List of mammals of Andorra explained

This is a list of the mammal species recorded in Andorra. There are seventy-nine mammal species in Andorra, of which three are vulnerable, ten are near threatened, and two are endangered.^[1]

The following tags are used to highlight each species' conservation status as assessed by the International Union for Conservation of Nature:

EX	Extinct	No reasonable doubt that the last individual has died.
EW	Extinct in the wild	Known only to survive in captivity or as a naturalized populations well outside its previous range.
CR	Critically endangered	The species is in imminent risk of extinction in the wild.
EN	Endangered	The species is facing an extremely high risk of extinction in the wild.
VU	Vulnerable	The species is facing a high risk of extinction in the wild.
NT	Near threatened	The species does not meet any of the criteria that would categorise it as risking extinction but it is likely to do so in the future.
LC	Least concern	There are no current identifiable risks to the species.
DD	Data deficient	There is inadequate information to make an assessment of the risks to this species.

Order: Artiodactyla (even-toed ungulates)

The even-toed ungulates are ungulates whose weight is borne about equally by the third and fourth toes, rather than mostly or entirely by the third as in perissodactyls. There are about 360 artiodactyl species, including many that are of great economic importance to humans. This group also includes cetaceans.

• Family: Suidae (pigs)
  • Subfamily: Suinae
    • Genus: Sus
      • Wild boar, S. scrofa ^[2]
• Family: Cervidae (deer)
  • Subfamily: Cervinae
    • Genus: Cervus
      • Red deer, C. elaphus^{[3] [4]}
        • European red deer, C. e. elaphus
    • Genus: Dama
      • Fallow deer, D. dama^{[5] [6]} introduced
  • Subfamily: Capreolinae
    • Genus: Capreolus
      • Western roe deer, C. capreolus ^[7]
        • European roe deer, C. c. capreolus
• Family: Bovidae (bovids)
  • Subfamily: Bovinae
    • Genus: Bos
      • Domestic cattle, B. taurus^[8] (introduced)
    • Genus: Bison
      • European bison, B. bonasus ^{[9] [10]}
  • Subfamily: Antilopinae
    • Genus: Capra
      • Iberian Ibex, Capra pyrenaica ^[11]
        • Pyrenean ibex, C. p. pyrenaica
        • Western Spanish ibex, C. p. victoriae (introduced)
    • Genus: Ovis
      • Domestic sheep, O. aries^{[12] [13]}
      • Mouflon, O. gmelini^{[14] [15]} (introduced)
    • Genus: Rupicapra
      • Pyrenean chamois, R. pyrenaica ^[16]
        • Pyrenean chamois, R. p. pyrenaica

Order: Perissodactyla (odd-toed ungulates)

The odd-toed ungulates are browsing and grazing mammals. They are usually large to very large, and have relatively simple stomachs and a large middle toe. There are only around 20 species of perrissodactyl.

• Family: Equidae (horses & allies)
  • Genus: Equus
    • Wild horse, E. ferus^[17]
      • Domestic horse, E. f. caballus introduced
      • Wild horse, E. f. ferus
    • Donkey, E. asinus^[18] introduced

Order: Carnivora (carnivorans)

There are over 310 species of carnivorans, the majority of which feed primarily on meat. They have a characteristic skull shape and dentition.

• Suborder: Feliformia
  • Family: Felidae (cats)
    • Subfamily: Felinae
      • Genus: Felis
        • European wildcat, F. silvestris ^[19]
  • Family: Viverridae
    • Subfamily: Viverrinae
      • Genus: Genetta
        • Common genet, G. genetta introduced^[20]
• Suborder: Caniformia
  • Family: Canidae (dogs, foxes)
    • Genus: Canis
      • Golden jackal, C. aureus vagrant^[21]
      • Gray wolf, C. lupus ^[22]
        • Eurasian wolf, C. l. lupus
    • Genus: Vulpes
      • Red fox, V. vulpes ^[23]
  • Family: Mustelidae (weasels, stoats, & allies)
    • Subfamily: Guloninae
      • Genus: Martes
        • European pine marten, M. martes^{[24] [25]}
        • Stone marten, M. foina^{[26] [27]}
    • Subfamily: Lutrinae
      • Genus: Lutra
        • European otter, L. lutra ^[28]
    • Subfamily: Melinae
      • Genus: Meles
        • European badger, M. meles ^[29]
    • Subfamily: Mustelinae
      • Genus: Mustela
        • Stoat, M. erminea ^[30]
        • Least weasel, M. nivalis ^[31]
        • European polecat, M. putorius ^[32]
  • Family: Ursidae (bears)
    • Genus: Ursus
      • Brown bear, U. arctos ^[33]
        • Eurasian brown bear, U. a. arctos

Order: Chiroptera (bats)

The bats' most distinguishing feature is that their forelimbs are developed as wings, making them the only mammals capable of flight. Bat species account for about 20% of all mammals, with almost 1,500 species described.

