List of largest reptiles explained
This list of largest reptiles takes into consideration both body length and mass of large reptile species, including average ranges and maximum records. The crocodilians reaching a length of 4m (13feet) and a mass of 500kg (1,100lb) or more. It is worth mentioning that unlike the upper weight of mammals, birds or fish, mass in reptiles is frequently poorly documented, thus subject to conjecture and estimation.
The saltwater crocodile is considered to be the largest extant reptile, verified at up to 6.32m (20.73feet) in length and around 1000kg-1500kgkg (2,000lb-3,300lbkg) in mass.[1] Larger specimens have been reported albeit not fully verified,[2] the maximum of which is purportedly 7m (23feet) long with an estimated mass of 2000kg (4,000lb).[3]
The following table below lists the 15 largest extant reptile species ranked according to their average mass range, with maximum reported/reliable/estimated mass also being provided.
Overall
Lizards and snakes (Squamata)
- The most massive living member of this highly diverse reptilian order is the green anaconda (Eunectes murinus) of the neotropical riverways. These may exceed and, although such reports are not fully verified.[54] Rumors of larger anacondas also persist.[55] The reticulated python (Malayopython reticulatus) of Southeast Asia is longer but more slender, and has been reported to measure as much as in length and weigh up to .[3] [56] [57] The Burmese python, a south-east Asian species is known to reach up to and weigh as much as and is generally among the three heaviest species of snakes.[58] Several other species of python can reach or exceed in length and in weight. The fossil of the largest snake ever, the extinct boa Titanoboa were found in coal mines in Colombia. This snake was estimated to reach a length of and weighed about .[59]
- The longest venomous snake is the king cobra (Ophiophagus hannah), with lengths (recorded in captivity) of up to and a weight of up to .[3] It is also the largest elapid. The second-longest venomous snake in the world is possibly the African black mamba (Dendroaspis polylepis), which can grow up to . Among the genus Naja, the longest member arguably may be the forest cobra (Naja melanoleuca), which can reportedly grow up to . The King brown snake, reaching lengths of up to and weights of or more, is the largest venomous snake in Australia.[64] The Yellow sea snake (Hydrophis spiralis) is the largest of the sea snakes growing up to a length of . Few other elapids can reach or exceed in length and in weight.[65] [66]
- The Gaboon viper, a very bulky species with a maximum length of around, is typically the heaviest non-constrictor snake and the biggest member of the viper family, with unverified specimens reported to as much as .[3] [67] The Eastern diamondback rattlesnake is nearly as large, with the maximum length being and maximum weight being . The rattlesnake genus Crotalus, which includes the aforementioned eastern diamondback rattlesnake and western diamondback rattlesnake (Crotalus atrox), reaches a maximum length of, and according to W. A. King one large specimen had a length of and a mass of .[68] The third largest rattlesnake is the Mexican west coast rattlesnake (Crotalus basiliscus), which reaches long and mass,[68] and one captive-raised male was weighed at in 2020.[69] While not quite as heavy, other members of the viper family are longer still, the South American bushmaster (Lachesis muta) and Central American bushmaster (Lachesis stenophrys), with a maximum length in the range of, with the former being considered as the third-longest venomous snake in the world.[70]
- The largest of the monitor lizards (and the largest extant lizard in genera) is the Komodo dragon (Varanus komodoensis), endemic to the island of its name, at a maximum size of long and, although this is currently the only record that places the mass above .[3] Crocodile monitor (Varanus salvadorii) is probably the longest living lizard, known to grow as much as, with reported lengths of up to and weights of up to .[71] [72] [73] The Asian water monitor is also one of the largest lizards in the world, with sizes of up to and reported weights of up to .[74] Few other species (such as perentie and nile monitor) can reach or possibly exceed in length and in weight. The prehistoric Australian megalania (Varanus priscus), which may have existed up to 40,000 years ago, is the largest terrestrial lizard known to exist, but the lack of a complete skeleton has resulted in a wide range of size estimates. Molnar's 2004 assessment resulted in an average weight of and length of, and a maximum of at in length, which is toward the high end of the early estimates.[75]
- Iguanas are very large lizards, some of which can reach lengths of up to 2 m (6.6 ft) and weigh more than 10kg (20lb). They are the largest lizards after some large species of monitor lizards, and the largest lizards in the New World. Iguanas vary considerably in size and form, but even the smallest lizards in this family are still quite large. Many sources describe the green iguana (Iguana iguana) as the largest iguanid, often reaching lengths up to 1.5 metres (4.91 ft) and masses of 4kg (09lb),[76] and with a maximum length of 2 m (6.6 ft) and a mass of 8kg (18lb)[77] and in some cases even 9.1kg (20.1lb). However, the heaviest species in this family is the blue iguana (Cyclura lewisi), with a total length of up to 1.5 m (4.91 ft), a SVL of 51–76 cm (30 in) and a mass of up to 14kg (31lb)[77] [78] It is the eighth-heaviest and largest extant lizard.[79] Other large species in this family include the Galapagos land iguana (Conolophus subcristataus), with a length of about 1.5 m (4.91 ft) and a mass of up to 13kg (29lb). It is the second-heaviest iguanid after the blue iguana and the ninth-heaviest and largest lizard in the world. Another large species from the same genus is the Santa Fe land iguana (Conolophus pallidus), reaching a SVL of 56.4cm (22.2inches) and a mass of 7.2kg (15.9lb). The Galapagos pink land iguana (Conolophus marthae) have snout-vent length 57.5cm (22.6inches) and the mass of 8kg (18lb).[80] The marine iguana (Amblyrhynchus cristatus) is also among the largest iguanas in the world, and the largest reptile on Galapagos Islands after the Galapagos land iguana, not including turtles reaching a maximum total length of 1.4 m (4.59 ft), a SVL of from 12 till 56 cm (from 4.72 till 22 in)[81] [82] and a mass of from 1to[83] depending on islands.
- The largest extant gecko is the New Caledonian giant gecko (Rhacodactylus leachianus) of New Caledonia, which can grow to 14 inches in length.[84] It was surpassed in size by the extinct Kawekaweau (Hoplodactylus delcourti) of New Zealand, which grew to a length of .[85]
Tuataras (Sphenodontia)
The larger of the two extant species of the New Zealand native tuataras is the Brothers Island tuatara (Sphenodon guntheri). The maximum size is 1.4kg (03.1lb) and 76cm (30inches).
