V.11117 (1-2)	Institute of Vertebrate Paleontology and Paleoanthropology	Early Cretaceous^[190]	Gongpoquan Basin	China	Nomen dubium^[191]
Sirindhorna khoratensis	Sirindhorna		NRRU3001-166^[192]	Northeastern Research Institute of Petrified Wood and Mineral Resources, Nakhon Ratchasima Rajabhat University	Aptian	Khok Kruat Formation	Thailand
Syngonosaurus macrocercus	Syngonosaurus		CAMSM B55570-78, B55580-86	Sedgwick Museum of Earth Sciences	Upper Albian	Gault Formation	United Kingdom
Talenkauen santacrucensis	Talenkauen		MPM-10001^[193]	Museo Padre Molina	Maastrichtian	Pari Aike Formation	Argentina
Tanius sinensis	Tanius		PMU 24720^[194]	Museum of Evolution of Uppsala University	Upper Campanian-Lower Maastrichtian	Jiangjunding Formation^[195]	China
Telmatosaurus transsylvanicus	Telmatosaurus		BMNH R.3386^[196]	Natural History Museum, London	Upper Maastrictian	Sânpetru Formation^[197]	Romania
Tenontosaurus dossi	Tenontosaurus		FWMSH 93B1^[198]	Fort Worth Museum of Science and History	Early Cretaceous	Twin Mountains Formation	United States
Tenontosaurus tilletti	Tenontosaurus		AMNH 3040^[199]	American Museum of Natural History	Aptian-Albian	Cloverly Formation	United States
Tethyshadros insularis	Tethyshadros		SC 57021^[200]	Italian State Collections	Late Campanian-Early Maastrichtian	Liburnian Formation	Italy
Theiophytalia kerri	Theiophytalia		YPM 1887	Peabody Museum of Natural History	Lower Cretaceous	Purgatoire Formation	United States
Thespesius occidentalis	Thespesius		USNM 219, 220, 221^[201]	National Museum of Natural History	Late Cretaceous	Lance Formation^[202]	United States	Nomen dubium
Tlatolophus galorum	Tlatolophus		CIC/P/147^[203]	Colección Paleontológica del Centro INAH	Upper Campanian	Cerro del Pueblo Formation	Mexico
Trachodon mirabilis	Trachodon		ANSP 9260	Academy of Natural Sciences of Philadelphia	Late Cretaceous	Judith River Formation	United States	Nomen dubium
Transylvanosaurus platycephalus	Transylvanosaurus		LPB (FGGUB) R.2070^[204]	University of Tübingen	Maastrichtian	Pui Beds	Romania
Trinisaura santamartaensis	Trinisaura		MLP08-III-1-1^[205]	Museo de La Plata	Upper Campanian	Snow Hill Island Formation	Antarctica
Tsintaosaurus spinorhinus	Tsintaosaurus		IVPP V 725^[206]	Institute of Vertebrate Paleontology and Paleoanthropology	Late Campanian-Early Maastrichtian	Jingangkou Formation^[207]	China
Ugrunaaluk kuukpikensis	Ugrunaaluk		UAMES 12995^[208]	University of Alaska Museum	Lower Maastrichtian	Prince Creek Formation	United States
Uteodon aphanoecetes	Uteodon		CM 11337	Carnegie Museum of Natural History	Lower-Middle Tithonian	Brushy Basin Member, Morrison Formation	United States
Valdosaurus canaliculatus	Valdosaurus		BMNH R185, R186	Natural History Museum, London	Barremian	Wealden Group	United Kingdom
Velafrons coahuilensis	Velafrons		CPC-59^[209]	Paleontological Collection of Coahuila	Late Campanian	Cerro del Pueblo Formation	Mexico
Weewarrasaurus pobeni	Weewarrasaurus		LRF 3067^[210]	Australian Opal Centre	Early Cenomanian	Wallangulla Sandstone, Griman Creek Formation	Australia
Willinakaqe salitralensis	Willinakaqe		MPCA-Pv SM 8^[211]	Museo Provincial Carlos Ameghino	Late Cretaceous	Allen Formation	Argentina	Nomen vanum^[212]
Wulagasaurus dongi	Wulagasaurus		GMH W184^[213]	Geological Museum of Heilongjiang	Maastrichtian	Yuliangze Formation	China
Xuwulong yueluni	Xuwulong		GSGM F00001^[214]	Gansu Geological Museum	Aptian-Albian	Xinminpu Group	China
Yamatosaurus izanagii	Yamatosaurus		MNHAH D1-033516^[215]	Museum of Nature and Human Activities	Early Maastrictian (71.94-71.69 Ma)	Kita-ama Formation	Japan
Yueosaurus tiantaiensis	Yueosaurus		ZMNH M8620^[216]	Zhejiang Museum of Natural History	Albian-Cenomanian	Liangtoutang Formation	China
Yunganglong datongensis	Yunganglong		SXMG V 00001^[217]	Shanxi Museum of Geology	Upper Cretaceous	Zhumapu Formation	China
Zalmoxes robustus	Zalmoxes		BMNH R.3392^[218]	Natural History Museum, London	Upper Maastrichtian	Sânpetru Formation	Romania
Zalmoxes shqiperorum	Zalmoxes		BMNH R.4900	Natural History Museum, London	Upper Maastrichtian		Romania
Zhanghenglong yangchengensis	Zhanghenglong		XMDFEC V0013^[219]	XIxia Museum of Dinosaur Fossil Eggs of China	Middle Santonian	Majiacun Formation	China
Zuoyunlong huangi	Zuoyunlong		SXMG V 00004^[220]	Shanxi Museum of Geology	Cenomanian?	Zhumapu Formation	China

See also

• List of marginocephalian type specimens
• List of thyreophoran type specimens
• List of other ornithischian type specimens
• List of non-avian theropod type specimens
• List of sauropodomorph type specimens
• List of Mesozoic birds

Notes and References

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3. 10.1080/08912963.2020.1793979 . A new phylogeny of cerapodan dinosaurs . 2021 . Dieudonné . P. -E. . Cruzado-Caballero . P. . Godefroit . P. . Tortosa . T. . Historical Biology . 33 . 10 . 2335–2355 . 221854017 .
4. Madzia. Daniel. Boyd. Clint A.. Mazuch. Martin. A basal ornithopod dinosaur from the Cenomanian of the Czech Republic. Journal of Systematic Palaeontology. 16. 11. 2017. 967–979. 10.1080/14772019.2017.1371258 . 90008574.
5. 10.2992/007.079.0103. A Medium-Sized Ornithopod (Dinosauria: Ornithischia) from the Upper Cretaceous Bajo Barreal Formation of Lago Colhué Huapi, Southern Chubut Province, Argentina. 2010. Ibiricu. Lucio M.. Martínez. Rubén D.. Lamanna. Matthew C.. Casal. Gabriel A.. Luna. Marcelo. Harris. Jerald D.. Lacovara. Kenneth J.. 53407321. Annals of Carnegie Museum. 79. 39–50.
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