• Family: Molossidae
  • Genus: Tadarida
    • European free-tailed bat, T. teniotis ^[34]
• Family: Rhinolophidae
  • Subfamily: Rhinolophinae
    • Genus: Rhinolophus
      • Mediterranean horseshoe bat, R. euryale ^[35]
      • Lesser horseshoe bat, R. hipposideros ^[36]
      • Greater horseshoe bat, R. ferrumequinum ^[37]
• Family: Vespertilionidae
  • Subfamily: Myotinae
    • Genus: Myotis
      • Bechstein's bat, M. bechsteinii ^[38]
      • Lesser mouse-eared bat, M. blythii ^[39]
      • Long-fingered bat, M. capaccinii ^[40]
      • Cryptic myotis, M. crypticus ^[41]
      • Daubenton's bat, M. daubentonii ^[42]
      • Geoffroy's bat, M. emarginatus ^[43]
      • Escalera's bat, M. escalerai ^[44]
      • Greater mouse-eared bat, M. myotis ^[45]
      • Whiskered bat, M. mystacinus ^[46]
  • Subfamily: Vespertilioninae
    • Genus: Barbastella
      • Western barbastelle, B. barbastellus ^[47]
    • Genus: Eptesicus
      • Serotine bat, E. serotinus ^[48]
    • Genus: Hypsugo
      • Savi's pipistrelle, H. savii ^[49]
    • Genus: Nyctalus
      • Greater noctule, N. lasiopterus ^[50]
      • Lesser noctule, N. leisleri ^[51]
      • Common noctule, N. noctula ^[52]
    • Genus: Pipistrellus
      • Kuhl's Pipistrelle, P. kuhlii ^[53]
      • Common pipistrelle, P. pipistrellus ^[54]
      • Kuhl's Pipistrelle, P. kuhlii ^[55]
    • Genus: Plecotus
      • Brown long-eared bat, P. auritus ^[56]
      • Grey long-eared bat, P. austriacus ^[57]
      • Alpine long-eared bat, P. macrobullaris ^[58]

Order: Lagomorpha (hares, rabbits, & pikas)

Moderately small herbivores related to rodents, but with long ears, large hindlegs, and four incisors rather than two on the top jaw. They move primarily hopping and bounding. There are over 100 species of lagomorph.

• Family: Leporidae
  • Genus: Lepus
    • European hare, L. europaeus ^[59]
  • Genus: Oryctolagus
    • European rabbit, O. cuniculus ^[60]

Order: Rodentia (rodents)

Rodents make up the largest order of mammals, with over 40 percent of mammalian species, with almost 2,700 currently described species. They have two incisors in the upper and lower jaw which grow continually and must be kept short by gnawing.

• Suborder: Myomorpha
  • Family: Cricetidae (hamsters, voles, & allies)
    • Subfamily: Arvicolinae
      • Genus: Arvicola
        • European water vole, A. amphibius^{[61] [62]} (sometimes split into Arvicola scherman}^[63])
        • Southwestern water vole, A. sapidus^{[64] [65]}
      • Genus: Chionomys
        • European snow vole, C. nivalis^{[66] [67]}
      • Genus: Clethrionomys
        • Bank vole, C. glareolus^{[68] [69]}
      • Genus: Microtus
        • Field vole, M. agrestis ^[70]
        • Common vole, M. arvalis ^[71]
        • Mediterranean pine vole, M. duodecimcostatus ^[72]
        • Pyrenean pine vole, M. pyrenaicus ^[73]
  • Family: Muridae (Old World rats, Old World mice, & allies.)
    • Subfamily: Murinae
      • Genus: Apodemus
        • Yellow-necked mouse, A. flavicollis ^[74]
        • Wood mouse, A. sylvaticus ^[75]
      • Genus: Mus
        • House mouse, M. musculus ^[76]
      • Genus: Rattus
        • Brown rat, R. norvegicus^{[77] [78]}
        • Black rat, R. rattus^{[79] [80]}
• Suborder: Sciuromorpha
  • Family: Gliridae (dormice)
    • Subfamily: Glirinae
      • Genus: Glis
        • European edible dormouse, G. glis^{[81] [82]}
    • Subfamily: Leithiinae
      • Genus: Eliomys
        • Garden dormouse, E. quercinus ^[83]
  • Family: Sciuridae
    • Subfamily: Sciurinae
      • Genus: Sciurus
        • Eurasian red squirrel, S. vulgaris^{[84] [85]}
    • Subfamily: Xerinae
      • Genus: Marmota
        • Alpine marmot, M. marmota ^[86] introduced