Ichthyosaurs (Ichthyosauria)
Some of these marine reptiles were comparable in size to modern cetaceans. Until 2024, the largest ichthyosaur was believed to be the Late Triassic species Shastasaurus sikanniensis, at approximately 21m (69feet) long[87] [88] and in weight, from the Norian stage in what is now British Columbia.[89] However, in 2018, a specimen from the Rhaetian stage of Lilstock was discovered to be 25 percent larger, approximately in length.[90] [91] The specimen was referred in 2024 to Ichthyotitan severnensis, with revised estimates of 25m (82feet) making it the largest known marine reptile.[92] Shonisaurus popularis is another enormous ichthyosaur. It reached in length and in weight.[88] Cymbospondylus youngorum is the largest Middle Triassic ichthyosaur, with in length and in weight.[93] In 2022 the tooth of a giant predatory ichthyosaur (unofficially named Swiss Tyrant) was discovered in the Swiss Alps. Based on crown diameter, the animal was about 18 m (59 ft) long and weighed 55 t (121,000 lb), making it one of the largest marine reptiles ever.[94]
Pantestudines
Turtles and tortoises (Testudines)
- The largest extant turtle is the leatherback sea turtle (Dermochelys coriacea), reaching a maximum total length of and a weight of .[3] [95] The second-largest extant testudine is the Loggerhead sea turtle. It tends to weight slightly more average weight than the green sea turtle, and reaches more massive top sizes. The Loggerhead sea turtle (Caretta caretta) reaches a maximum size of and weight of, while the Green sea turtle (Chelonia mydas) reaches a maximum weight in the range of .[47] The Flatback sea turtle (Natator depressus) may reach a weight of up to .[96] Other species of Sea turtles are small-medium in size, but are still considered as large-sized for a typical turtle.
- The largest extant freshwater turtle is possibly the North American alligator snapping turtle (Macrochelys temminckii), which has an unverified maximum reported weight of, although this is challenged by several rare, giant softshell turtle from Asia (Rafetus and Pelochelys) unverified to and nearly in total length.[3] [97] [98] The Yangtze giant softshell turtle (Rafetus swinhoei), a critically endangered species of softshell turtle, is sometimes considered as the largest extant freshwater turtle. The heaviest recorded specimen was reported to weigh . Some researchers have claimed that a related species, the Asian giant softshell turtle (Pelochelys cantorii), is the largest living species of freshwater turtle. However, such claims are fraught with problems, because it now seems likely that this species is actually a composite, i.e., consisting of several separate, presently undifferentiated species that have all been traditionally but erroneously lumped together taxonomically as a single species.[99] In the genus chitra, two species of critically endangered turtles have been reported to grow to massive sizes. The Asian narrow-headed softshell turtle (Chitra chitra), at up to, and the Indian narrow-headed softshell turtle (Chitra indica), at up to, are also contenders for the title of the largest extant freshwater turtle.[100] [101]
- The Galápagos tortoise (Chelonoidis nigra) and the Aldabra giant tortoise (Aldabrachelys gigantea) are considered the largest truly terrestrial reptiles alive today.[3] While the Aldabra tortoise averages larger at, the more variable-sized Galapagos tortoise can reach a greater maximum size of and in total length.[102] [103] The Aldabra giant tortoise has a maximum recorded weight of . The African spurred tortoise (Centrochelys sulcata) is the third-largest extant tortoise (and the largest mainland tortoise) in the world. The large adults of this species may reach in length and weigh more than .[104] Other relatively large-sized tortoises include the Yellow-footed tortoise (Chelonoidis denticulatus) and Leopard tortoise (Stigmochelys pardelis), at up to, and the Asian forest tortoise (Manouria emys), at up to or more, can be rather large as well.[105] [106] The tortoise Megalochelys, of the Pleistocene epoch from what is now Pakistan and India, was even larger, at nearly in shell length[107] and .[108]
- The largest of side-necked turtles (Pleurodira) is the Arrau turtle (Podocnemis expansa). Its carapace length is up to and adults can reach up to in weight.[109] There are also reports of these turtles weighing up to .[110] The Mata mata (Chelus fimbriata) is another large species of side-necked turtle with a carapace length of up to and weight of more than .
