List of largest birds explained
The largest extant species of bird measured by mass is the common ostrich (Struthio camelus), closely followed by the Somali ostrich (Struthio molybdophanes). A male ostrich can reach a height of 2.8m (09.2feet) and weigh over 156.8kg (345.7lb), A mass of 200kg (400lb) has been cited for the ostrich but no wild ostriches of this weight have been verified.[1] Ostrich eggs are the largest of any bird, averaging 1.4kg (03.1lb).[2]
The largest wingspan of any extant bird is that of the wandering albatross (Diomedea exulans) of the Sub-Antarctic oceans. The largest dimensions found in this species are an approximate head-to-tail length of 1.44m (04.72feet) and a wingspan of 3.65m (11.98feet).
The largest bird of all time was likely the elephant bird Aepyornis maximus, which was estimated to have weighed NaNkg (-2,147,483,648lb) and stood at 3m (10feet) tall.[3]
The largest wingspan of all time likely belonged to Pelagornis sandersi at roughly 5.2 m (17 ft).[4] P. sandersi was also likely the largest bird to ever fly.[5]
Largest extinct birds
Table of heaviest bird species
Largest extant birds
Table of heaviest extant bird species
The following table is a list of the heaviest extant bird species based on maximum reported or reliable mass, with the average weight is also given for comparison. These species are almost all flightless, having denser bones and heavier bodies. Flightless birds comprise less than 1% of all extant bird species.
By families
- New World vultures are generally considered belonging to this order,[40] although their inclusion is not accepted by all.[41] If included, the largest species of this order, based on body weight and wingspan, is the Andean condor (Vultur gryphus) of western South America. The Andean condor can reach a wingspan of 3.2m (10.5feet)[42] and a weight of 15kg (33lb).[43]
- Excluding New World vultures, the largest extant species is the Eurasian black vulture (Aegypius monachus). The Eurasian black vulture can attain a maximum weight of 14kg (31lb), a height of up to 1.2m (03.9feet), and a wingspan of 3.1m (10.2feet).[44] Other vultures can be almost as large, with the Himalayan vulture (Gyps himalayensis) reaching lengths of up to 1.5m (04.9feet) due to its long neck.[45]
- The largest living member of this order, in terms of length and height, is the secretarybird (Sagittarius serpentarius) of sub-Saharan Africa. It measures NaNm (-2,147,483,648feet) in height and NaNm (-2,147,483,648feet) in length. Its wingspan can reach NaNm (-2,147,483,648feet) and have a weight of NaNkg (-2,147,483,648lb).[46]
- The largest predatory bird, specifically the largest eagle, is a source of contention.
- The harpy eagle (Harpia harpyja) of neotropical forests is often cited as the most massive eagle, with wild females up to 10kg (20lb) in weight and captive females occasionally growing to weights of over 12kg (26lb).
- The Steller's sea eagle (Haliaeetus pelagicus) of Asia's North Pacific, with unconfirmed weights of up to 12.7kg (28lb), and an average weight of 6.7kg (14.8lb), is sometimes regarded as the heaviest eagle.
- The up to 1.12m (03.67feet) Philippine eagle (Pithecophaga jefferyi) has the greatest length of any eagle. The harpy and Philippine eagles, due to having to navigate in deep forest, are relatively short-winged and do not exceed 2or, respectively, in wingspan.
- The golden eagle (Aquila chrysaetos) is of marginally larger wingspan, with the Himalayan subspecies recorded at 2.77m (09.09feet).
- The white-tailed eagle (Haliaeetus albicilla) measures 66– in length with a 1.78– wingspan. Its wingspan, with a midpoint of 2.18m (07.15feet), is on average the largest of any eagle.
- The white-tailed eagle is sometimes considered the fourth-largest eagle in the world, and is on average the fourth-heaviest.
- The martial eagle (Polemaetus bellicosus) is the largest eagle in Africa, and the fifth-heaviest (on average) eagle in the world, with a length of 78–, a weight of 3– and a wingspan of 188-.
- The longest wingspan of an eagle ever recorded was an Australian wedge-tailed eagle (Aquila audax) at 2.83m (09.28feet). However, less substantiated records indicate that the Steller's sea eagle may reach at least 2.74m (08.99feet).[47]
- The now extinct Haast's eagle (Hieraaetus moorei), which existed alongside early aboriginal people in New Zealand, was by far the largest eagle known and perhaps the largest raptor ever. Adult female Haast's are estimated to have averaged up to 1.4m (04.6feet) in length, weighing up to 15kg (33lb), with a relatively short 3m (10feet) wingspan.[48]
- The largest of the accipitrine hawks is the Eurasian goshawk (Accipiter gentilis) of temperate Eurasia. They range in size variably, but on average measure NaNcm (-2,147,483,648inches) in length, have a wingspan of NaNcm (-2,147,483,648inches) and weigh NaNkg (-2,147,483,648lb).[49] The Henst's goshawk (Accipiter henstii) and Meyer's goshawk (Accipiter meyerianus) do rival it in terms of wing size and body mass.
- Among the buteonine hawks, the largest species are the ferruginous hawk (Buteo regalis) and the upland buzzard (Buteo hemilasius) of North America and Asia respectively. The former can have a wingspan of NaNcm (-2,147,483,648inches), weigh NaNkg (-2,147,483,648lb) and measure NaNcm (-2,147,483,648inches) in length.[50] The weight of the upland buzzard, which can be in the range of NaNkg (-2,147,483,648lb), broadly overlaps that of the ferruginous hawk, even though it is slightly larger at NaNcm (-2,147,483,648inches) long and with a wingspan of NaNcm (-2,147,483,648inches).
- The swamp harrier (Circus approximans) of Australasia is believed to be the largest species of harrier, measuring NaNcm (-2,147,483,648inches) long, having a wingspan of NaNcm (-2,147,483,648inches) and weighing NaNkg (-2,147,483,648lb).[51]
- The largest species of kite is the red kite (Milvus milvus). With a wingspan of NaNcm (-2,147,483,648inches), it measures NaNcm (-2,147,483,648inches) in length and weighs NaNkg (-2,147,483,648lb).
- The largest waterfowl species by average size is the trumpeter swan (Cygnus buccinator) of Northern North America, which can reach a length of 1.82m (05.97feet), a wingspan of 3.1m (10.2feet) and a weight of 17.3kg (38.1lb).[52] The heaviest single waterfowl ever recorded was a cob (Cygnus olor) from Poland which weighed 23kg (51lb), and was allegedly too heavy to take flight.
