List of non-avian theropod type specimens explained

This list of non-avian theropod type specimens is a list of fossils that are the official standard-bearers for inclusion in the Mesozoic species and genera of the dinosaur clade Theropoda, which includes the carnivorous dinosaurs like Tyrannosaurus and Velociraptor, their herbivorous relatives like the therizinosaurs, and birds. Type specimens are those that are definitionally members of biological taxa, and additional specimens can only be "referred" to these taxa if an expert deems them sufficiently similar to the type. Theropods have been found all over the globe, and are alive today as birds

The list

Species	Genus	Nickname	Catalogue number	Institution	Age	Unit	Country	Notes	Images
Abelisaurus comahuensis	Abelisaurus		MC 11098^[1]	Museo de Cipolleti	Maastrichtian	Allen Formation	Argentina
Acheroraptor temertyorum	Acheroraptor		ROM 63777^[2]	Royal Ontario Museum	Upper Maastrichtian	Hell Creek Formation	United States
Achillesaurus manazzonei	Achillesaurus		MACN-PV-RN 1116^[3]	Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”	Santonian	Bajo de la Carpa Formation	Argentina
Achillobator giganticus	Achillobator		MNUFR-15^[4]	Mongolian National University	Santonian-Campanian	Bayan Shireh Formation	Mongolia
Acrocanthosaurus atokensis	Acrocanthosaurus		Holotype: OMNH 10146^[5] Paratype: OMNH 10147	Oklahoma Museum of Natural History	Albian-Aptian	Antlers Formation	United States^[6]
Adasaurus mongoliensis	Adasaurus		IGM 100/20^[7]	Mongolian Institute of Geology	Late Cretaceous	Nemegt Formation	Mongolia
Aepyornithomimus tugrikinensis	Aepyornithomimus		MPC-D 100/130^[8]	Institute of Paleontology and Geology, Mongolian Academy of Sciences	Campanian	Djadokhta Formation	Mongolia
Aerosteon riocoloradensis	Aerosteon		MCNA-PV-3137^[9]	Museo de Ciencias Naturales y Antropológicas (J. C. Moyano) de Mendoza	Santonian	Anacleto Formation	Argentina
Afromimus tenerensis	Afromimus		MNBH GAD112^[10]	Musée National Boubou Hama	Aptian-Albian	Elrhaz Formation	Niger
Afrovenator abakensis	Afrovenator		UC OBA 1^[11]	University of Chicago	Middle Jurassic^[12]	Tiouaren Formation	Niger
Albertavenator curriei	Albertavenator		TMP 1993.105.0001^[13]	Royal Tyrell Museum of Paleontology	Earliest Maastrichtian	Horsethief Member, Horseshoe Canyon Formation	Canada
Albertonykus borealis	Albertonykus		TMP 2001.45.91^[14]	Royal Tyrell Museum of Paleontology	Maastrichtian	Horseshoe Canyon Formation	Canada
Albertosaurus arctunguis	Albertosaurus		ROM 807^[15]	Royal Ontario Museum	Campanian-Maastrichtian^[16]	Horseshoe Canyon Formation	Canada	Synonymous with Albertosaurus sarcophagus
Albertosaurus sarcophagus	Albertosaurus		CMN 5600	Canadian Museum of Nature	Campanian-Maastrichtian	Horseshoe Canyon Formation	Canada
Albinykus baatar	Albinykus		IGM 100/3004^[17]	Mongolian Institute of Geology	Santonian-Campanian	Javkhlant Formation	Mongolia
Alectrosaurus olseni	Alectrosaurus		AMNH 6368^[18]	American Museum of Natural History	Upper Cretaceous	Iren Dabasu Formation	Mongolia
Alioramus altai	Alioramus		IGM 100/1844^[19]	Mongolian Institute of Geology	Maastrichtian	Nemegt Formation	Mongolia