- There are many extinct turtles that vie for the title of the largest ever.[3] The largest freshwater turtle seems to be Stupendemys, with an estimated parasagittal carapace length of and weight of up to .[111] A close contender is Archelon ischyros, a sea turtle, which reached a length of and a weight of .[112]
Meiolaniformes
A terrestrial relative of turtles survived until about 2,000 years ago, the Australasian Meiolania at about long and a weight of over .[3] Later research suggests the maximum length possibly over .[113]
Plesiosaurs (Plesiosauria)
Plesiosaurs were aquatic reptiles of the Mesozoic era. They had a broad flat body, a short tail, and strong flippers. Most of the Plesiosauroidea group are identified by their long necks, while Pliosauroidea are usually short-necked. The largest known plesiosauroid is Aristonectes, with a body length of 10m-11.86mm (30feet-38.91feetm) and body mass of .[114] The largest well known pliosauroid is Pliosaurus funkei at in length.[115]
Crocodilians (Crocodilia)
- Some species of crocodiles such as Saltwater crocodile (Crocodylus porosus), Nile crocodile (Crocodylus niloticus), American crocodile (Crocodylus acutus) and the critically endangered Orinoco crocodile (Crocodylus intermedius) can reach lengths of or more. The largest known specimen among the living crocodilians was an Orinoco crocodile with a length of .[1] One of the largest known Saltwater crocodile measured and was shot in Papua New Guinea.[1] A long individual was captured alive in Mindanao in 2011.[2] The largest confirmed Saltwater crocodile on record was long, and weighed about .[116] In 2006, Guinness World Records accepted claims of a 7m (23feet), 2000kg (4,000lb) male saltwater crocodile, living within Bhitarkanika National Park.[117] Due to the difficulty of trapping and measuring a large living crocodile, the accuracy of these dimensions is yet to be verified. These observations and estimations have been made by park officials over the course of ten years, from 2006 to 2016, however, regardless of the skill of the observers it cannot be compared to a verified tape measurement, especially considering the uncertainty inherent in visual size estimation in the wild.[118] The largest Nile crocodile accurately measured, shot near Mwanza, Tanzania, measured and weighed about .[116] Another large Nile crocodile specimen was purported to be a man-eater from Burundi named Gustave; it was thought to have been more than long. The American crocodile is also one of the largest crocodile species, with large males in the southern part of their range reported to approach in size. Based on projections from various skulls, the largest males may have reached in length, and their predicted mass reached up to .[119] Other crocodiles can also grow to large sizes, such the Mugger crocodile, which typically reaches an average maximum length of, and has a maximum reported length of . The extinct Crocodylus thorbjarnarsoni was the largest species in its genus, growing up to in length.[120] The largest true crocodile ever existed is Euthecodon which estimated to have reached or even long.[121] [122]
- A long gharial was killed in the Ghaghara River in Faizabad in August 1920.[123] Male gharials may grow up to a length of .[124] [28] The heaviest recorded gharial was a male measuring in total length and weighing .[125] The False gharial is also a large crocodilian with males reaching in length, weighing up to at least .[34] The largest gavialid to ever exist was the extinct Rhamphosuchus from the Miocene of Asia. It was originally thought to be long and more than in weight but later estimations suggest and . Based on its fossils, the latter species was less massive and heavy than the other giant crocodilians, weighing an estimated .[126] [127]
- The largest member of the family Alligatoridae is the Black caiman (Melanosuchus niger) with the American alligator (Alligator mississippiensis) sometimes growing to similar lengths. Black caimans can reach more than in length and weigh up to .[128] American alligators can be almost as large, with males reaching in length and weighing over .[129] Unverified reports suggest lengths of up to for the black caiman and for the American alligator, reaching weights of over, but such lengths are probably exaggerated.
- The giant prehistoric caiman, Purussaurus, from northern South America during the Miocene epoch grew up to long and could weigh up to 8 tonnes, making it one of the largest crocodilians ever.[130] [131] Other contenders for the largest crocodilian ever include the late Cretaceous period Deinosuchus, at up to and .[132] Relative of crocodilians, Sarcosuchus imperator of the early Cretaceous was found in the Sahara desert and could measure up to and weigh an estimated .[133]
Pterosaurs (Pterosauria)
A Mesozoic reptile is believed to have been the largest flying animal that ever existed: the pterosaur Quetzalcoatlus northropi, from North America during the late Cretaceous. This species is believed to have weighed up to 126kg (278lb), measured 7.9m (25.9feet) in total length (including a neck length of over 3m (10feet)) and measured up to 10- across the wings.[134] [135] Another possible contender for the largest pterosaur is Hatzegopteryx, which is estimated to have had an 11- wingspan.[135] [134] An unnamed Mongolian pterodactyloid pterosaur[136] and Arambourgiania from Jordan could reach a wingspan of nearly .[137]
Non-avian dinosaurs (Dinosauria)
Notes and References
- Whitaker . R. . Whitaker . N. . 2008 . Who's got the biggest? . Crocodile Specialist Group Newsletter . 27 . 4 . 26–30 .
- Britton . A. R. C. . Whitaker . R. . Whitaker . N. . Here be a Dragon: Exceptional Size in Saltwater Crocodile (Crocodylus porosus) from the Philippines . Herpetological Review . 2012 . 43 . 4 . 541–546 .
- Book: Wood, G. . 1983 . The Guinness Book of Animal Facts and Feats . 978-0-85112-235-9 . registration . 256. Guinness Superlatives .
- Book: Ogamba, E.N. and Abowei, JFN . Some Aquatic Reptiles in Culture Fisheries Management . July 2012 . International Journal of Fishes and Aquatic Sciences . 5–15 .
- Webb, G. J.. Hollis, G. J.. Manolis, S. C.. amp . 1991. Feeding, growth, and food conversion rates of wild juvenile saltwater crocodiles (Crocodylus porosus). Journal of Herpetology. 462–473. 1564770. 25. 4. 10.2307/1564770.
- Wood, G. (1983). The Guinness Book of Animal Facts and Feats. Sterling Pub Co Inc.
- Book: Guggisberg, C.A.W. . Crocodiles: Their Natural History, Folklore, and Conservation . 1972 . 978-0-7153-5272-4 . 195. David & Charles .
- Brazaitis, P. (1989). The forensic identification of crocodilian hides and products. In: Crocodiles: Their Ecology, Management, and Conservation. IUCN Special Publication of Crocodile Specialist Groups of the Species Survival Commission. pp. 17–43.
- Graham, A. D. (1968). The Lake Rudolf Crocodile (Crocodylus niloticus Laurenti) Population. Masters of Science Thesis, The University of East Africa.
- Book: Stefanie B. Ganswindt . Non-invasive assessment of adrenocortical function in captive Nile crocodiles (Crocodylus niloticus) and its relation to housing conditions . October 2012 . University of Pretoria .
- Web site: Nile Crocodile (Crocodylus niloticus). 2013-12-14. 2018-09-23. https://web.archive.org/web/20180923081324/http://wildliferanching.com/content/nile-crocodile-crocodylus-niloticus. dead.
- Book: "TERRAPENE CAROLINA TRIUNGUIS (Three-toed Box." Herpetological Review, 41(4) . ORNATA, T. O. . 489 .
- Book: FESTIVA, AMEIVA and Ameivafestiva . Herpetological Review . December 2010 . 490 .
- Web site: ADW: Crocodylus acutus: INFORMATION. Jake Fishman. Animal Diversity Web.
- http://www.arkive.org/orinoco-crocodile/crocodylus-intermedius/ Orinoco crocodile videos, photos and facts – Crocodylus intermedius
- Zoo . Gladys Porter . Critically Endangered Orinoco Crocodiles Coming to Gladys Porter Zoo . www.prnewswire.com . en.
- Web site: Orinoco Crocodile. WAZA. 2013-05-02. https://web.archive.org/web/20131110142610/http://www.waza.org/en/zoo/choose-a-species/reptiles/crocodiles/crocodylus-intermedius. 2013-11-10. dead.
