List of ichthyosauromorph genera explained

This list of ichthyosauromorphs is a comprehensive listing of all genera that have ever been included in the clade Ichthyosauromorpha, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered ichthyopterygian. Non-ichthyosaur ichthyopterygians shall be noted as such. This list contains 108 genera.

Scope and terminology

There is no official, canonical list of ichthyosauromorph genera but one of the most thorough attempts can be found at the "Ichthyosauromorpha" section of Mikko Haaramo's Phylogeny Archive.

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

• Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
• Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen. In this case, these nomina nuda will be deleted from this list in favor of the published name.
• Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
• Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

See also

• Ichthyosaur
• Ichthyopterygia
• List of dinosaurs
• List of mosasaurs
• List of plesiosaurs
• List of pterosaurs
• List of ichthyosaur type specimens
• Timeline of ichthyosaur research

External links

• USMP Berkeley's ichthyosaur introduction
• Ryosuke Motani's detailed Ichthyosaur homepage, with vivid graphics
• Eureptilia: Ichthyosauria - Palaeos
• Ichthyosauria - cladogram (Mikko's Phylogeny Archive)
• Hauff Museum, Germany - exhibiting the finds of Holzmaden
• http://www.collectionscanada.gc.ca/obj/s4/f2/dsk2/ftp02/NQ41566.pdf