Alioramus remotus	Alioramus		PIN 3141/1^[20]	Paleontological Institute, Russian Academy of Sciences	Late Cretaceous		Mongolia
Allosaurus europaeus	Allosaurus		ML415^[21]	Museu da Lourinhã	Kimmeridgian	Porto Novo Member, Lourinhã Formation	Portugal
Allosaurus fragilis	Allosaurus		YPM 1930^[22]	Peabody Museum of Natural History	Upper Jurassic	Lower Morrison Formation	Colorado
Allosaurus jimmadseni	Allosaurus		DINO 11541^[23]	Dinosaur National Monument	Kimmeridgian	Morrison Formation	United States
Almas ukhaa	Almas		IGM 100/1323^[24]	Mongolian Institute of Geology	Campanian	Djadokhta Formation	Mongolia
Alnashetri cerropoliciensis	Alnashetri		MPCA-477^[25]	Museo Provincial Carlos Ameghino	Cenomanian-Turonian	Candeleros Formation	Argentina
Altispinax dunkeri	Altispinax		NHMUK R1828^[26]	Natural History Museum, London	Berriasian-Valanginian^[27]	Hasting Beds Group	United Kingdom
Alvarezsaurus calvoi	Alvarezsaurus		MUCPv 54^[28]	Museo de Ciencias Naturales, Universidad Nacional del Comahue	Coniacian-Santonian	Bajo de la Carpa Member, Río Colorado Formation	Argentina
Alxasaurus elesitaiensis	Alxasaurus		IVPP 88402^[29]	Institute of Vertebrate Paleontology and Paleoanthropology	Albian	Bayin Gobi Formation	China
Ambopteryx longibrachium	Ambopteryx		IVPP V24192^[30]	Institute of Vertebrate Paleontology and Paleoanthropology	Oxfordian		China
Anchiornis huxleyi	Anchiornis		IVPP V14378^[31]	Institute of Vertebrate Paleontology and Paleoanthropology	Oxfordian^[32]	Tiaojishan Formation	China
Aniksosaurus darwini	Aniksosaurus		MDT-PV 1/48^[33]	Museo Desiderio Torres	Late Cretaceous	Bajo Barreal Formation	Argentina
Anomalipes zhaoi	Anomalipes		ZCDM V0020^[34]	Zhucheng Dinosaur Museum	Upper Cretaceous	Wangshi Group	China
Anserimimus planinychus	Anserimimus		IGM 100/300^[35]	Mongolian Institute of Geology	Maastrichtian^[36]	Nemegt Formation	Mongolia
Antrodemus valens	Antrodemus		USNM 218	National Museum of Natural History	Upper Jurassic	Morrison Formation	United States	Nomen dubium
Anzu wyliei	Anzu		CM 78000^[37]	Carnegie Museum of Natural History	Upper Maastrichtian	Hell Creek Formation	United States
Aoniraptor libertatem	Aoniraptor		MPCA-Pv 804/1 to 804/25^[38]	Museo Provincial “Carlos Ameghino”	Cenomanian-Turonian	Huincul Formation	Argentina
Aorun zhaoi	Aorun		IVPP V15709^[39]	Institute of Vertebrate Paleontology and Paleoanthropology	Oxfordian-Callovian	Shishugou Formation	China
Apatoraptor pennatus	Apatoraptor		TMP 1993.051.0001^[40]	Royal Tyrell Museum of Paleontology	Campanian-Maastrichtian	Horsethief Member, Horseshoe Canyon Formation	Canada
Appalachiosaurus montgomeriensis	Appalachiosaurus		RMM 6670^[41]	McWane Science Center	Campanian	Demopolis Formation	United States
Aratasaurus museunacionali	Aratasaurus		MPSC R 2089^[42]	Museu de Paleontologia, Universidade Regional do Cariri	Aptian	Romualdo Formation	Brazil
Archaeopteryx albersdoerferi	Archaeopteryx	The Daiting specimen	SNSB BSPG VN-2010/1^[43]	Bayerische Staatssammlung für Paläontologie und Geologie	Lower Tithonian	Mörnsheim Formation	Germany
Archaeopteryx lithographica	Archaeopteryx	The London specimen	NHMUK 37001^[44]	Natural History Museum, London	Lower Tithonian	Solnhofen Limestone	Germany^[45]