- http://www.kwata.net/medias/images/upload/PUBLI_de%20Thoisy_2003_CSG.pdf French Guiana
- Book: Vliet . Kent A. . Alligators: The Illustrated Guide to Their Biology, Behavior, and Conservation . 3 March 2020 . JHU Press . 978-1-4214-3338-7 . 150 . en.
- Book: Chris Johnson, Sarah R. Anderson, James Moore, David Warrell, Jon Dallimore, Chris Imray · . Oxford handbook of expedition and wilderness medicine . 2015 . Oxford University Press . Oxford . 9780199688418 . 545 . 2nd .
- http://www.dinosaurcorporation.com/caimanskull.html Black Caiman, Black Caiman Skull
- http://crocodilian.com/cnhc/csp_mnig.htm Crocodilian Species – Black Caiman (Melanosucus niger)
- Martonos . Cristian . Dumitru . Ioana . Dezdrobitu . Cristian . Blendea . Alexandra . Irimescu . Irina . Damian . Aurel . Anatomical Studies Concerning the Skeleton of the Head in the Alligator (Alligator mississippiensis) . Bulletin of University of Agricultural Sciences and Veterinary Medicine Cluj-Napoca. Veterinary Medicine . 2014 . English . 1843-5270.
- Web site: American Alligator. 25 April 2016 .
- Book: Stevenson, C. . Whitaker, R. . 2010 . Gharial Gavialis gangeticus . 139–143 . Crocodiles. Status Survey and Conservation Action Plan . Third . Manolis, S. C. . Stevenson, C. . Crocodile Specialist Group . Darwin . http://www.iucncsg.org/365_docs/attachments/protarea/23_G-6764939a.pdf.
- Web site: Gavial . www.aquaticcommunity.com.
- https://web.archive.org/web/20070625033007/http://animals.nationalgeographic.com/animals/reptiles/gavial.html Gavials (Gharials), Gavial (Gharial) Pictures, Gavial (Gharial) Facts
- Web site: Gharial . September 26, 2011 . dead . https://web.archive.org/web/20111018085525/http://wwfindia.org/about_wwf/priority_species/gharial/ . October 18, 2011 .
- http://www.euroturtle.org/outline/Leatherback%20Sea%20Turtle.pdf Leatherback Sea Turtle
- Web site: AquaFacts . 2013-12-14 . 2011-09-07 . https://web.archive.org/web/20110907073934/http://www.vanaqua.org/education/aquafacts/leatherbacks.html . dead .
- Web site: Largest turtle/chelonian . Guinness World Records.
- http://www.zoonegaramalaysia.my/RPFalseGharial.pdf
- Erickson . Gregory M. . Gignac . Paul M. . Steppan . Scott J. . Lappin . A. Kristopher . Vliet . Kent A. . Brueggen . John D. . Inouye . Brian D. . Kledzik . David . Webb . Grahame J. W. . Insights into the Ecology and Evolutionary Success of Crocodilians Revealed through Bite-Force and Tooth-Pressure Experimentation . PLOS ONE . 14 March 2012 . 7 . 3 . e31781 . 10.1371/journal.pone.0031781 . 22431965 . 3303775 . 2012PLoSO...731781E . 1932-6203. free .
- Ahmad . A. A. . Dorrestein . G. M. . Oh . S. J. W. Y. . Hsu . C. D. . Multi-organ Metastasis of Fibrolamellar Hepatocellular Carcinoma in a Malayan Gharial (Tomistoma schlegelii) . Journal of Comparative Pathology . 1 August 2017 . 157 . 2 . 80–84 . 10.1016/j.jcpa.2017.06.007 . 28942308 . en . 0021-9975.
- Web site: False gharial – Tommy . Ferme aux crocodiles.
- Book: Kyle Shaney, Bruce Shwedick, Boyd Simpson and Colin Stevenson . Tomistoma (Tomistoma schlegelii) . Crocodile Specialist Group (IUCN) .
- Web site: Marsh Crocodile . www.wii.gov.in . en.
- Lang . J. W. . Andrews . H. . Whitaker . R. . 1989 . Sex determination and sex ratios in Crocodylus palustris . American Zoologist . 29 . 3. 935–952 . 10.1093/icb/29.3.935. free .
- Web site: African Slender-Snouted Crocodile – The Maryland Zoo in Baltimore. The Maryland Zoo in Baltimore.
- Web site: Slender-Snouted Crocodile – San Diego Zoo Animals.
- Web site: African Slender-snouted Crocodile . The Maryland Zoo.
- Book: Michael J. Lynch, Stephen F. Pires . Quantitative studies in green and conservation criminology : the measurement of environmental harm and crime . 2019 . Taylor & Francis . Abingdon, Oxon . 9780429844232 . 96 .
- Book: J Milan, R Hedegaard . Interspecific variation in tracks and trackways from extant crocodylians . 2010 . New Mexico Museum of Natural History and Science Bulletin . 15–29 .
- Web site: Loggerhead Turtle Sea Turtles Species WWF . World Wildlife Fund . en.
- Web site: ECOS: Species Profile . ecos.fws.gov.
- Web site: Loggerhead sea turtle, facts and photos . https://web.archive.org/web/20210412074734/https://www.nationalgeographic.com/animals/reptiles/facts/loggerhead-sea-turtle . dead . April 12, 2021 . Animals . en . 11 April 2010.
- Web site: Largest hardshell sea turtle . Guinness World Records . 6 March 2018 . en-gb.
- Wood . S. C. . Gatz . R. N. . Glass . M. L. . Oxygen transport in the green sea turtle . Journal of Comparative Physiology B . 1 April 1984 . 154 . 3 . 275–280 . 10.1007/BF02464407 . 20253025 . en . 1432-136X.
- Gatz . R. N. . Glass . M. L. . Wood . S. C. . Pulmonary function of the green sea turtle, Chelonia mydas . Journal of Applied Physiology . 1 February 1987 . 62 . 2 . 459–463 . 10.1152/jappl.1987.62.2.459 . 3558205 . en . 8750-7587.