Notes and References

1. Fischer,V. . Maisch, M.W. . Naish, D. . Kosma, R. . Liston, J. . Joger, U. . Krüger, F.J. . Pérez,J.P. . Tainsh, J. . Appleby, R.M.. 2012 . New Ophthalmosaurid Ichthyosaurs from the European Lower Cretaceous Demonstrate Extensive Ichthyosaur Survival across the Jurassic–Cretaceous Boundary . PLOS ONE . 7 . 1 . e29234 . 10.1371/journal.pone.0029234 . 22235274 . 3250416. 2012PLoSO...729234F . free .
2. Jair Israel Barrientos Lara . Jesús Alvarado Ortega . Marta S.Fernández . 2020 . Acuetzpalin carranzai gen et sp. nov. A new ophthalmosauridae (Ichthyosauria) from the Upper Jurassic of Durango, North Mexico . Journal of South American Earth Sciences . 98 . Article 102456 . 10.1016/j.jsames.2019.102456 . 2020JSAES..9802456B . 213102661 .
3. Bardet . N . Fernández . M . 2000 . A new ichthyosaur from the Upper Jurassic lithographic limestones of Bavaria . Journal of Paleontology . 74 . 3. 503–511 . 10.1666/0022-3360(2000)074<0503:aniftu>2.0.co;2. 131190803 .
4. Fernández . M. . 2007 . Redescription and phylogenetic position of Caypullisaurus (Ichthyosauria: Ophthalmosauridae) . Journal of Paleontology . 81 . 2. 368–375 . 10.1666/0022-3360(2007)81[368:RAPPOC]2.0.CO;2 . 130457040 .
5. Zverkov. N.G.. Grivoriev. D.V.. Wolnievicz. A.S.. Konstantinov. A.G.. Sobolev. E.S.. 2021. Ichthyosaurs from the Upper Triassic (Carnian–Norian) of the New Siberian Islands, Russian Arctic, and their implications for the evolution of the ichthyosaurian basicranium and vertebral column. Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 113 . 51–74 . 10.1017/S1755691021000372. 245153589 .
6. Dal Sasso, Cristiano and Giovanni Pinna, 1996. Besanosaurus leptorhynchus n. gen. n. sp., a new shastasaurid ichthyosaur from the Middle Triassic of Besano (Lombardy, N. Italy). Paleontologia Lombarda, Nuova seire Volume IV: 1-23.
7. Sepkoski . Jack . A compendium of fossil marine animal genera (entry on Reptilia) . Bulletins of American Paleontology . 364 . 560 . 2002 . 2008-09-28.
8. Efimov . VM. . 1998 . An Ichthyosaur, Otschevia pseudoscythica gen. et sp. nov from the Upper Jurassic Strata of the Ulyanovsk Region (Volga Region) . Paleontologicheskii Zhurnal . 32 . 2. 187–191 .
9. Maisch . MW . Matzke . AT. . 2000 . The Ichthyosauria . Stuttgarter Beiträge zur Naturkunde . Serie B (Geologie und Paläontologie) . 298 . 1–159 .
10. Ryosuke Motani . Da-Yong Jiang . Guan-Bao Chen . Andrea Tintori . Olivier Rieppel . Cheng Ji . Jian-Dong Huang . amp . 2015 . A basal ichthyosauriform with a short snout from the Lower Triassic of China . Nature . 517 . 7535 . 485–488 . 10.1038/nature13866 . 25383536 . 2015Natur.517..485M . 4392798 .
11. Fernández. Marta S.. Campos. Lisandro. Maxwell. Erin E.. Garrido. Alberto C.. 2021-01-02. Catutosaurus gaspariniae, gen. et sp. nov. (Ichthyosauria, Thunnosauria) of the Upper Jurassic of Patagonia and the evolution of the ophthalmosaurids. Journal of Vertebrate Paleontology. 41. 1. e1922427. 10.1080/02724634.2021.1922427. 2021JVPal..41E2427F . 236312664. 0272-4634.
12. Web site: Haaramo . Mikko . Euichthyosauria . Mikko's Phylogeny Archive . 11 March 2004. 9 October 2008.
13. Maisch . Matzke . A. T. . 2000 . The mixosaurid ichthyosaur Contectopalatus from the Middle Triassic of the German Basin . Lethaia . 33 . 2. 71–74 . 10.1080/00241160050150186. 2000Letha..33...71M .
14. Patrick S. Druckenmiller . Jørn H. Hurum . Espen M. Knutsen . Hans Arne Nakrem . amp . 2012 . Two new ophthalmosaurids (Reptilia: Ichthyosauria) from the Agardhfjellet Formation (Upper Jurassic: Volgian/Tithonian), Svalbard, Norway . Norwegian Journal of Geology . 92 . 2–3 . 311–339 .
15. Stephen L. Brusatte . Mark T. Young . Thomas J. Challands. Neil D. L. Clark . Valentin Fischer. Nicholas C. Fraser . Jeff J. Liston. Colin C. J. MacFadyen . Dugald A. Ross . Stig Walsh . Mark Wilkinson . amp . 2015 . Ichthyosaurs from the Jurassic of Skye, Scotland . Scottish Journal of Geology . 51 . 1 . 43–55 . 10.1144/sjg2014-018 . 2015ScJG...51...43B . 2268/176434 . 54614526 . free .
16. Xiao-hong Chen . Ryosuke Motani . Long Cheng . Da-yong Jiang . Olivier Rieppel . amp . 2015 . A New Specimen of Carroll's Mystery Hupehsuchian from the Lower Triassic of China . PLOS ONE . 10 . 5 . e0126024 . 10.1371/journal.pone.0126024 . 26017585 . 4446317 . 2015PLoSO..1026024C. free .