- The largest species of goose is the Canada goose (Branta canadensis), more specifically the subspecies known as the giant Canada goose (Branta canadensis maxima). Individuals can reach more than 9.1kg (20.1lb) in weight.[53]
- The largest 'duck' species is the Muscovy duck (Cairina moschata) of the Americas. Males can weigh from NaNkg (-2,147,483,648lb) and can measure up to 86cm (34inches).[54] However, its genus is now considered to be paraphyletic with the species currently being placed in the subfamily Tadorninae (shelducks and shelgeese). If so, the largest species of the true ducks or dabbling ducks (Anatinae) is the mallard (Anas platyrhynchos).[55] They can measure NaNcm (-2,147,483,648inches) in length, have a wingspan of NaNcm (-2,147,483,648inches) and a weight of NaNkg (-2,147,483,648lb).[56]
- The largest species of Apodiformes is the white-naped swift (Streptoprocne semicollaris), endemic to southern Mexico, and the purple needletail (Hirundapus celebensis), of the Philippine islands. Both reach weights of up to 225g, lengths of up to 25cm (10inches) and wingspans as long as 0.6m (02feet).[57]
- Traditionally included in this order, by far the largest hummingbird species is the giant hummingbird (Patagona gigas) of the Andes Mountains. "Giant" is a relative term among the hummingbirds, the smallest-bodied variety of birds, with the giant hummingbird species weighing up to 24g with a length of 23cm (09inches).[58]
- The longest hummingbird species, indeed the longest in the order, is the adult male black-tailed trainbearer (Lesbia victoriae), which can measure up to 25.5cm (10inches). The majority of this length is due to the hummingbird's extreme tail streamers. Another size champion among hummingbirds is the sword-billed hummingbird, a fairly large species of which approximately half of its 21cm (08inches) length derives from its bill. This is by far the largest bill-to-body-size ratio of any bird.[59]
- The largest species of this order of nocturnal birds is the neotropical great potoo (Nycitbius grandis), which can grow to a weight of 680g and a height of 60cm (20inches). Heavier Caprimulgiformes have been recorded in juvenile specimens of the Australian tawny frogmouth (Podargus strigoides), which can weigh up to 1.4kg (03.1lb). Other species nearly as large as the potoo are the Papuan frogmouth (Podargus papuensis) of New Guinea and the neotropic, cave-dwelling oilbird (Steatornis caripensis), both growing as large as 48cm (19inches). The wingspan of the great potoo and the oilbird can be more than 1m (03feet), the largest of the order.[60] [61]
- The largest species of the nightjar family, the great eared nightjar (Eurostopodus macrotis) of East Asia, is of smaller proportions. Great eared nightjars can reach 150g in weight and 41cm (16inches) in height.[62]
- The largest species in this diverse order is the great black-backed gull (Larus marinus) of the North Atlantic, attaining a height as large as 0.79m (02.59feet), a wingspan of 1.7m (05.6feet) and a weight of up to 2.3kg (05.1lb). The glaucous gull (L. hyperboreus) is smaller on average than the black-back but has been weighed as heavy as 2.7kg (06lb).[63]
- Among the most prominent family of "small waders", the sandpipers reach their maximum size in the Far Eastern curlew (Numenius madagascariensis) at up to 0.6m (02feet) in length and 1.1m (03.6feet) across the wings. The more widespread Eurasian curlew (N. arquata) can weigh up to 1.36kg (03lb).[64] [65]
- Less variable in size, the largest species of plovers is the Australasian masked lapwing (Vanellus miles) which grows up to 0.4m (01.3feet) long with a 0.85m (02.79feet) wingspan and a weight of 400g.[66] The widely distributed Caspian tern (Hydroprogne caspia), is relatively large and heavily built. Caspians can range up to 782g in weight, with a 1.4m (04.6feet) wingspan and a length of 0.6m (02feet).[67] [68]
- The largest extant alcid is the sub-Arctic thick-billed murre (Uria lomvia), which can weigh up to 1.48kg (03.26lb), with a length of 0.48m (01.57feet) and a wingspan of 0.76m (02.49feet).[69] However, until its extinction, the flightless great auk (Pinguinus impennis) of the North Atlantic was both the largest alcid and the second-largest member of the order. Great auks could range up to 6.8kg (15lb) and 0.9m (03feet) tall.[70]
- Miomancalla howardi was the largest charadriiform of all time, weighing approximately 1.5feet(?) more than the Great Auk with a height of approximately 1m (03feet).[71]
- The longest-bodied and tallest species in this order is the saddle-billed stork of Africa (Ephippiorhynchus senegalensis), which often exceeds 1.5m (04.9feet) tall and has a wingspan of up to 2.7m (08.9feet).[72] Reaching a similar height but more heavily built among the storks are the neotropical jabiru (Jabiru mycteria), the Asian greater adjutant (Leptoptilos dubius) and the African marabou stork (L. crumeniferus), all of which weigh up to 8to.[73] The greater adjutant and marabou nearly equal the Andean condor in maximum wingspan, with all three birds believed to reach or exceed a wingspan of 3.16m (10.37feet). Standing up to 1.53m (05.02feet), with a wingspan of up to 2.3m (07.5feet) and a weight up to 5kg (11lb), the African goliath heron (Ardea goliath) is the largest of the herons and egrets. Juvenile white-bellied heron (A. insignis) have been reported to weigh up to 8.5kg (18.7lb) with heights of 1.58m (05.18feet).[74]
- Many of the largest flying birds in the fossil record may have been members of the Ciconiiformes. The heaviest flying bird ever, Argentavis magnificens, is part of a group, the teratorns, that is considered an ally of the New World vultures.[75]
- The largest ibis is the giant ibis (Thaumatibis gigantea). Adults can grow to 102– long, with a standing height of up to 100cm (00inches) and are estimated to weigh approximately 4.2kg (09.3lb). Among standard measurements, the wing chord is 52.3-, the tail is 30cm (10inches), the tarsus is 11cm (04inches) and the culmen is 20.8-. The crested ibis (Nipponia nippon) of Japan is as large as 78.5cm (30.9inches) in height and 30.9cm (12.2inches) in length.