- http://animaldiversity.ummz.umich.edu/site/accounts/information/Geochelone_nigra.html ADW: Geochelone nigra: Information
- Bastille-Rousseau . Guillaume . Yackulic . Charles B. . Frair . Jacqueline L. . Cabrera . Freddy . Blake . Stephen . Allometric and temporal scaling of movement characteristics in Galapagos tortoises . Journal of Animal Ecology . September 2016 . 85 . 5 . 1171–1181 . 10.1111/1365-2656.12561 . 27336221 . en . 0021-8790. free . 2016JAnEc..85.1171B .
- Web site: Largest tortoise (specimen) . Guinness World Records . en-gb.
- Web site: Aldabra tortoise. 25 April 2016.
- Barker . David G. . Barten . Stephen L. . Ehrsam . Jonas P. . Daddono . Louis . 2012 . The Corrected Lengths of Two Well-known Giant Pythons and the Establishment of a new Maximum Length Record for Burmese Pythons, Python bivittatus . Bulletin Chicago Herpetology Society . 47 . 1 . 1–6 . 2016-02-04.
- https://web.archive.org/web/20100204214402/http://animals.nationalgeographic.com/animals/reptiles/green-anaconda Green Anacondas, Green Anaconda Pictures, Green Anaconda Facts
- http://www.cotswoldwildlifepark.co.uk/meet-animals/python.htm Cotswold Wildlife Park and Gardens | Reticulated python
- Web site: Largest albino snake in captivity . Guinness World Records . 7 December 2012 . en-gb.
- Web site: Burmese Python National Geographic . https://web.archive.org/web/20210228013329/https://www.nationalgeographic.com/animals/reptiles/facts/burmese-python . dead . February 28, 2021 . Animals . en . 10 September 2010.
- Head . J. J. . Bloch . J. I. . Hastings . A. K. . Bourque . J. R. . Cadena . E. A. . Herrera . F. A. . Polly . P. D. . Jaramillo . C. A. . 10.1038/nature07671 . Giant boid snake from the Palaeocene neotropics reveals hotter past equatorial temperatures . Nature . 457 . 7230 . 715–717 . 2009 . 19194448 . 2009Natur.457..715H . 4381423 .
- Web site: Keeled Rat Snake – Ptyas carinata – Not Dangerous | Thailand Snakes. 20 January 2013.
- Auliya, M. (2010). Conservation Status and Impact of Trade on the Oriental Rat Snake Ptyas mucosa in Java, Indonesia. TRAFFIC Southeast Asia.
- Das, I. (2015). A field guide to the reptiles of South-East Asia. Bloomsbury Publishing.
- Book: Spilotes pullatus (Tiger Rat Snake or Clibo) . The Online Guide to the Animals of Trinidad and Tobago .
- Web site: Mulga Snake. Animal Species. Cecilie. Beatson. 25 November 2018. 26 August 2019. Australian Museum.
- Web site: Oxyuranus scutellatus – General Details. Clinical Toxinology Resource. University of Adelaide. 17 April 2012.
- Web site: Massive red-bellied black snake surprises Newcastle wrangler called in to remove it . ABC News . 3 October 2014 . 4 Dec 2018 . Australian Broadcasting Corporation . 10 February 2019 . https://web.archive.org/web/20190210060005/https://www.abc.net.au/news/2014-10-02/snake-wrangler-catches-massive-red-bellied-black-snake-newcastle/5786950 . live .
- http://research.calacademy.org/redirect?url=http://researcharchive.calacademy.org/research/herpetology/frogs/frogsimp/reptiles/e31.htm Gaboon Viper, Bitis gabonica
- Book: Laurence M. Klauber. 1972. University of California Press. Rattlesnakes: Their Habits, Life Histories, and Influence on Mankind. 1. 0-520-01775-7.
- Monster Basiliscus Weighed – 20 lb Rattlesnake . April 1, 2020 . SnakeBytesTV . 2022-06-27.
- Mehrtens JM. 1987. Living Snakes of the World in Color. New York: Sterling Publishers. .
- Wood, Gerald (1983). The Guinness Book of Animal Facts and Feats. .
- Web site: Longest lizard . Guinness World Records . en-gb.
- Web site: Wojtasek . Gregory . Varanus salvadorii (Crocodile Monitor) . Animal Diversity Web . en.
- Web site: The Largest Monitor Lizards by Paleonerd01 on DeviantArt . www.deviantart.com . 18 August 2019 . en.
- Encyclopedia: Molnar. R. E.. Varanoid Lizards of the World . The Long and Honorable History of Monitors and Their Kin . Indiana University Press. 2004. 588 . http://www.iupress.indiana.edu/catalog/product_info.php?products_id=21751 . 978-0-253-34366-6.
- Rivas, J.A. (2008). Pers. comm.
- Book: Philippe De Vosjoli. Susan Donoghue. Roger Klingenberg. David Blair.. The Green Iguana Manual. BowTie Press. 111. 2003. 1882770676.
- Dorge. Ray. A Tour of the Grand Cayman Blue Iguana Captive-Breeding Facility. Reptiles: Guide to Keeping Reptiles and Amphibians . 4. 9. 1996. 32–42.
- Web site: The 10 Largest Lizards in The World. A-z-animals.com. 18 August 2021 . 12 February 2022.
- News: es. Cruz M. Márquez B.. Estado poblacional de las iguanas terrestres (Conolophus subcristatus, C. pallidusy C.marthae: Squamata, Iguanidae), Islas Galápagos. Boletín Técnico, Serie Zoológica. Sangolquí, Équateur. ESPE. 9. June 2010. 19–37. .
- Miralles . Macleod . Rodríguez . Ibáñez . Jiménez-Uzcategui . Quezada . Vences . Steinfartz . amp. 2017 . Shedding Light On the Imps of Darkness: An Integrative Taxonomic Revision of the Galápagos Marine Iguanas (Genus Amblyrhynchus) . Zoological Journal of the Linnean Society . XX . 3 . 1–33 . 10.1093/zoolinnean/zlx007. free .
- Wikelski, M. . Trillmich, F. . 1997 . Body Size and Sexual Size Dimorphism in Marine Iguanas Fluctuate as a Result of Opposing Natural and Sexual Selection: An Island Comparison . Evolution . 51 . 3 . 922–936 . 10.1111/j.1558-5646.1997.tb03673.x . 28568579 . 205780374 . free .