Archaeopteryx siemensii	Archaeopteryx	The Berlin specimen	HMN 1880/81^[46]	Museum für Naturkunde	Lower Tithonian	Solnhofen Limeston	Germany
Archaeornithoides deinosauriscus	Archaeornithoides		ZPAL MgD-II/29	Institute of Paleobiology, Polish Academy of Sciences	Late Santonian-Early Campanian	Djadokhta Formation	Mongolia
Archaeornithomimus asiaticus	Archaeornithomimus		AMNH 6565^[47]	American Museum of Natural History	Cenomanian?	Iren Dabasu Formation	China
Arcovenator escotae	Arcovenator		MHNA-PV-2011.12.1, MHNA-PV-2011.12.2, MHNA.PV.2011.12.15, MHNA.PV.2011.12.5, MHNA.PV.2011.12.4, MHNA.PV.2011.12.3^[48]	Muséum d’Histoire Naturelle d’Aix-en-Provence	Late Campanian	Argiles Rutilantes Formation	France
Aristosuchus pusilius	Aristosuchus		NHMUK R178^[49]	Natural History Museum, London	Barremian	Wessex Formation	United Kingdom
Arkansaurus fridayi	Arkansaurus		UAM 74-16^[50]	University of Arkansas Museum	Albian-Aptian	Trinity Group	United States
Asfaltovenator vialidadi	Asfaltovenator		MPEF PV 3440^[51]	Museo Paleontológico Egidio Feruglio	Late Toarcian-Bajocian	Cañadón Asfalto Formation	Argentina
Atrociraptor marshalli	Atrociraptor		TMP 95.166.1^[52]	Royal Tyrell Museum of Paleontology	Upper Campanian-Lower Maastrichtian	Horseshoe Canyon Formation	Canada
Aublysodon mirandus	Aublysodon		ANSP 9535^[53]	Academy of Natural Sciences of Philadelphia		Judith River Formation^[54]	United States	Type specimen now lost, might be a nomen dubium
Aucasaurus garridoi	Aucasaurus		MCF-PVPH-236^[55]	Museo Municipal Carmen Funes	Campanian	Anacleto Member, Río Colorado Formation	Argentina
Aurornis xui	Aurornis		YFGP-T5198	Yizhou Fossil and Geology Park	Middle-Late Jurassic	Tiaojishan Formation	China
Australovenator wintonensis	Australovenator		AODF 604^[56]	Australian Age of Dinosaurs	Albian	Winton Formation	Australia
Austrocheirus isasii	Austrocheirus		MPM-PV 10003^[57]	Museo Padre Molina	Campanian^[58]	Pari Aike Formation	Argentina
Austroraptor cabazai	Austroraptor		MML-195^[59]	Museo Municipal de Lamarque	Campanian-Maastrichtian	Allen Formation	Argentina
Aviatyrannis jurassica	Aviatyrannis		IPFUB Gui Th 1^[60]	Institut für Geologische Wissenschaften, Freie Universtät Berlin	Kimmeridgian	Alcobaça Formation	Portugal
Avimimus nemegtensis	Avimimus		MPC-D 102/81^[61]	Institute of Paleontology and Geology, Mongolian Academy of Sciences	Late Cretaceous	Nemegt Formation	Mongolia
Avimimus portentosus	Avimimus		PIN 3907-1	Paleontological Institute, Russian Academy of Sciences	Upper Cretaceous^[62]	Nemegt Formation	Mongolia
Bagaraatan ostromi	Bagaraatan		ZPAL MgD-I/108^[63]	Institute of Paleobiology, Polish Academy of Sciences	Middle Maastrichtian?	Nemegt Formation	Mongolia
Bahariasaurus ingens	Bahariasaurus		BSP 1922 X 47^[64]	Bayerische Staatssammlung für Paläontologie und historische Geologie, München	Cenomanian^[65]	Bahariya Formation	Egypt	Type specimen was destroyed in World War II
Balaur bondoc	Balaur		EME PV.313^[66]	Transylvanian Museum Society, Cluj-Napoca	Maastrichtian	Sebeş Formation	Romania
Bambiraptor feinbergi	Bambiraptor	Bambi^[67]	AMNH FR 30556^[68]	American Museum of Natural History	Upper Cretaceous	Two Medicine Formation	Montana
Banji long	Banji		IVPP V 16896^[69]	Institute of Vertebrate Paleontology and Paleoanthropology	Upper Cretaceous	Nanxiong Formation?	China