- The largest mousebird species, the speckled mousebird (Colius striatus), weighs 2oz with a height of over 14inches.[76]
Pigeons (Columbiformes)
- The largest species of the pigeon/dove complex is the Victoria crowned pigeon (Goura victoria) of Northern New Guinea. Some exceptionally large Victoria crowned pigeons have reached 3.7kg (08.2lb) and 85cm (33inches). The largest arboreal pigeon is the Marquesan imperial pigeon (Ducula galeata), which can grow approximately 0.8m (02.6feet) across the wings and can weigh 1kg (02lb).[77]
- The largest pigeons and doves known to have existed were the dodo (Raphus cucullatus) and the Rodrigues solitaire (Pezophaps solitaria). Both flightless species may have exceeded 1m (03feet) in height. The dodo is frequently cited as the largest-ever pigeon, potentially weighing as much as 28kg (62lb), although recent estimates have indicated that an average wild dodo weighed much less at approximately 10.2kg (22.5lb).[78] [79]
Rollers, Kingfishers, Bee-eaters, motmots, and todies (Coraciiformes)
- The largest kingfisher is the giant kingfisher (Megaceryle maxima), at up to 48cm (19inches) long and 425g in weight.[80] The common Australian species, the laughing kookaburra (Dacelo novaeguineae), may be heavier still, as individuals exceeding 450g are not uncommon. A kookaburra's wingspan can range up to 0.9m (03feet).[81]
Hornbills, hoopoe, and wood-hoopoes (Order Bucerotiformes)
- The largest species of Coraciiformes is the southern ground hornbill (Bucorvus leadbeateri), which can reach weights of up to 6.2kg (13.7lb) and grow as long as 1.3m (04.3feet).[82] Several arboreal, Asian hornbills can also grow very large, with the great hornbill (Buceros bicornis) weighing up to 4kg (09lb), and the helmeted hornbill (Rhinoplax vigil) measuring as much as 1.7m (05.6feet) in total length.[83] [84] The larger hornbills have a wingspan of up to 1.83m (06feet).[85]
Cuckoos, coucals and roadrunners (Cuculiformes)
- The largest of the cuckoos is the Australasian channel-billed cuckoo (Scythrops novaehollandiae), which can range up to a weight of 0.93kg (02.05lb), a 1m (03feet) wingspan and a length of 0.66m (02.17feet).[86] [87]
- Many authorities now support the split of falcons from the Accipitriformes, despite similar adaptations, due to the genetic evidence showing they are not closely related.[88] The largest species of falcon is the gyrfalcon (Falco rusticolus). Large females of this species can range up to 2.1kg (04.6lb), span 1.6m (05.2feet) across the wings and measure 0.66m (02.17feet) long.
Gamebirds (Galliformes)
- The heaviest member of this order is the North American wild turkey (Meleagris gallopavo). The largest specimen ever recorded was shot in 2015, and weighed 17.05kg (37.59lb).[89] The heaviest domesticated turkey on record weighed 37kg (82lb).
- The longest gamebirds species, if measured from the tip of the bill to the end of the tail coverts, is the male green peafowl (Pavo muticus) of Southeast Asia at a length of up to 3m (10feet), with two-thirds of the length being made up by the tail coverts. It has a relatively large wingspan for a gamebird, spanning as much as 1.6m (05.2feet) across the wings.[90]
- The largest member of the grouse family is the Eurasian western capercaillie (Tetrao urogallus), weighing up to 6.7kg (14.8lb) with a length of 1m (03feet).
- A prehistoric, flightless family, sometimes called (incorrectly) "giant megapodes" (Sylviornis) of New Caledonia were the most massive galliformes ever, with lengths of up to 1.7m (05.6feet) weights up to approximately 40kg (90lb).[91]
- The largest species on average is the yellow-billed loon (Gavia adamsii) of the Arctic, at up to 1m (03feet) and 7kg (15lb). One exceptionally large North American Great northern diver (Gavia immer) was weighed at 8kg (18lb), heavier than any recorded yellow-billed loon. Wingspans of the largest loons can reach 1.52m (04.99feet).[92]
- The males of the Eurasian great bustard (Otis tarda) and the African kori bustard (Ardeotis kori) are the heaviest birds capable of flight, averaging up to 16kg (35lb) and weighing 2 to 3 times as much as their female counterparts. It is not resolved if one of these species is larger than the other, but both can reach a weight of at least 21kg (46lb) and measure up to 1.53m (05.02feet) long.[93] Some kori bustards have been reported from 23kg (51lb) to even 40kg (90lb), but all such reports are unverified or dubious.[94]
- The tallest flying bird on earth, also represented in the Gruiformes, is the sarus crane (Grus antigone) of Southern Asia and Australia, which can reach a height of 2m (07feet).[95] Heavier cranes are reported in other species, the red-crowned crane (Grus japonensis) and the Siberian crane (G. leucogeranus), both from Northeast Asia and both at up to 15kg (33lb), as opposed to a top weight of 12.8kg (28.2lb) in the sarus.[96] Wingspan in both the largest cranes and the largest bustards can range up to 2.5–.[97] [98]
- The most species-rich family in this order, the rails, reaches their largest size in the bulky takahē (Porphyrio hochstetteri) of New Zealand, an endangered species that can weigh up to 4.2kg (09.3lb) and measure 0.65m (02.13feet) long.[99] The aforementioned "terror bird", Brontornis burmeisteri, has traditionally been classified as a member of this order, although this may not be an accurate classification.