- Endangered animals of the world pp. 48
- Allison Ballance and Rod Morris, "Island Magic; wildlife of the south seas", David Bateman publishing, 2003
- Book: Wilson, Kerry-Jayne. Canterbury University Press. 978-0-908812-52-3 . 411. Flight of the Huia: Ecology and Conservation of New Zealand's Frogs, Reptiles, Birds and Mammals . Christchurch, N.Z. 2004.
- https://www.bbc.co.uk/science/seamonsters/factfiles/giantmosasaur.shtml Science & Nature – Sea Monsters – Fact File: Giant Mosasaur
- Web site: Triassic Giant. https://web.archive.org/web/20110716093513/http://www.tyrrellmuseum.com/exhibits/triassic_giant.htm. live. 13 October 2022. 16 July 2011.
- Sander, P.M.. Romero Pérez de Villar, P.. Furrer, H.. Wintrich, T.. 2022. Giant Late Triassic Ichthyosaurs from the Kössen Formation of the Swiss Alps and Their Paleobiological Implications. Journal of Vertebrate Paleontology. 41. 6. e2046017. 10.1080/02724634.2021.2046017. 248444094. https://web.archive.org/web/20220920173105/https://www.parc-ela.ch/sites/parc_ela/files/2022-05/Sander%20et%20al.%202022_02724634.2021.pdf. 20 September 2022. live.
- Nicholls . E. L. . Manabe . M. . Giant ichthyosaurs of the Triassic—a new species of Shonisaurus from the Pardonet Formation (Norian: Late Triassic) of British Columbia . Journal of Vertebrate Paleontology . 24 . 4 . 838–849 . 2004 . 10.1671/0272-4634(2004)024[0838:GIOTTN]2.0.CO;2 . 130274656 .
- Web site: Researchers have found a 205-million-year-old jawbone from one of the largest animals that ever lived. Newsweek. 9 April 2018 . https://web.archive.org/web/20220921213249/http://www.newsweek.com/giant-ichthyosaur-205-million-year-old-jawbone-discovered-one-largest-animals-877542?yptr=yaho. 21 September 2022. live.
- De la Salle . Paul . R. Lomax . Dean . A. Massare . Judy . Gallois . Ramues . vanc . 2018 . A giant Late Triassic ichthyosaur from the UK and a reinterpretation of the Aust Cliff 'dinosaurian' bones . PLOS ONE . 10.6084/m9.figshare.5975440 . 13 . 4 . https://web.archive.org/web/20220227083139/https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0194742. 27 February 2022. live.
- Lomax . D. R. . de la Salle . P. . Perillo . M. . Reynolds . J. . Reynolds . R. . Waldron . J. F. . The last giants: New evidence for giant Late Triassic (Rhaetian) ichthyosaurs from the UK . 2024 . . 19 . 4 . e0300289 . 10.1371/journal.pone.0300289 . free . 38630678 . 11023487 .
- Sander . P.M. . Griebeler. E.M.. Klein. N.. Juarbe. J.V.. Wintrich. T.. Revell. L.J.. Schmitz. L.. 2021. Early giant reveals faster evolution of large body size in ichthyosaurs than in cetaceans. Science. 374. 6575. eabf5787 . 10.1126/science.abf5787. 34941418 . 245444783 . https://web.archive.org/web/20221010231502/http://www.faculty.umb.edu/liam.revell/pdfs/Sander_etal_2021.Science.pdf . 10 October 2022 . live.
- https://www.newscientist.com/article/2317948-ichthyosaur-tooth-from-the-swiss-alps-is-largest-ever-discovered/
- Web site: Largest turtle/chelonian . Guinness World Records . en-gb.
- Book: Hoagland . Porter . Steele . John H. . Thorpe . Steve A. . Turekian . Karl K. . Marine Policy & Economics . 2010 . Academic Press . 978-0-08-096481-2 . 162 . en.
- Lukas I. Alpert Sacred century-old turtle pulled from Vietnam lake in effort to save its life. New York Daily News (2011-04-04)
- Seth Mydans. "How to Survive in Cambodia: For a Turtle, Beneath Sand," New York Times (2007-05-18).
- Web site: Largest freshwater turtle (living) . Guinness World Records . 27 October 2016 . en-gb.
- Web site: Asian Narrow-headed Softshell Turtle . EDGE of Existence.
- Web site: Narrow-headed Softshell Turtle . EDGE of Existence.
- Ebersbach, V.K. (2001). Zur Biologie und Haltung der Aldabra-Riesenschildkröte (Geochelone gigantea) und der Galapagos-Riesenschildkröte (Geochelone elephantopus) in menschlicher Obhut unter besonderer Berücksichtigung der Fortpflanzun (PhD thesis). Hannover: Tierärztliche Hochschule. http://elib.tiho-hannover.de/dissertations/ebersbachk_2001.pdf.
- Fritts, T.H. (1983). Morphometrics of Galapagos tortoises: evolutionary implications. In Bowman, I.R.; Berson, M.; Leviton, A.E. Patterns of evolution in Galapagos organisms. San Francisco: American Association for the Advancement of Science. pp. 107–122.
- Book: Pizzi, R and Goodman, G and Gunn-Moore, D and Meredith, A and Keeble, Emma . Pieris japonica intoxication in an African spurred tortoise (Geochelone sulcata) . 2005 . London: The Association . 487–488 .
- Web site: Baker . Hillary H. . Grubb . Jordan N. . Psammobates pardalis (Leopard Tortoise) . Animal Diversity Web . en.
- Book: Iain J. Gordon, Herbert H. T. Prins . The ecology of browsing and grazing II . 2019 . Berlin . 978-3-030-25865-8 . 339–404 .
- Hirayama . Ren . Sonoda . Teppei . Takai . Masanaru . Htike . Thaung . Thein . Zin Maung Maung . Takahashi . Akio . Megalochelys: gigantic tortoise from the Neogene of Myanmar . 6 April 2015 . PeerJ PrePrints . 10.7287/peerj.preprints.961v1 . en . free .