Bannykus wulatensis	Bannykus		IVPP V25026^[70]	Institute of Vertebrate Paleontology and Paleoanthropology	Aptian	Bayingobi Formation	China
Baryonyx walkeri	Baryonyx	Claws^[71]	NHMUK VP R9951^[72]	Natural History Museum, London	Barremian	Weald Clay	United Kingdom
Beibeilong sinensis	Beibeilong	Baby Louie^[73]	HGM 41HIII1219	Henan Geological Museum	Cenomanian-Turonian	Gaogou Formation	China
Beipiaosaurus inexpectus	Beipiaosaurus		IVPP V11559^[74]	Institute of Vertebrate Paleontology and Paleoanthropology	Lower Cretaceous	Yixian Formation	China
Beishanlong grandis	Beishanlong		FRDC-GS GJ (06) 01-18^[75]	Fossil Research and Development Center, Third Geology and Mineral Resources Exploration Academy, Gansu Provincial Bureau of Geo-Exploration and Mineral Development	Aptian-Albian	Xinminpu Group	China
Berberosaurus liassicus	Berberosaurus		MHNM-Pt9, Pt23, Pt22, Pt19, Pt21, Pt16, Pt20^[76]	Muséum d’Histoire Naturelle de Marrakech	Pliensbachian-Toarcian	Toundoute Continental Series	Morocco
Berthasaura leopoldinae	Berthasaura		MN 7821-V	Museu Nacional, Universidade Federal do Rio de Janeiro	Aptian-Albian	Goio Erê Formation	Brazil
Betasuchus bredai	Betasuchus		NHMUK R 42997^[77]	Natural History Museum, London	Maastrichtian^[78]	Maastricht Formation?	Netherlands
Bicentenaria argentina	Bicentenaria		MPCA 865^[79]	Museo Provincial "Carlos Ameghino"	Cenomanian	Candeleros Formation	Argentina
Bistahieversor sealeyi	Bistahieversor		NMMNH P-27469^[80]	New Mexico Museum of Natural History and Science	Campanian	Hunter Wash Member, Kirtland Formation	United States
Bonapartenykus ultimus	Bonapartenykus		MPCA 1290^[81]	Museo Provincial "Carlos Ameghino"	Campanian-Maastrichtian	Allen Formation	Argentina
Boreonykus certekorum	Boreonykus		TMP 1989.055.0047^[82]	Royal Tyrell Museum of Paleontology	Campanian	Wapiti Formation	Canada
Borogovia gracilicrus	Borogovia		ZPAL MgD-I/174^[83]	Institute of Paleobiology, Polish Academy of Sciences	Maastrichtian	Nemegt Formation	Mongolia
Bradycneme draculae	Bradycneme		BMNH A1588^[84]	Natural History Museum, London	Maastrichtian^[85]	Hațeg Basin^[86]	Romania
Buitreraptor gonzalezorum	Buitreraptor		MPCA 245^[87]	Museo Provincial "Carlos Ameghino"	Cenomanian-Turonian	Candeleros Formation	Argentina
Byronosaurus jaffei	Byronosaurus		IGM 100/983^[88]	Mongolian Institute of Geology	Late Cretaceous	Djadokhta Formation	Mongolia
Caenagnathasia martinsoni	Caenagnathasia		CCMGE 401/12457^[89]	Chernyshev's Central Museum of Geological Exploration	Turonian^[90]	Bissekty Formation	Uzbekistan
Caenagnathus collinsi	Caenagnathus		CMN 8776^[91]	Canadian Museum of Nature	Campanian	Dinosaur Park Formation	Canada
Caihong juji	Caihong		PMoL-B00175^[92]	Paleontological Museum of Liaoning	Oxfordian	Tiaojishan Formation	China
Calamosaurus foxi	Calamosaurus		BMNH R901	Natural History Museum, London	Barremian	Wealden Group^[93]	United Kingdom
Calamospondylus oweni	Calamospondylus				Upper Barremian	Vectis Formation	United Kingdom	Type specimen has been lost
Camarillasaurus cirugedae	Camarillasaurus		MPG-KPC1-46^[94]	Museo Paleontológico de Galve	Lower Barremian	Camarillas Formation	Spain