Order: Eulipotyphla (shrews, moles, and solenodons)

These are insectivorous mammals. The shrews and solenodons closely resemble mice while the moles are stout-bodied burrowers. There are almost 600 species of eulipotyphlans.

• Family: Soricidae (shrews)
  • Subfamily: Crocidurinae
    • Genus: Crocidura
      • Greater white-toothed shrew, C. russula^{[87] [88]}
    • Genus Suncus
      • Pygmy white-toothed shrew, S. etruscus ^[89]
  • Subfamily: Soricinae
    • Genus: Sorex
      • Common shrew, S. araneus^{[90] [91]}
      • Crowned shrew, S. coronatus ^[92]
      • Eurasian pygmy shrew, S. minutus^{[93] [94]}
    • Genus: Neomys
      • Southern water shrew, N. anomalus^{[95] [96]}
      • Eurasian water shrew, N. fodiens^{[97] [98]}
• Family: Talpidae (moles)
  • Subfamily: Talpinae
    • Genus: Talpa
      • European mole, T. europaea ^[99]
    • Genus: Galemys
      • Pyrenean desman, G. pyrenaicus ^[100]

See also

• List of chordate orders
• List of prehistoric mammals
• Mammal classification
• List of mammals described in the 2000s

External links

• Web site: Mammal Species of the World. Smithsonian National Museum of Natural History. 2005. 22 May 2007. https://web.archive.org/web/20070427043030/http://nmnhgoph.si.edu/msw/ . 27 April 2007.
• Web site: Animal Diversity Web. University of Michigan Museum of Zoology. 1995–2006. 22 May 2007.