- The passerine or songbird order comprises more than half of all bird species, and are known for their generally small size, their strong voices and their frequent perching. Corvids are the largest of passerines, particularly the large races of the common raven (Corvus corax) and the Northeast African thick-billed raven (C. crassirostris). Large ravens can weigh 2kg (04lb), attain a 1.5m (04.9feet) wingspan and measure 0.8m (02.6feet) long.[100]
- The closest non-corvid contender to largest size is the Australian superb lyrebird (Menura novaehollandiae), which can reach a length of 1m (03feet), much of it comprised by their spectacular tail, and a weight of 1kg (02lb).[101]
- The largest species in the most species-rich passerine family, Tyrannidae or tyrant-flycatchers, is the great shrike-tyrant of the South Andes (Agriornis lividus), at 99.2g and 31cm (12inches), although the fork-tailed flycatcher (Tyrannus savana), to 41cm (16inches), is longer thanks to its extreme tail.[102]
- The namesake of the previous family, the Old World flycatchers, reaches its maximum size in the blue whistling thrush of India Southeast Asia (Myophonus caeruleus), if it is indeed a proper member of the family, at up to 122g and a length of 29cm (11inches).[103]
- Closely related to the Old World flycatchers, the thrush family's largest representative is the Great thrush of South America (Turdus fuscater), at up to 175g and 28to.[104]
- The largest bird family in Eurasia is the Old World warblers. As previously classified these warblers could get fairly large, up to 57g and 28cm (11inches) in the striated grassbird of Southeast Asia (Megalurus palustris). The Old World warblers have been split into several families, however, which leaves the barred warbler of central Eurasia (Sylvia nisoria), up to 36g and 17cm (07inches), as the largest "true warbler".[105]
- Not to be confused with the previous family, the largest of the well-known New World warblers is the aberrant yellow-breasted chat (Icteria virens), which can exceptionally measure up to 22cm (09inches) and weigh 53g.[106] [107]
- Another large family is the bulbuls, the largest of which is the south Asian straw-headed bulbul (Pycnonotus zeylanicus), to 94g and 29cm (11inches). The diverse, large family of babblers can reach 35cm (14inches) and 170g in the south Asian greater necklaced laughingthrush (Garrulax pectoralis).[108]
- The familiar domesticated species, the Java sparrow (Padda oryzivora), is (in the wild) the largest estrildid, at up to 28.3g and 17cm (07inches). The largest honeyeater, perhaps the most diverse Australasian bird family, is the crow honeyeater (Gymnomyza aubryana), at up to 290g and 30cm (10inches). The largest of the "true finches" is the collared grosbeak (Mycerobas affinis) of central and south Asia at up to 23cm (09inches) and 80g.[109]
- Among the largest bird families, the emberizids, reaches its largest size in the Abert's towhee (Pipilo aberti) of Southwest United States and north Mexico at up to 23cm (09inches) and 80g.[110]
- Closely related to the previous family is the tanagers, which can range up to 140g in the Andean-forest-dwelling white-capped tanager (Sericossypha albocristata). Another species-rich neotropical family is the ovenbirds, the largest of which, the great rufous woodcreeper (Xiphocolaptes major) of the Amazonian rainforest, can weigh up to 162g and 35cm (14inches). The specialized antbird family can range up to 156g and 35.5cm (14inches) in the giant antshrike (Batara cinerea). Among the most variably sized passerine families is the icterids.
- The largest icterid is the olive oropendola (Psarocolius bifasciatus), in which males can range up to 52cm (20inches) and 550g.[111] The latter species competes with the similarly sized Amazonian umbrellabird (Cephalopterus ornatus) as the largest passerine in South America.
- Pelicans rank amongst the largest flying birds. The largest species of pelican is the Eurasian Dalmatian pelican (Pelecanus crispus), which can attain a length of 1.83m (06feet) and a body weight of 15kg (33lb). The great white pelican (P. onocrotalus) of Europe and Africa is almost as large. The Australian pelican (P. conspicillatus) is slightly smaller but has the largest bill of any bird, at as much as 49cm (19inches) long. A large pelican can attain a wingspan of 3.6m (11.8feet), second only to the great albatrosses among all living birds.[112]
- The largest of the cormorants is the flightless cormorant of the Galapagos Islands (Nannopterum harrisi), at up to 5kg (11lb) and 1m (03feet), although large races in the great cormorant (Phalacrocorax carbo) can weigh up to 5.3kg (11.7lb).[113] [114] The spectacled cormorant of the North Pacific (Urile perspicillatus), which became extinct around 1850, was larger still, averaging around 6.4kg (14.1lb) and 1.15m (03.77feet).
- The widely distributed magnificent frigatebird is of note for having an extremely large wingspan, up to 2.5m (08.2feet), for its relatively light body, at up to only 1.9kg (04.2lb).[115]
- Pelagornithidae or pseudotooth birds included several species that were behind only Argentavis magnificens in size among all flying birds. Characterized by the tooth-like protrusions along their bills, this unique family has been variously allied with the Pelecaniformes, tubenoses, large waders and even waterfowl. Their true linkage to extant birds remains in question, though pelecaniformes are the group most regularly considered related. Some of the largest pseudotooth birds have included, Osteodontornis of the late Miocene from the North Pacific, Gigantornis eaglesomei, from the Eocene era in what is now Nigeria and Dasornis, from Eocene era Europe. A new, unnamed species has been discovered which may outsize even these giants. Superficially albatross-like, each of these pseudotooth species may have attained lengths of 2.1-longNaN-long and wingspans of at least 6m (20feet). Body mass in these slender birds was probably only up to around 29kg (64lb).[116] [117] [118]
- The largest tropicbirds is the red-billed tropicbird (Phaethon aethereus). The adult is a slender, mainly white bird, 48 cm long, excluding the central tail feathers which double the total length, and a one-meter wingspan.
- The largest flamingo is the greater flamingo (Phoenicopterus roseus) of Eurasia and Africa. One of the tallest flying birds in existence when standing upright (exceeded only by the tallest cranes), this species typically weighs 3.5kg (07.7lb) and stands up to 1.53m (05.02feet) tall. At maximum, a male can weigh up to 4.55kg (10.03lb) and stand as high as 1.87m (06.14feet).[119] Wingspan is relatively small in flamingos, ranging up to 1.65m (05.41feet).[120]
Woodpeckers and allies (Piciformes)
- The largest species of this order is the toco toucan (Ramphastos toco) of the neotropic forest. Large specimens of this toucan can weigh to 870g and 0.65m (02.13feet), at which size the beak alone can measure approximately 20cm (10inches).[121]
- Until the 20th century, the largest woodpecker was the imperial woodpecker (Campephilus imperialis) of Mexico, with a length of up to 0.6m (02feet). This species is generally believed to have gone extinct following habitat destruction and hunting.[122] The closely related ivory-billed woodpecker (Campephilus principalis) of the Southeast United States and Cuba approached similar sizes at up to 0.5m (01.6feet) in length, with a wingspan of 0.78m (02.56feet) and a mass of at least 530g. Despite possibilities that it has survived in some deep swamp forests in Arkansas or Florida, the ivory-billed is also generally considered to have gone extinct.[123] The great slaty woodpecker (Mulleripicus pulverulentus) of southeast Asia is the largest woodpecker certain to exist, with a weight of up to 500g and a length of up to 0.58m (01.9feet).[124]
- Less well-known than the woodpeckers and toucans, barbets can range up to 273g and 33cm (13inches) in the great barbet (Megalaima virens).