- Gregory S. Paul . Guy D. Leahy. 1994. Terramegathermy in the time of the titans: Restoring the metabolics of colossal dinosaurs. The Paleontological Society Special Publications. 7: Dino Fest. 177–198. 10.1017/S2475262200009515. https://web.archive.org/web/20220920172524/http://gspauldino.com/TerraTimeofTitans.pdf. 20 September 2022.
- Web site: Podocnemis expansa . . 28 September 2017.
- Book: Cesar L. Barrio-Amoros, Ínigo Narbaiza . Turtles of the Venezuelan Estado Amazonas . 2008 . Radiata 17(1) . 2–19 .
- Cadena. E.-A.. Scheyer. T.M.. Carrillo-Briceño. J.D.. Sánchez. R.. Aguilera-Socorro. O.A.. Vanegas. A.. Pardo. M.. Hansen. D.M.. Sánchez-Villagra. M.R.. 12 Feb 2020. The anatomy, paleobiology, and evolutionary relationships of the largest extinct side-necked turtle. Science Advances. 6. 7. eaay4593 . 10.1126/sciadv.aay4593. 32095528. 7015691. 2020SciA....6.4593C . free.
- Colston, T. J., Kulkarni, P., Jetz, W., & Pyron, R. A.. Phylogenetic and spatial distribution of evolutionary diversification, isolation, and threat in turtles and crocodilians (non-avian archosauromorphs). BMC Evolutionary Biology. 2020 . 20. 1. 81 . 10.1186/s12862-020-01642-3. 32650718 . 7350713 . free . 2020BMCEE..20...81C .
- Lauren E. Brown, Don Moll. The enigmatic palaeoecology and palaeobiogeography of the giant, horned, fossil turtles of Australasia: a review and reanalysis of the data. Herpetological Journal. 29. October 2019. 4 . 252–263. 10.33256/hj29.4.252263. 0268-0130. https://web.archive.org/web/20220618074344/https://www.thebhs.org/publications/the-herpetological-journal/volume-29-number-4-october-2019/1979-08-the-enigmatic-palaeoecology-and-palaeobiogeography-of-the-giant-horned-fossil-turtles-of-australasia-a-review-and-reanalysis-of-the-data/file. 18 June 2022. live.
- 2019-10-01. A giant elasmosaurid (Sauropterygia; Plesiosauria) from Antarctica: New information on elasmosaurid body size diversity and aristonectine evolutionary scenarios. Cretaceous Research. en. 102. 37–58. 10.1016/j.cretres.2019.05.004. 0195-6671. O'Gorman. J.P.. Santillana. S.. Otero. R.. Reguero. M.. 2019CrRes.102...37O . 181725020.
- Knutsen . Espen M. . Druckenmiller . Patric S. . Hurum . Jørn H. . 2012 . A new species of Pliosaurus (Sauropterygia: Plesiosauria) from the Middle Volgian of central Spitsbergen, Norway . Norwegian Journal of Geology . 92 . 235–258 . 1502-5322 . https://web.archive.org/web/20200301151304/https://njg.geologi.no/images/NJG_articles/NJG_2_3_2012_13_Knutsen_etal_Scr.pdf . 1 March 2020.
- Book: Wood . Gerald L. . The Guinness book of animal facts and feats . 1982 . Guinness Superlatives . Enfield, Middlesex . 978-0-85112-235-9 . 3rd.
- Web site: World's Largest Reptile Found in India . ohmynews.com . Mishra . Braja Kishore . 14 June 2006 . dead . https://web.archive.org/web/20080108203350/http://english.ohmynews.com/articleview/article_view.asp?no=298369&rel_no=1 . 8 January 2008 .
- Bayliss, P. (1987). Survey methods and monitoring within crocodile management programmes. Surrey Beatty & Sons, Chipping Norton, pages 157–175
- Book: Juan Gabriel Abarca, Charles R. Knapp . Herpetological Review . December 2010 . 490 .
- Brochu . C. A. . Storrs . G. W. . 10.1080/02724634.2012.652324 . A giant crocodile from the Plio-Pleistocene of Kenya, the phylogenetic relationships of Neogene African crocodylines, and the antiquity of Crocodylus in Africa . Journal of Vertebrate Paleontology . 32 . 3 . 587 . 2012 . 2012JVPal..32..587B . 85103427 .
- Storrs, G. W. (2003). Late Miocene-Early Pliocene crocodilian fauna of Lothagam, southwest Turkana Basin, Kenya. In: Lothagam: The Dawn of Humanity in Eastern Africa pp. 137–159. New York. Columbia University Press. .
- Web site: Крупнейшие крокодиломорфы . 6 September 2022 . vk.com . ru.
- Pitman, C. R. S. . Charles Pitman (game warden) . 1925 . The length attained by and the habits of the Gharial (G. gangeticus) . Journal of the Bombay Natural History Society . 30 . 3 . 703 .
- Whitaker, R. . Basu, D. . 1982 . The Gharial (Gavialis gangeticus): A review . Journal of the Bombay Natural History Society . 79 . 3 . 531–548 .
- Hone . David . Mallon . Jordan C. . Hennessey . Patrick . Witmer . Lawrence M. . Ontogeny of a sexually selected structure in an extant archosaur Gavialis gangeticus (Pseudosuchia: Crocodylia) with implications for sexual dimorphism in dinosaurs . PeerJ . 12 May 2020 . 8 . e9134 . 10.7717/peerj.9134 . 32435543 . 7227661 . en . 2167-8359 . free .
- Heilprin, Angelo (1974). The Geographical and Geological Distribution of Animals: The International Science Series Vol. 57. Ayer Publishing. .
- Web site: Prehistoric Crocodile Evolution. ThoughtCo. https://web.archive.org/web/20220627235449/https://www.thoughtco.com/prehistoric-crocodile-profile-4047616. 27 June 2022.
- Book: Vliet . Kent A. . Alligators : the illustrated guide to their biology, behavior, and conservation . 2020 . Baltimore . 978-1-4214-3338-7 . 150.
- Coulson . Roland A. . Metabolic rate and the flow theory: A study in chemical engineering . Comparative Biochemistry and Physiology Part A: Physiology . 1 January 1986 . 84 . 2 . 217–229 . 10.1016/0300-9629(86)90607-9 . 2873921 . en . 0300-9629.
- Schwimmer, David R. (2002).