Camposaurus arizonensis	Camposaurus		UCMP 34498^[95]	University of California Museum of Paleontology	Lower-Middle Norian	Mesa Redondo Member, Chinle Formation	United States
Carcharodontosaurus iguidensis	Carcharodontosaurus		MNN IGU2^[96]	Musée National du Niger	Cenomanian	Echkar Formation	Niger
Carcharodontosaurus saharicus	Carcharodontosaurus		SGM-Din 1	Ministére de l’Energie et des Mines, Rabat	Cenomanian	Kem Kem Group	Morocco
Carnotaurus sastrei	Carnotaurus		MACN-CH 894^[97]	Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”	Maastrichtian^[98]	La Colonia Formation	Argentina
Caudipteryx dongi	Caudipteryx		IVPP V12344^[99]	Institute of Vertebrate Paleontology and Paleoanthropology	Barremian^[100]	Yixian Formation	China
Caudipteryx zoui	Caudipteryx		NGMC 97-4-A^[101]	National Geological Museum of China	Barremian	Jiulongsong Member, Chaomidianzi Formation	China
Ceratonykus oculatus	Ceratonykus		MPC 100/124^[102]	Paleontological Center, Mongolian Academy of Sciences	Santonian	Baruungoyot Formation	Mongolia
Ceratosaurus dentisulcatus	Ceratosaurus		UMNH 5278^[103]	Utah Museum of Natural History	Late Jurassic	Brushy Basin Member, Morrison Formation	United States	Possible junior synonym of Ceratosaurus nasicornis
Ceratosaurus magnicornis	Ceratosaurus		MWC 1	Museum of Western Colorado	Late Jurassic	Brushy Basin Member, Morrison Formation	United States	Possible junior synonym of Ceratosaurus nasicornis
Ceratosaurus nasicornis	Ceratosaurus		USNM 4735	National Museum of Natural History	Late Jurassic^[104]	Morrison Formation	United States
Ceratosuchops inferodios	Ceratosuchops		IWCMS 2014.95.1-3, IWCMS 2014.95.5, IWCMS 2021.30^[105]	Dinosaur Isle Museum	Barremian	Wessex Formation	United Kingdom
Changyuraptor yangi	Changyuraptor		HG B016^[106]	Paleontological Center, Bohai University	Early Cretaceous	Yixian Formation	China
Chenanisaurus barbaricus	Chenanisaurus		OCP DEK-GE 772^[107]	Office Cherifien des Phosphates	Maastrichtian	Ouled Abdoun Basin	Morocco
Chienkosaurus ceratosauroides	Chienkosaurus		IVPP V237^[108]	Institute of Vertebrate Paleontology and Paleoanthropology	Tithonian	Kuangyuan Series	China	Dubious
Chilantaisaurus tashuikouensis	Chilantaisaurus		IVPP V2884.1^[109] ^[110]	Institute of Vertebrate Paleontology and Paleoanthropology	Late Cretaceous	Ulansuhai Formation	China
Chingkankousaurus fragilis	Chingkankousaurus		IVPP V836^[111]	Institute of Vertebrate Paleontology and Paleoanthropology	Campanian - Maastrichtian	Wangshi Series	China
Chirostenotes pergracilis	Chirostenotes		CMN 2367^[112]	Canadian Museum of Nature	Upper Campanian	Dinosaur Park Formation	Canada
Chuandongocoelurus primitivus	Chuandongocoelurus		CCG 20010	Chengdu University of Geology	Middle Jurassic	Xiashaximiao Formation	China
Citipati osmolskae	Citipati		IGM 100/978^[113]	Mongolian Institute of Geology	Upper Cretaceous	Djadokhta Formation	Mongolia
Citipes elegans	Citipes		ROM 781^[114]	Royal Ontario Museum	Campanian	Dinosaur Park Formation	Canada
Coelophysis bauri	Coelophysis		AMNH 7224^[115]	American Museum of Natural History	Upper Triassic^[116]	Chinle Formation	United States	Neotype
Coeluroides largus	Coeluroides		GSI K27/562, K27/574, K27/595^[117]	Geological Survey of India	Maastrichtian	Lameta Formation	India