- The largest jacamar is the great jacamar (Jacamerops aureus). It measures 29.5to in length and weighs between 63and.[125]
- The largest species of grebe is the South American great grebe (Podiceps major). It can reach a length of 0.8m (02.6feet), with a wingspan of 1m (03feet) and a weight of over 2kg (04lb).[126]
Tubenoses (Procellariiformes)
- The largest species of Procellariiformes is the wandering albatross (Diomedea exulans) of the sub-Antarctic oceans, which has the largest wingspan of any living bird. The maximum dimensions of this species are a length of 1.44m (04.72feet) and a wingspan of 3.65m (11.98feet). Unverified specimens have been reported to measure 5.3m (17.4feet). Immature wandering albatrosses have weighed as much as 15.9kg (35.1lb) at the time of their first flights, with the maximum reported weight of adults being 12.7kg (28lb).
- The Southern royal albatross (Diomedea epomophora) is slightly lesser in length, wingspan and weight.
- The largest procellarids is the southern giant petrel (Macronectes giganteus). It can reach a body length of 1 m (39 in), with a wingspan of 2.1 m (83 in) and a weight of 8 kg (17.6 lb).
- Hoatzin (Opisthocomus hoazin), the only member of its order, is a pheasant-sized South American bird, with a total length of 65cm (26inches) and a maximum weight of 1kg (02lb).
- The largest parrot by length and wingspan is the endangered hyacinth macaw (Anodorhynchus hyacinthinus) of the neotropic lowlands, reaching a length of nearly 1.2m (03.9feet) with a wingspan of 1.4m (04.6feet) and weighing as little as 2kg (04lb).[127] The heaviest parrot is the nearly extinct kākāpō (Strigops habroptilus),[128] which is part of the New Zealand parrot family. The flightless kākāpō does not exceed 0.68m (02.23feet) in length, but weighs up to 4.1kg (09lb).[129]
- The largest parakeet is the Alexandrine parakeet (Palaeornis eupatoria), reaching lengths of up to 60 centimetres and a mass of 250 grams.
- The largest species in the cockatoo family is the Australasian palm cockatoo (Probosciger aterrimus), at up to 0.6m (02feet) long with a weight of 1.2kg (02.6lb).[130]
Penguins (Sphenisciformes)
- The largest species of Sphenisciformes is the emperor penguin (Aptenodytes forsteri) of the Antarctic, with a maximum height of 1.35m (04.43feet) and a weight of 46kg (101lb).[39] The next largest living species is the king penguin, which grows to a maximum of 1m (03feet) in height and 18kg (40lb) in weight.[132] Now extinct, Anthropornis nordenskjoeldi, is believed to have reached a height of 1.8m (05.9feet) and a weight of up to 108kg (238lb).[133]
- The most massive owl is certainly either the Eurasian eagle-owl (Bubo bubo) or the endangered and similarly sized Blakiston's fish owl (Bubo blakistoni) of coastal Russia and Japan. Record-sized specimens of both species have weighed approximately 4.5kg (09.9lb) and measured over 0.75m (02.46feet) long.[134] In either species, the wingspan can range up to 2m (07feet).[135] [136] Longer still, but not as massive as the previous species (never more than 1.8kg (04lb) in weight), a large female great gray owl (Strix nebulosa) from the northern boreal forest can range up to 0.83m (02.72feet).[137]
- The largest of the barn or masked owl family is the Tasmanian masked owl (Tyto novaehollandiae castanops), which weighs up to 1.4kg (03.1lb) and measures up to 0.6m (02feet). The largest owl known to have existed was Ornimegalonyx oteroi of Cuba, a uniquely cursorial owl. The giant bird was estimated to stand over 1.1m (03.6feet) on the ground and to weigh at least 9.05kg (19.95lb).[138]
- The largest ratite is the ostrich (Struthio camelus), from the plains of Africa and Arabia. A large male ostrich can reach a height of 2.8m (09.2feet) and weigh over 156kg (344lb). A mass of 200kg (400lb) has been cited for the ostrich but no wild ostriches of this weight have been verified. Eggs laid by the ostrich are the largest in the world, weighing 1.4kg (03.1lb). The emu (Dromaius novaehollandiae) of Australia reaches 1to(-) at the shoulder with a full height of 150to(-). In length measured from the bill to the tail, emus range from 139to. The southern cassowary (Casuarius casuarius) from Australia and Papua New Guinea has a height of 127to. The greater rhea (Rhea americana) from South America weighs up to 20- and often measures 127to long from beak to tail with a height of approximately 1.5m (04.9feet).
- The largest bird in the fossil record may be the extinct elephant birds (Vorombe/Aepyornis) of Madagascar, which were related to the ostrich. They exceeded 3m (10feet) in height and 500kg (1,100lb) in weight. The last of the elephant birds became extinct approximately 1000 years ago.
- The tallest bird ever was the South Island giant moa (Dinornis robustus), part of the moa family of New Zealand that went extinct about 500 years ago. The moa stood up to 3.7m (12.1feet) tall, and weighed approximately half as much as a large elephant bird or mihirung due to its comparatively slender frame.
- The largest species of tinamou, a group of chunky, elusive ground-birds from neotropical forests, is the grey tinamou (Tinamus tao) of western South America. It can reach a weight over 2kg (04lb) and length of over 55cm (22inches).[139]
- The resplendent quetzal (Pharomachrus mocinno) of the montane forest of Central America is the largest trogon, though a few other quetzals approach similar sizes. It can weigh more than 226g and, in females and non-breeding or immature males, they can measure up to 0.4m (01.3feet) from the head to the tail. Upon developing tail streamers, adult males can reach lengths of up to 0.6m (02feet).[140]
See also
Notes and References
- http://www.avianmedicine.net/ampa/48.pdf Avian Medicine: Principles and Application
- Web site: Largest egg from a bird (living, specimen). 2020-11-05. Guinness World Records. en-GB.
- Book: Hume. J. P.. M.. Walters. 2012. Extinct Birds. A & C Black. London. 19–21. 978-1-4081-5725-1.
- News: 2014-07-07 . Fossil of 'largest flying bird' identified . 2024-05-29 . BBC News . en-GB.
- ibid.
- Handley . Warren D. . Chinsamy . Anusuya . Yates . Adam M. . Worthy . Trevor H. . 2016-09-02 . Sexual dimorphism in the late Miocene mihirung Dromornis stirtoni (Aves: Dromornithidae) from the Alcoota Local Fauna of central Australia . Journal of Vertebrate Paleontology . en . 36 . 5 . e1180298 . 10.1080/02724634.2016.1180298 . 2016JVPal..36E0298H . 88784039 . 0272-4634.
- Web site: Lazaro . Enrico de . 28 June 2019 . Giant Birds Roamed Europe Two Million Years Ago Paleontology Sci-News.com . 3 March 2023 . Sci.News: Breaking Science News . en-US.