Notes and References

1. This list is derived from the IUCN Red List which lists species of mammals and includes those mammals that have recently been classified as extinct (since 1500 AD). The taxonomy and naming of the individual species is based on those used in existing Wikipedia articles as of 15 August 2024 and supplemented by the common names and taxonomy from the IUCN and GBIF where no Wikipedia article was available.
2. Sus scrofa. Keling, O.. Leus, K.. 2019. amp. e.T41775A44141833. 10.2305/IUCN.UK.2019-3.RLTS.T41775A44141833.en.
3. 2440958. Cervus elaphus Linnaeus, 1758. 15 August 2024.
4. Cervus elaphus . Lovari, S. . Lorenzini, R. . Masseti, M. . Pereladova, O. . Carden, R.F. . Brook, S.M. . Mattioli, S. . amp . e.T55997072A142404453. 2018. }
5. 5220136. Dama dama (Linnaeus, 1758). 15 August 2024.
6. Dama dama . Masset, M.. Mertzanidou, D.. amp . e.T42188A10656554. 2008. 10.2305/IUCN.UK.2008.RLTS.T42188A10656554.en.
7. Capreolus capreolus . Lovari, S.. Herrero, J.. Masseti, M.. Ambarli, H.. Lorenzini, R.. Giannatos, G.. amp . 2016. e.T42395A22161386. 10.2305/IUCN.UK.2016-1.RLTS.T42395A22161386.en.
8. 2441022. Bos taurus Linnaeus, 1758. 15 August 2024.
9. Web site: Biggest bison transport ever Rewilding Europe. OKIA. 9 October 2012.
10. News: Segovia recibe nueve bisontes europeos para evitar su extinción . 24 September 2019 . 20 Minutos . 20 April 2018.
11. Capra pyrenaica . Herrero, J. . Acevedo, P. . Arnal, M. C. . Fernández de Luco, D.. Fonseca, C. . García-González, R.. Pérez, J. M.. Sourp, E. . amp . 2021. 2020 . e.T3798A195855497. 10.2305/IUCN.UK.2021-1.RLTS.T3798A195855497.en.
12. 2441110. Ovis aries Linnaeus, 1758. 15 August 2024.
13. Ovis aries . Aulagnier, S.. Giannatos, G.. Lovari, S.. amp . 2007 . e.T136720A4331580.
14. 9542814. Ovis gmelini Linnaeus, 1758. 15 August 2024.
15. Ovis gmelini . Michel, S.. Ghoddousi, A.. amp . 2020. e.T54940218A22147055. 10.2305/IUCN.UK.2020-2.RLTS.T54940218A22147055.en.
16. Rupicapra pyrenaica . Herrero, J. . Lovari, S. . Nores, C. . Toigo, C. . amp . 2022. 2020 . e.T19771A217715455. 10.2305/IUCN.UK.2022-1.RLTS.T19771A217715455.en.
17. 2440886. Equus caballus Linnaeus, 1758. 15 August 2024.
18. 2440891. Equus asinus Linnaeus, 1758. 15 August 2024.
19. Felis silvestris . Gerngross, P. . Ambarli, H. . Angelici, F.M. . Anile, S. . Campbell, R. . Ferreras de Andres, P. . Gil-Sanchez, J.M. . Götz, M. . Jerosch, S. . Mengüllüoglu, D. . Monterosso, P. . Zlatanova, D. . amp . e.T181049859A224982454. 10.2305/IUCN.UK.2023-1.RLTS.T181049859A224982454.en. 2023. 2022.
20. Genetta genetta . Gaubert, P. . Carvalho, F. . Camps, D. . Do Linh San, E. . amp . 2015 . e.T41698A45218636. 10.2305/IUCN.UK.2015-4.RLTS.T41698A45218636.en.
21. Canis aureus . Hoffmann, M. . Arnold, J. . Duckworth, J. W. . Jhala, Y. . Kamler, J. F. . Krofel, M. . amp . 2018 . e.T118264161A46194820.
22. Canis lupus . Boitani, L. . Phillips, M. . . amp . 2018 . e.T3746A119623865.
23. Vulpes vulpes . Hoffmann, M. . Sillero-Zubiri, C. . 2021 . e.T23062A193903628. 10.2305/IUCN.UK.2021-1.RLTS.T23062A193903628.en.
24. 5218878. Martes martes (Linnaeus, 1758). 15 August 2024.
25. Martes martes . Herrero, J.. Kranz, A.. Skumatov, D.. Abramov, A. V.. Maran, T.. Monakhov, V. G. . amp . e.T12848A45199169. 2016. 10.2305/IUCN.UK.2016-1.RLTS.T12848A45199169.en.