Coelurus fragilis	Coelurus		YPM 1991, 1993^[118]	Yale Peabody Museum	Upper Jurassic	Morrison Formation	United States	Syntypes
Compsognathus longipes	Compsognathus		SNSB-BSPG AS I 563^[119]	Bavarian State Collection for Paleontology and Geology	Tithonian	Solnhofen Limestone	Germany
Compsosuchus solus	Compsosuchus		GSI K27/578	Geological Survey of India	Maastrichtian	Lameta Formation	India
Concavenator corcovatus	Concavenator		MCCM-LH 6666^[120]	Museo de las Ciencias de Castilla-La Mancha	Upper Barremian	Calizas de La Huérguina Formation	Spain
Conchoraptor gracilis	Conchoraptor		MPC-D 100/20	Institute of Paleontology and Geology, Mongolian Academy of Sciences	Late Cretaceous	Barun Goyot Formation	Mongolia
Condorraptor currumilli	Condorraptor		MPEF-PV 1672^[121]	Museo Paleontológico Egidio Feruglio	Callovian	Cañadón Asfalto Formation	Argentina
Corythoraptor jacobsi	Corythoraptor		JPM-2015-001^[122]	Jinzhou Paleontological Museum	Campanian - Maastrichtian	Nanxiong Formation	China
Cristatusaurus lapparenti	Cristatusaurus		MNHN GDF 366^[123]	Muséum National d'Histoire Naturelle	Aptian	Elrhaz Formation^[124]	Niger
Cruxicheiros newmanorum	Cruxicheiros		WARMS G15770^[125]	Warwickshire Museum Service	Lower Bathonian	Chipping Norton Limestone	United Kingdom
Cryolophosaurus ellioti	Cryolophosaurus		FMNH PR1821^[126]	Field Museum of Natural History	Early Jurassic^[127]	Hanson Formation	Antarctica
Tyrannosaurus rex	Tyrannosaurus		CM 9380	Carnegie Museum of Natural History			United States

See also

• List of ornithopod type specimens
• List of marginocephalian type specimens
• List of thyreophoran type specimens
• List of other ornithischian type specimens
• List of Mesozoic birds

Notes and References

1. Bonaparte, J., and Fernando E. Novas. "Abelisaurus comahuensis, ng, n sp, Carnosauria from the Late Cretaceous of Patagonia." Ameghiniana 21, no. 2-4 (1985): 259-265.
2. Evans . David C. . Larson . Derek W. . Currie . Philip J. . 2013 . A new dromaeosaurid (Dinosauria: Theropoda) with Asian affinities from the latest Cretaceous of North America . Naturwissenschaften . en . 100 . 11 . 1041–1049 . 10.1007/s00114-013-1107-5 . 24248432 . 2013NW....100.1041E . 253638918 . 0028-1042.
3. Martinelli . Agustín G. . Vera . Ezequiel I. . 2007-09-12 . Achillesaurus manazzonei, a new alvarezsaurid theropod (Dinosauria) from the Late Cretaceous Bajo de la Carpa Formation, Río Negro Province, Argentina . Zootaxa . 1582 . 1 . 1–17 . 10.11646/zootaxa.1582.1.1 . 1175-5334.
4. Perle, Altangerel, Mark Norell, and James Matthew Clark. A new maniraptoran theropod, Achillobator giganticus (Dromaeosauridae), from the Upper Cretaceous of Burkhant, Mongolia. Geology and Mineralogy Chair, National University of Mongolia, 1999.
5. Currie, Philip J., and Kenneth Carpenter. "A new specimen of Acrocanthosaurus atokensis (Theropoda, Dinosauria) from the lower Cretaceous Antlers formation (lower Cretaceous, Aptian) of Oklahoma, USA." Geodiversitas 22, no. 2 (2000): 207-246.
6. Stovall . J. Willis . Langston . Wann . 1950 . Acrocanthosaurus atokensis, a New Genus and Species of Lower Cretaceous Theropoda from Oklahoma . American Midland Naturalist . 43 . 3 . 696 . 10.2307/2421859 . 2421859 . 0003-0031.