- Degrange, F. J., Noriega, J. I., & Areta, J. I. (2012). Diversity and paleobiology of the Santacrucian birds. Early Miocene paleobiology in Patagonia: high-latitude paleocommunities of the Santa Cruz Formation, 138-155.
- Buffetaut . Eric . Angst . Delphine . 2021. A Giant Ostrich from the Lower Pleistocene Nihewan Formation of North China, with a Review of the Fossil Ostriches of China . Diversity . en . 13 . 2 . 47 . 10.3390/d13020047 . free . 1424-2818.
- Murray, P. F. & Vickers-Rich, P. (2004)
- Buffetaut1 . Eric . Angst . Delphine. 2017 . How Large was the Giant Ostrich of China?. EVOLUÇÃO - Revista de Geistória e Pré-História . 2 . 1 . 6–8 . 9 January 2023.
- News: Brennan . Imogen . Bones from ancestor of 'giant duck' discovered in Australia . PM . www.abc.net.au . 19 February 2016 . en-AU.
- Book: Molina-Pérez . Rubén . Dinosaurs Facts and Figures: The Theropods . Larramendi . Asier . Princeton University Press . 2019 . 978-0-691-18031-1 . 179.
- Grealy . Alicia . Miller . Gifford H. . Phillips . Matthew J. . Clarke . Simon J. . Fogel . Marilyn . Patalwala . Diana . Rigby . Paul . Hubbard . Alysia . Demarchi . Beatrice . Collins . Matthew . Mackie . Meaghan . Sakalauskaite . Jorune . Stiller . Josefin . Clarke . Julia A. . Legendre . Lucas J. . 2023-02-28 . Molecular exploration of fossil eggshell uncovers hidden lineage of giant extinct bird . Nature Communications . en . 14 . 1 . 914 . 10.1038/s41467-023-36405-3 . 36854679 . 9974994 . 2041-1723. free . 2023NatCo..14..914G .
- Brassey . Charlotte A. . Holdaway . Richard N. . Packham . Abigail G. . Anné . Jennifer . Manning . Philip L. . Sellers . William I. . 2013-12-18 . Kupczik . Kornelius . More than One Way of Being a Moa: Differences in Leg Bone Robustness Map Divergent Evolutionary Trajectories in Dinornithidae and Emeidae (Dinornithiformes). PLOS ONE. en . 8 . 12 . e82668 . 10.1371/journal.pone.0082668 . 1932-6203. 3867382. 24367537 . free . 2013PLoSO...882668B .
- Tambussi, C. P., & Degrange, F. J. (2013). The dominance of zoophagous birds: just a cliché?. In South American and Antarctic Continental Cenozoic Birds (pp. 87-102). Springer, Dordrecht.
- Titanis walleri: Bones of Contention. Gould. G. C.. Quitmyer. I. R.. Bulletin of the Florida Museum of Natural History. 45. 4. 201–229. 2005. 10.58782/flmnh.xumx1681 .
- Eric Buffetaut und Delphine Angst: Stratigraphic distribution of large flightless birds in the Palaeogene of Europe and its palaeobiological and palaeogeographical implications. Earth-Science Reviews 32 (2), 2014, S. 394–408.
- Book: 9780691190594. Dinosaur Facts and Figures: The Theropods and Other Dinosauriformes. Princeton University Press. June 25, 2019. 30 August 2022. 282. Rubén Molina-Pérez, Asier Larramendi, David Connolly, Gonzalo Ángel Ramírez Cruz, Andrey Atuchin.
- Hospitaleche. Carolina A.. 2014. New giant penguin bones from Antarctica: Systematic and paleobiological significance. Comptes Rendus Palevol. 13. 7. 555–560. 10.1016/j.crpv.2014.03.008. free. 2014CRPal..13..555A . 11336/32571. free.
- Ksepka . Daniel T. . Flights of fancy in avian evolution . American Scientist . 2014 . 102 . 1 . 39 . 10.1511/2014.106.36 . 43707746 . 0003-0996.
- Alexander . David E. . Ancient Argentavis soars again . Proceedings of the National Academy of Sciences of the United States of America . 24 July 2007 . 104 . 30 . 12233–12234 . 10.1073/pnas.0705515104 . 17640902 . 1941455 . 2007PNAS..10412233A . free .
- Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of Patagonia" [1], geredigeerd door Sergio F. Vizcaíno, Sergio F. Vizcno, Richard F. Kay en M. Susana Bargo
- Meat Weights and Nutritional Yield Values for New Zealand Archaeofauna
- Amadon (1947), Rasmussen et al. (1987, 2001)
- Book: Hume. J. P.. Walters. M.. 2012. Extinct Birds. A & C Black. London. 978-1-4081-5725-1. 137–138.
- Ksepka . Daniel T. . Flight performance of the largest volant bird . 7 July 2014 . . 10.1073/pnas.1320297111 . 111 . 29 . 10624–10629 . 25002475 . 4115518. free . 2014PNAS..11110624K .
- del Hoyo, et al.,Handbook of the Birds of the World. Volume 1: Ostrich to Ducks. Lynx Edicons (1992),
- Christopher P. Kofron (1999). Attacks to humans and domestic animals by the southern cassowary (Casuarius casuarius johnsonii) in Queensland, Australia. Journal of Zoology, 249, pp 375–381
- CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), .
- Marion, Remi, Penguins: A Worldwide Guide. Sterling Publishing Co. (1999),
- http://www.authorstream.com/Presentation/dawnvea09-169623-Leopard-Seals-group-Penguin-Slideshow-g-Entertainment-ppt-powerpoint/ Leopard Seals Group Penguin Slideshow Ppt Presentation
- Web site: The turkeys we eat today weigh twice as much as they did a few decades ago. Business Insider.
- Web site: Turkey Facts - Turkey for Holidays - University of Illinois Extension.
- Wood, Gerald (1983). The Guinness Book of Animal Facts and Feats. .
- del Hoyo, J; Elliot, A; Sargatal, J (1996). Handbook of the Birds of the World 3. Barcelona: Lynx Edicions. .