- Aguilera . O. A. . Riff . D. . Bocquentin-Villanueva . J. . 10.1017/S147720190600188X . A new giant Purussaurus (Crocodyliformes, Alligatoridae) from the Upper Miocene Urumaco Formation, Venezuela . Journal of Systematic Palaeontology . 4 . 3 . 221–232 . 2006 . 2006JSPal...4..221A . 85950121.
- The Size of Deinosuchus, King of the Crocodylians: The Paleobiology of Deinosuchus. Indiana University Press. pp. 42–63. .
- O'Brien, H. D. . Lynch, L. M. . Kent A Vliet . Brueggen, J. . Erickson, G.M. . Gignac, P.M. . Crocodylian Head Width Allometry and Phylogenetic Prediction of Body Size in Extinct Crocodyliforms. Integrative Organismal Biology . 2019 . 1 . 1]. obz006 . 10.1093/iob/obz006. 33791523 . 7671145 . free .
- Mark P. Witton, David M. Martill and Robert F. Loveridge, 2010, "Clipping the Wings of Giant Pterosaurs: Comments on Wingspan Estimations and Diversity", Acta Geoscientica Sinica, 31 Supp.1: 79–81
- Witton . M. P. . Naish . D. . McClain . Craig R . 10.1371/journal.pone.0002271 . A Reappraisal of Azhdarchid Pterosaur Functional Morphology and Paleoecology . PLOS ONE . 3 . 5 . e2271 . 2008 . 18509539. 2386974 . 2008PLoSO...3.2271W . free .
- Web site: Ancient Winged Terror Was One of the Largest Animals to Fly. 31 October 2017 . https://web.archive.org/web/20190330060740/https://news.nationalgeographic.com/2017/10/new-pterosaur-found-mongolia-largest-fossils-science/ . 30 March 2019.
- Takanobu Tsuihiji, Brian Andres, Patrick M O'Connor, Mahito Watabe, Khishigjav Tsogtbaatar, Mainbayar Buuvei . 2017 . Gigantic pterosaurian remains from the Upper Cretaceous of Mongolia . Journal of Vertebrate Paleontology . 37 . 5 . e1361431 . 10.1080/02724634.2017.1361431. 2017JVPal..37E1431T . 134424023 .
- Carpenter, K. (2006). "Biggest of the big: a critical re-evaluation of the mega-sauropod Amphicoelias fragillimus". In Foster, J. R. and Lucas, S. G., eds., 2006, Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36: 131–138.
- Book: MOLINA-PEREZ, RUBEN. LARRAMENDI, ASIER.. Dinosaur facts and figures: the sauropods and other sauropodomorphs. Princeton University Press. 2020. 978-0-691-19069-3. New Jersey. 254. 1125972915.
- Book: Peter Matthews. The Guinness Book of Records. 1992. Guinness Publishing. 42. 9780851129785 .
- Book: Tim Footman, Mark C. Young. Guinness World Records 2001. Bantam Books. May 2001. 276. 9780553583755.
- Web site: Curtice. Brian. 2021. New Dry Mesa Dinosaur Quarry Supersaurus vivianae (Jensen 1985) axial elements provide additional insight into its phylogenetic relationships and size, suggesting an animal that exceeded 39 meters in length.
- Paul. Gregory S.. 2019. Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals. Annals of the Carnegie Museum. 85. 4. 335–358. 10.2992/007.085.0403. 210840060.
- Wedel. Mathew J.. Cifelli, R. L.. Sanders, R.. K.. 59141243. 2000. Osteology, paleobiology, and relationships of the sauropod dinosaur Sauroposeidon. https://web.archive.org/web/20200626075134/https://pdfs.semanticscholar.org/9efe/8db6eb32247f90a760110d5a9695977fb761.pdf. dead. June 26, 2020. Acta Palaeontologica Polonica. 45. 343–388.
- Book: Guinness World Records. Bantam Books. 110. 2004. 9780553587128 .
- Therrien . F. . Henderson, D.M. . 2007 . My theropod is bigger than yours...or not: estimating body size from skull length in theropods . Journal of Vertebrate Paleontology . 27 . 1 . 108–115 . 10.1671/0272-4634(2007)27[108:MTIBTY]2.0.CO;2 . 86025320 . 0272-4634.
- dal Sasso . C. . Maganuco, S. . Buffetaut, E. . Mendez, M.A. . 2005 . New information on the skull of the enigmatic theropod Spinosaurus, with remarks on its sizes and affinities . Journal of Vertebrate Paleontology . 25 . 4 . 888–896. 10.1671/0272-4634(2005)025[0888:NIOTSO]2.0.CO;2. 85702490 .
- Vickaryous, M.K., Maryanska, T., & Weishampel, D.B. 2004. Ankylosauria. In: Weishampel, D.B., Dodson, P., & Osmólska, H. (Eds.). The Dinosauria (2nd edition). Berkeley: University of California Press. Pp. 363–392.
- Coombs . Walter P.. Theoretical Aspects of Cursorial Adaptations in Dinosaurs . . 53 . December 1978 . 393–418 . 10.1086/410790 . 4. 84505681.
- Alexander . R.M. . 1985 . Mechanics of posture and gait of some large dinosaurs . Zoological Journal of the Linnean Society . 83 . 1–25 . 10.1111/j.1096-3642.1985.tb00871.x.
- Seebacher . F. . 2001 . A new method to calculate allometric length-mass relationships of dinosaurs . Journal of Vertebrate Paleontology . 21 . 1 . 51–60. 10.1671/0272-4634(2001)021[0051:ANMTCA]2.0.CO;2 . 4524171 . 53446536 .
- Book: Holtz, Thomas R. Jr. . 2012 . Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages . Winter 2011 Appendix.
- Morris. William J.. 1981 . A new species of hadrosaurian dinosaur from the Upper Cretaceous of Baja California: ?Lambeosaurus laticaudus. Journal of Paleontology. 55. 2. 453–462.
- Zhao. X.. Li, D. . Han, G. . Hao, H. . Liu, F. . Li, L. . Fang, X. . 2007 . Zhuchengosaurus maximus from Shandong Province. Acta Geoscientia Sinica. 28. 2 . 111–122 . 10.1007/s10114-005-0808-x. 119700784.