26. 5218887. Martes foina (Erxleben, 1777). 15 August 2024.
27. Martes foina . Abramov, A. V.. Kranz, A.. Herrero, J.. Choudhury, A.. Maran, T. . amp . e.T29672A45202514. 2016. 10.2305/IUCN.UK.2016-1.RLTS.T29672A45202514.en.
28. Lutra lutra . Loy, A.. Kranz, A.. Oleynikov, A.. Roos, A.. Savage, M.. Duplaix, N. . amp . 2022. 2021. e.T12419A218069689. 10.2305/IUCN.UK.2022-2.RLTS.T12419A218069689.en.
29. Meles meles . Kranz, A. . Abramov, A. V. . Herrero, J. . Maran, T. . amp . 2016 . 2016 . e.T29673A45203002 .
30. Mustela erminea . Reid, F.. Helgen, K.. Kranz, A.. 2016 . e.T29674A45203335. 10.2305/IUCN.UK.2016-1.RLTS.T29674A45203335.en.
31. Mustela nivalis . McDonald, R.A.. Abramov, A. V.. Stubbe, M.. Herrero, J.. Maran, T.. Tikhonov, A.. Cavallini, P.. Kranz, A.. Giannatos, G.. Kryštufek, B.. Reid, F. . 2019. 2016 . e.T70207409A147993366. 10.2305/IUCN.UK.2016-1.RLTS.T70207409A147993366.en.
32. Mustela nivalis . Skumatov, D.. Abramov, A. V.. Herrero, J.. Kitchener, A.. Maran, T.. Kranz, A.. Sándor, A.. Saveljev, A.. Savourè-Soubelet, A.. Guinot-Ghestem, M.. Zuberogoitia, I.. Birks, J. D. S.. Weber, A.. Melisch, R.. Ruette, S.. 2016 . e.T41658A45214384. 10.2305/IUCN.UK.2016-1.RLTS.T41658A45214384.en.
33. Ursus arctos . McLellan, B. N. . Proctor, M. F. . Huber, D. . Michel, S. . amp . 2017 . e.T41688A121229971 . 2017 . 10.2305/IUCN.UK.2017-3.RLTS.T41688A121229971.en.
34. Tadarida teniotis . amp . Benda, P. . Piraccini, R.. 2016. e.T21311A22114995. 10.2305/IUCN.UK.2016-2.RLTS.T21311A22114995.en.
35. Rhinolophus euryale . amp . Juste, J. . Alcaldé, J. . 2016. e.T19516A21971185. 10.2305/IUCN.UK.2016-2.RLTS.T19516A21971185.en.
36. Rhinolophus hipposideros . amp . Taylor, P. . 2016. e.T19518A21972794. 10.2305/IUCN.UK.2016-2.RLTS.T19518A21972794.en.
37. Rhinolophus ferrumequinum . amp . Piraccini, R. . 2016. e.T19517A21973253. 10.2305/IUCN.UK.2016-2.RLTS.T19517A21973253.en.
38. Myotis bechsteinii . amp . Paunović, M. . 2016. e.T14123A22053752. 10.2305/IUCN.UK.2016-2.RLTS.T14123A22053752.en.
39. Myotis blythii . amp . Juste, J. Paunović, M. . 2016. e.T14124A22053297. 10.2305/IUCN.UK.2016-2.RLTS.T14124A22053297.en.
40. Myotis capaccinii . amp. Paunović, M. . 2016. e.T14124A22053297. 10.2305/IUCN.UK.2016-2.RLTS.T14124A22053297.en.
41. Myotis crypticus . amp . Russo, D. . Cistrone, L. . 2023. 2024 . e.T215154989A254355251.
42. Myotis daubentonii . amp . Kruskop, S. V. . Godlevska, L.. Bücs, S.. Çoraman, E.. Gazaryan, S.. 2020. 2021 . e.T85342710A195858793. 10.2305/IUCN.UK.2020-2.RLTS.T85342710A195858793.en.
43. Myotis emarginatus . amp . Piraccini, R. . 2016 . e.T14129A22051191. 10.2305/IUCN.UK.2016-2.RLTS.T14129A22051191.en.
44. Myotis escalerai . amp . Russo, D.. Cistrone, L. . 2023. e.T85733126A211003991. 10.2305/IUCN.UK.2023-1.RLTS.T85733126A211003991.en.
45. Myotis myotis . amp . Coroiu, I. . Juste, J. . Paunović, M. . 2016 . e.T14133A22051759. 10.2305/IUCN.UK.2016-2.RLTS.T14133A22051759.en.
46. Myotis mystacinus . amp . Coroiu, I. . 2016 . e.T14134A22052250. 10.2305/IUCN.UK.2016-2.RLTS.T14134A22052250.en.
47. Barbastella barbastellus . amp . Piraccini, R.. 2016. e.T2553A22029285. 10.2305/IUCN.UK.2016-2.RLTS.T2553A22029285.en.
48. Eptesicus serotinus . amp . Godlevska, L. . Kruskop, S. V. Gazaryan, S. . 2021. 2020. e.T85199559A195834153.