- BirdLife International (2007) Species factsheet: California Condor Gymnogyps californianus . Retrieved August 17, 2007
- Raptors of the World by Ferguson-Lees, Christie, Franklin, Mead & Burton. Houghton Mifflin (2001),
- Wood, Gerald The Guinness Book of Animal Facts and Feats (1983)
- American Ornithologists' Union (2010)
- Remsen et al. (2008)
- http://alltoptens.com/top-10-largest-birds-in-the-world/ Largest flying animals/birds in The World/Universe AllTopTens.com
- http://www.arkive.org/andean-condor/vultur-gryphus/ Andean condor videos, photos and facts – Vultur gryphus
- Christie, David A. & Ferguson-Lee, James, Raptors of the World. Princeton University Press (2006),
- http://www.oiseaux-birds.com/card-himalayan-griffon-vulture.html Himalayan Griffon Vulture
- Web site: Sagittarius serpentarius (Secretary bird). .
- https://web.archive.org/web/20071217063227/http://ngm.nationalgeographic.com/ngm/0301/feature5/zoom1.html Japan's Winter Wildlife Zoom In @ National Geographic Magazine
- Notes on the weight, flying ability, habitat, and prey of Haast's Eagle (Harpagornis moorei). https://web.archive.org/web/20061108034612/http://www.notornis.org.nz/free_issues/Notornis_39-1992/Notornis_39_4_239.pdf. 2006-11-08. 1992. Brathwaite, D. H.. Notornis. 39. 4. 239–247.
- Web site: Northern Goshawk Identification, All About Birds, Cornell Lab of Ornithology.
- Web site: Ferruginous Hawk Identification, All About Birds, Cornell Lab of Ornithology.
- Web site: Swamp Harriers | Beauty of Birds.
- Price, Christopher ed., Firefly Encyclopedia of Birds. Firefly Books (2003),
- Web site: Canada Goose | Types of Ducks & Geese.
- Web site: Muscovy Duck. 12 May 2021.
- Web site: The biggest dabbling duck . 21 April 2015 .
- Web site: Mallard Identification, All About Birds, Cornell Lab of Ornithology.
- Chantler, Phil, Swifts: A Guide to the Swifts and Treeswifts of the World. Pica Press (1993),
- Fjeldsa, Jon; Krabbe, Niels (1990). Birds of the High Andes. Zoological Museum, University of Copenhagen, Denmark. p. 876.
- Ridgely, Robert S., The Birds of Ecuador, Vol. 2: Field Guide. Cornell University Press (2001),
- http://en.caiman.com.br/2010/12/06/great-potoo-truths-and-legends/ Great Potoo- Truths & Legends
- http://www.oiseaux-birds.com/card-oilbird.html Oilbird Card
- Cleere, Nigel, Nightjars: A Guide to the Nightjars, Nighthawks, and Their Relatives. Yale University Press (1998),
- Howell, Steven N.G. & Dunn, Jon, Peterson Reference Guides: Gulls of the Americas. Houghton Mifflin Harcourt (2007), .
- Brazil, Mark, Birds of East Asia. Christopher Helm Ornithology (2009),
- http://www.birdfellow.com/birds/far-eastern-curlew-numenius-madagascariensis Far Eastern Curlew
- http://www.oiseaux-birds.com/card-masked-lapwing.html Masked Lapwing Card
- http://birdsinbackyards.net/species/Sterna-caspia Caspian Tern (Sterna caspia) – Birds in Backyards Fact sheet
- Web site: 2024-02-06 . How Big Is A Caspian Tern Actually? Find Out Here! . 2024-02-06 . en-GB.
- http://www.nhptv.org/natureworks/thickbilledmurre.htm Thick-billed Murre
- Fuller, Errol, Great Auk. Harry N. Abrams (1999), .
- Smith, N. 2015. Evolution of body mass in the Pan-Alcidae (Aves, Charadriiformes): the effects of combining neontological and paleontological data. Paleobiology. doi: 10.1017/pab.2015.24
- http://www.oiseaux-birds.com/card-saddle-billed-stork.html Saddle-billed Stork card
- Hancock & Kushan, Storks, Ibises and Spoonbills of the World. Princeton University Press (1992),
- The Bhutan Observer The Independent Voice (August 29th 2008)
- Miller, Loye H. (1909): Teratornis, a new avian genus from Rancho La Brea. University of California Publications, Bulletin of the Department of Geology 5: 305–317.
- http://www.mousebirds.com/speckled.html Speckled – Welcome to the wonderful world of Mousebirds
- Gibbs, David, Pigeons and Doves. A&C Black (2001),
- Vermeij, Geerat J. (2004). Nature: An Economic History. Princeton University Press,
- 10.1007/s00114-010-0759-7. Angst . D.. Buffetaut . E.. Abourachid . A.. 2011. The end of the fat dodo? A new mass estimate for Raphus cucullatus. Naturwissenschaften. 98. 3. 233–236. 21240603. . 2011NW.....98..233A . 29215473 .
- http://www.oiseaux-birds.com/card-giant-kingfisher.html Giant Kingfisher
- Knowles, Leslie, Kingfishers of the World. Times Edn (1995),
- http://www.oiseaux.net/birds/southern.ground-hornbill.html Southern Ground-Hornbill – Bucorvus cafer
- Alan Kemp, The Hornbills: Bucerotiformes. Oxford University Press (1995),
- http://www.arkive.org/helmeted-hornbill/rhinoplax-vigil/#text=Facts Helmeted hornbill videos, photos and facts - Rhinoplax vigil
- http://www.sandiegozoo.org/animalbytes/t-hornbill.html Hornbill Profile
- Payne, Robert B.,The Cuckoos. Oxford University Press (2005),
- http://www.naturalhistory.org.au/tmsite/index.php?option=com_content&view=article&id=266:channel-billed-cuckoo&catid=66:nature-notes&Itemid=200029 Channel-billed Cuckoo
- http://www.museum.lsu.edu/~Remsen/SACCprop383.html Separate Accipitriformes from Falconiformes
- http://www.nwtf.org/hunt/records/search?rank=weight&keyword=&harvest-location=&species=&type=&method=&multiple-spurs=&hunter-gender= National Wild Turkey Federation
- http://www.sandiegozoo.org/animalbytes/t-peafowl.html San Diego Zoo's Animal Bytes: Peafowl
- Jones, Darryl & Göth, Ann, Mound-Builders (Australian Natural History Series). CSIRO Publishing (2009),
- http://animaldiversity.ummz.umich.edu/site/accounts/information/Gavia_immer.html Gavia immer: Information
- http://scienceblogs.com/tetrapodzoology/2010/04/the_great_bustard_returns.php The Great bustard returns: Tetrapod Zoology
- http://ecoport.org/ep?Bird=89231&entityType=BI****&entityDisplayCategory=BI****1000%7CKori Entity Display : Ardeotis kori
- http://gruitag.org/uploads/media/Sarus_Crane_Studbook_Second_Edition_2009.pdf Sarus Crane
- The Wildlife Year, The Reader's Digest Association, (1991).