7. Turner, Alan Hamilton. Phylogenetic relationships of paravian theropods. Columbia University, 2008.
8. Chinzorig . Tsogtbaatar . Kobayashi . Yoshitsugu . Tsogtbaatar . Khishigjav . Currie . Philip J. . Watabe . Mahito . Barsbold . Rinchen . 2017 . First Ornithomimid (Theropoda, Ornithomimosauria) from the Upper Cretaceous Djadokhta Formation of Tögrögiin Shiree, Mongolia . Scientific Reports . en . 7 . 1 . 5835 . 10.1038/s41598-017-05272-6 . 2045-2322 . 5517598 . 28724887.
9. Sereno . Paul C. . Martinez . Ricardo N. . Wilson . Jeffrey A. . Varricchio . David J. . Alcober . Oscar A. . Larsson . Hans C. E. . 2008-09-30 . Kemp . Tom . Evidence for Avian Intrathoracic Air Sacs in a New Predatory Dinosaur from Argentina . PLOS ONE . en . 3 . 9 . e3303 . 10.1371/journal.pone.0003303 . 1932-6203 . 2553519 . 18825273. free . 2008PLoSO...3.3303S .
10. Sereno . Paul C. . 2017 . Early Cretaceous Ornithomimosaurs (Dinosauria: Coelurosauria) from Africa . Ameghiniana . en . 54 . 5 . 576–616 . 10.5710/AMGH.23.10.2017.3155 . 134718338 . 0002-7014.
11. Sereno, Paul C., Jeffrey A. Wilson, Hans CE Larsson, Didier B. Dutheil, and Hans-Dieter Sues. "Early Cretaceous dinosaurs from the Sahara." Science 266, no. 5183 (1994): 267-271.
12. Rauhut . Oliver W.M. . López-Arbarello . Adriana . 2009 . Considerations on the age of the Tiouaren Formation (Iullemmeden Basin, Niger, Africa): Implications for Gondwanan Mesozoic terrestrial vertebrate faunas . Palaeogeography, Palaeoclimatology, Palaeoecology . en . 271 . 3–4 . 259–267 . 10.1016/j.palaeo.2008.10.019. 2009PPP...271..259R .
13. Evans . David C. . Cullen . Thomas M. . Larson . Derek W. . Rego . Adam . 2017 . A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada . Canadian Journal of Earth Sciences . en . 54 . 8 . 813–826 . 10.1139/cjes-2017-0034 . 2017CaJES..54..813E . 0008-4077.
14. Longrich . Nicholas R. . Currie . Philip J. . 2009 . Albertonykus borealis, a new alvarezsaur (Dinosauria: Theropoda) from the Early Maastrichtian of Alberta, Canada: implications for the systematics and ecology of the Alvarezsauridae . Cretaceous Research . en . 30 . 1 . 239–252 . 10.1016/j.cretres.2008.07.005. 2009CrRes..30..239L .
15. Book: A., Russell, Dale . Tyrannosaurs from the late cretaceous of Western Canada. . 1970 . Queen's Printer . 421445857.
16. Tanke . Darren H. . Currie . Philip J. . 2010 . Sues . Hans-Dieter . A history of Albertosaurus discoveries in Alberta, Canada . Canadian Journal of Earth Sciences . en . 47 . 9 . 1197–1211 . 10.1139/E10-057 . 2010CaJES..47.1197T . 0008-4077.
17. Nesbitt . Sterling J. . Clarke . Julia A. . Turner . Alan H. . Norell . Mark A. . 2011-02-10 . A small alvarezsaurid from the eastern Gobi Desert offers insight into evolutionary patterns in the Alvarezsauroidea . Journal of Vertebrate Paleontology . en . 31 . 1 . 144–153 . 10.1080/02724634.2011.540053 . 2011JVPal..31..144N . 85283009 . 0272-4634.
18. Gilmore, Charles Whitney. "On the dinosaurian fauna of the Iren Dabasu Formation. Bulletin of the AMNH; v. 67, article 2." (1933).
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