49. Hypsugo savii . amp . Juste, J.. Paunović, M.. 2016 . e.T44856A22072380.
50. Nyctalus lasiopterus . amp . Alcadé, J.. Juste, J.. Paunović, M.. 2016 . e.T14918A22015318. 10.2305/IUCN.UK.2016-2.RLTS.T14918A22015318.en.
51. Nyctalus leisleri . amp . Juste, J. . Paunović, M. . 2016 . e.T14919A22016159. 10.2305/IUCN.UK.2016-2.RLTS.T14919A22016159.en.
52. Nyctalus noctula . amp . Csorba, G.. Hutson, A. M. . 2016 . e.T14920A22015682. 10.2305/IUCN.UK.2016-2.RLTS.T14920A22015682.en.
53. Pipistrellus kuhlii . amp . Juste, J. . Paunović, M. . 2016 . e.T17314A22132946. 10.2305/IUCN.UK.2016-2.RLTS.T17314A22132946.en.
54. Pipistrellus pipistrellus . amp . Godlevska, L.. Bücs, S. Kruskop, S. V.. Gazaryan, S.. Benda, P.. Paunović, M.. 2020. 2021. e.T85333513A196581936. 10.2305/IUCN.UK.2020-2.RLTS.T85333513A196581936.en.
55. Pipistrellus kuhlii . amp . Juste, J. . Paunović, M. . 2016 . e.T17314A22132946. 10.2305/IUCN.UK.2016-2.RLTS.T17314A22132946.en.
56. Plecotus auritus . amp . Russo, D. . Cistrone, L. . 2023 . e.T85535522A211015413. 10.2305/IUCN.UK.2023-1.RLTS.T85535522A211015413.en.
57. Plecotus austriacus . amp . Russo, D. . Cistrone, L. . 2023 . e.T85533333A211018209. 10.2305/IUCN.UK.2023-1.RLTS.T85533333A211018209.en.
58. Plecotus macrobullaris . amp . Piraccini, R. . 2016. e.T136229A22002229. 10.2305/IUCN.UK.2016-2.RLTS.T136229A22002229.en.
59. Lepus europaeus . Hacklander, K. . Schai-Braun, S. . 2019 . 2019 . e.T41280A45187424 . 10.2305/IUCN.UK.2019-1.RLTS.T41280A45187424.en.
60. Oryctolagus cuniculus . 2019. 2020. Villafuerte, R.. Delibes-Mateos, M.. e.T41291A170619657. 10.2305/IUCN.UK.2019-3.RLTS.T41291A170619657.en.
61. 4265185. Arvicola amphibius . 15 August 2024.
62. Arvicola amphibius . 2016. Gazzard, A.. e.T219451656A219451755. 10.2305/IUCN.UK.2023-1.RLTS.T219451656A219451755.en.
63. Arvicola scherman . 2016. Cassola, F.. e.T136766A22358636. 10.2305/IUCN.UK.2016-3.RLTS.T136766A22358636.en.
64. 2438492. Arvicola sapidus Miller, 1908. 15 August 2024.
65. Arvicola amphibius . 2008. Rigaux, P.. Vaslin, M.. Noblet, J. F.. Amori, G.. Palomo, L. J.. e.T2150A9290712. 10.2305/IUCN.UK.2008.RLTS.T2150A9290712.en.
66. 2438162. Chionomys nivalis (Martins, 1842). 15 August 2024.
67. Chionomys nivalis . 2016. 2017 . Kryštufek, B.. e.T4659A115069366. 10.2305/IUCN.UK.2016-3.RLTS.T4659A22379147.en.
68. 2439140. Clethrionomys glareolus (Schreber, 1780). 15 August 2024.
69. Myodes glareolus . 2021. 2016 . Hutterer, R.. Kryštufek, B.. Yigit, N.. Mitsainas, G.. Palomo, L.. Henttonen, H.. Vohralík, V.. Zagorodnyuk, I.. Juškaitis, R.. Meinig, H.. Bertolino, S.. e.T4973A197520967. 10.2305/IUCN.UK.2021-1.RLTS.T4973A197520967.en.
70. Microtus agrestis . Kryštufek, B. . Vohralík, V.. Zima, J. . Zagorodnyuk, I.. 2016 . 2017. e.T13426A115112050. 10.2305/IUCN.UK.2016-3.RLTS.T13426A22349665.en.
71. Microtus arvalis . Yigit, N.. Hutterer, R.. Kryštufek, B.. Amori, G.. 2016 . e.T13488A22351133. 10.2305/IUCN.UK.2016-2.RLTS.T13488A22351133.en.
72. Microtus duodecimcostatus . Aulagnier, S.. 2016 . e.T13493A513875. 10.2305/IUCN.UK.2016-3.RLTS.T13493A513875.en.
73. Microtus pyrenaicus . Cassola, F.. 2016 . e.T39315A2896811. 10.2305/IUCN.UK.2016-2.RLTS.T39315A2896811.en.
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