- http://nationalzoo.si.edu/Animals/Birds/Facts/FactSheets/fact-redcrowncrane.cfm Red-Crowned Crane Fact Sheet
- Sterling, et al., Vietnam: A Natural History. Yale University Press (2006),
- Taylor, Barry, Rails: A Guide to the Rails, Crakes, Gallinules and Coots of the World. Yale University Press (1998),
- http://www.oiseaux-birds.com/card-common-raven.html Common Raven
- Bambaradeniya, et al., The Illustrated Atlas of Wildlife. University of California Press (2009),
- http://www.enature.com/fieldguides/detail.asp?recNum=BD0705 FieldGuides: Species Detail
- Web site: Blue Whistling-Thrush. 2021-01-09. www.oiseaux-birds.com.
- Escobar Riomalo. Maria Paula. Gongora. Esteban. Arsitizabal Leost. Sophie. Thomas S. Schulenberg. 2020-03-04. Great Thrush (Turdus fuscater). Birds of the World. 10.2173/bow.grethr1.01. 216306066. en.
- Handbook of the Birds of the World, Volume 11: Old World Flycatcher's to the Old World Warblers. Lynx Edicions (2006),
- http://www.allaboutbirds.org/guide/Yellow-breasted_Chat/lifehistory Yellow-breasted Chat, Life History, All About Birds – Cornell Lab of Ornithology
- http://www.tnwatchablewildlife.org/details.cfm?displayhabitat=forest&sort=aounumber&typename=FOREST&uid=09041521203597078&commonname=Yellow-breasted%20Chat Tennessee Watchable Wildlife. Yellow-breasted Chat
- http://www.zootierliste.de/en/?klasse=2&ordnung=222&familie=22212&art=2221204 ZootierlisteHomepage
- Clement, Peter, Finches & Sparrows. Princeton University Press (1999),
- Byers, Clive, Sparrows and Buntings: A Guide to the sparrows and Buntings of North America and the World. Houghton Mifflin (1997), ASIN B000ZFNLXS
- Hilty, Steven L., Birds of Venezuela. Princeton University Press (2003),
- Harrison, Peter, Seabirds: An Identification Guide. Houghton Mifflin Harcourt (1991),
- http://www.ecostravel.com/en/ecuador/galapagos/wildlife/cormorants.php Galapagos Wildlife, Flightless Cormorant
- Philip H.R. Stepney. http://www.thecanadianencyclopedia.com/index.cfm?PgNm=TCE&Params=A1ARTA0001923Cormorant. The Canadian Encyclopedia
- http://www.allaboutbirds.org/guide/Magnificent_Frigatebird/lifehistory Magnificent Frigatebird, Life History, All About Birds – Cornell Lab of Ornithology
- http://www.britannica.com/animal/procellariiform Procelariiform
- http://dinosaurs.about.com/od/otherprehistoriclife/tp/dinowhats.htm 10 Giant Prehistoric Creatures – A List of History's Strangest Prehistoric Giants
- Mayr . G. . Rubilar-Rogers . D. . 10.1080/02724634.2010.501465 . Osteology of a new giant bony-toothed bird from the Miocene of Chile, with a revision of the taxonomy of Neogene Pelagornithidae . Journal of Vertebrate Paleontology . 30 . 5 . 1313 . 2010 . 2010JVPal..30.1313M . 84476605.
- http://en.tourduvalat.org/content/download/7378/73859/version/2/file/All+About+Greater+Flamingo.pdf All About Flamingos
- http://www.seaworld.org/infobooks/flamingos/fphysical.html Flamingos
- Short, Lester & Horne, Jennifer, Toucans, Barbets and Honeyguides (Bird Families of the World). Oxford University Press (2002),
- http://www.birdlife.org/datazone/speciesfactsheet.php?id=718 Imperial Woodpecker (Campephilus imperialis) – BirdLife species factsheet
- http://www.birds.cornell.edu/ivory/ Welcome — Ivory-billed Woodpecker
- Winkler, et al., Woodpeckers: An Identification Guide to the Woodpeckers of the World. Houghton Mifflin (1995),
- http://wikiaves.com.br/jacamaracu Jacamaraçu
- del Hoyo, Elliott & Sargatal. Handbook of Birds of the World, Volume 1 Ostrich to Ducks.
- http://www.sarapetstudios.com/exotic-birds.html Exotic Birds – Scarlet Macaw, Hyacinth Macaw, Gold Blue Macaw and Green Wing Macaw Wholesaler & Retailer
- Walley . H. D. . Ruback . P. A. . Ornithological Literature . The Wilson Journal of Ornithology . 118 . 4 . 581 . 2006 . 10.1676/1559-4491(2006)118[581:OL]2.0.CO;2 . 198156686 .
- Elliott, Greaeme P (2006). A simulation of the future of kakapo. Society 53(1): 164-172
- Forshaw, Joseph, Parrots of the World. TFH Publications (1978),
- del Hoyo, Elliott & Sargatal. Handbook of Birds of the World, Volume 4: Sandgrouse to Cuckoos.
- Field Immobilization of King Penguins with Tiletamine-Zolazepam (Uso de tiletamine-zolazepam para inmobilizar Aptenodytes patagonicus en el campo). Marie-Anne Thil and René Groscolas. Journal of Field Ornithology. 73. 3. 2002. 308–317. 4131109.
- http://www.app.pan.pl/archive/published/app56/app20091107.pdf Body Mass of Extinct Penguins
- http://quillcards.com/blog/index.php/2009/09/23/take-a-peek-at-boo-the-eagle-owl/ Take A Peek At Boo, The Eagle Owl – QUILLCARDS BLOG
- http://www.pauldfrost.co.uk/eeagleowl.html Eagle Owl
- A Guide to the Owls of the World by Konig, Weick & Becking. Yale University Press (1999),
- http://www.owlpages.com/owls/species.php?s=1770|Great Grey Owl (Strix nebulosa)
- Arredondo, Oscar (1976) translated Olson, Storrs L. The Great Predatory Birds of the Pleistocene of Cuba pp. 169–187 in "Smithsonian Contributions to Paleobiology number 27; Collected Papers in Avian Paleontology Honoring the 90th Birthday of Alexander Wetmore"
- Davies, Stephen, Ratites and Tinamous. Oxford University Press (2002),
- [Paul Johnsgard|Johnsgard, Paul A.]