17. Robin S. Cuthbertson . Anthony P. Russell . Jason S. Anderson . amp . 2013 . Cranial morphology and relationships of a new grippidian (Ichthyopterygia) from the Vega-Phroso Siltstone Member (Lower Triassic) of British Columbia, Canada . Journal of Vertebrate Paleontology . 33 . 4 . 831–847 . 10.1080/02724634.2013.755989 . 2013JVPal..33..831C . 131501541 .
18. Web site: Haaramo . Mikko . Ichthyosauromorpha . Mikko's Phylogeny Archive . 14 July 2004 . 9 October 2008.
19. Barrientos-Lara. Jair Israel. Alvarado-Ortega. Jesús. 2021-11-01. A new ophthalmosaurid (Ichthyosauria) from the Upper Kimmeridgian deposits of the La Casita Formation, near Gómez Farías, Coahuila, northern Mexico. Journal of South American Earth Sciences. en. 111. 103499. 10.1016/j.jsames.2021.103499. 2021JSAES.11103499B. 0895-9811.
20. Aubrey Jane Roberts . Patrick Scott Druckenmiller . Glenn-Peter Sætre . Jørn Harald Hurum . amp . 2014 . A New Upper Jurassic Ophthalmosaurid Ichthyosaur from the Slottsmøya Member, Agardhfjellet Formation of Central Spitsbergen . PLOS ONE . 9 . 8 . e103152 . 10.1371/journal.pone.0103152 . 25084533 . 4118863. 2014PLoSO...9j3152R . free .
21. Болатовна. Якупова Джамиля. Максутович. Ахмеденов Кажмурат. 2021. НОВЫЙ ВИД KAZAKHSTANOSAURUS EFIMOVI YAKUPOVA ET AKHMEDENOV SP. NOV. (ICHTHYOSAURIA, UNDOROSAURIDAE) ИЗ ВЕРХНЕЮРСКИХ ОТЛОЖЕНИЙ СРЕДНЕГО ПОВОЛЖЬЯ РОССИИ. Ученые записки Казанского университета. Серия Естественные науки. 163. 2. 251–263. 2542-064X.
22. Lene Liebe Delsett . Aubrey J. Roberts . Patrick S. Druckenmiller . Jørn H. Hurum . 2017 . A New Ophthalmosaurid (Ichthyosauria) from Svalbard, Norway, and Evolution of the Ichthyopterygian Pelvic Girdle . PLOS ONE . 12 . 1 . e0169971 . 10.1371/journal.pone.0169971 . 28121995 . 2017PLoSO..1269971D . 5266267. free .
23. Cortés . D. . Maxwell . E.E. . Larsson . H.C.E. . 2021 . Re-appearance of hypercarnivore ichthyosaurs in the Cretaceous with differentiated dentition: revision of Platypterygius' sachicarum (Reptilia:Ichthyosauria, Ophthalmosauridae) from Colombia . Journal of Systematic Palaeontology. 19 . 14 . 969–1002 . 10.1080/14772019.2021.1989507. 2021JSPal..19..969C . 244512087 .
24. Valentin Fischer . Maxim S. Arkhangelsky . Gleb N. Uspensky. Ilya M. Stenshin . Pascal Godefroit . Pascal Godefroit . amp . 2014 . A new Lower Cretaceous ichthyosaur from Russia reveals skull shape conservatism within Ophthalmosaurinae . Geological Magazine . 151 . 1 . 60–70 . 10.1017/S0016756812000994 . 2014GeoM..151...60F . free .
25. W. Maisch . Michael . T. Matzke . Andreas . Magnipterygius huenei n. gen. n. sp., a new small stenopterygiid (Reptilia: Ichthyosauria) from the Posidonienschiefer Formation of SW Germany . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 2022 . 303 . 2 . 169–201 . 10.1127/njgpa/2022/1042 . 246797718 . 10 February 2022.
26. Valentin Fischer. Robert M. Appleby. Darren Naish. Jeff Liston. James B. Riding. Stephen Brindley. Pascal Godefroit. amp . 2013 . A basal thunnosaurian from Iraq reveals disparate phylogenetic origins for Cretaceous ichthyosaurs . Biology Letters . 9 . 4 . 20130021 . 10.1098/rsbl.2013.0021 . 23676653 . 3730615.
27. Web site: Haaramo . Mikko . Ichthyosauria. Mikko's Phylogeny Archive . 11 March 2008. 9 October 2008.
28. Maisch . M. W . Matzke . A. T. . 1998 . Mikadocephalus gracilirostris n. gen., n. sp., a new ichthyosaur from the Grenzbitumenzone (Anisian-Ladinian) of Mont San Giorgio (Switzerland) . Paläontologische Zeitschrift . 71 . 3–4. 267–289 . 10.1007/bf02988496. 129713553 .
29. Web site: Haaramo . Mikko . Ophthalmosauria . Mikko's Phylogeny Archive . 11 March 2008. 9 October 2008.
30. Shang Qing-Hua . Li Chun . 2009 . On the occurrence of the ichthyosaur Shastasaurus in the Guanling Biota (Late Triassic), Guizhou, China . Vertebrata PalAsiatica . 47 . 3 . 178–193 .
31. Sander . P.M. . Chen . X. . Cheng . L. . Wang . X. . 2011 . Short-Snouted Toothless Ichthyosaur from China Suggests Late Triassic Diversification of Suction Feeding Ichthyosaurs . PLOS ONE . 6 . 5. e19480 . 10.1371/journal.pone.0019480 . Claessens . Leon . 21625429 . 3100301. 2011PLoSO...619480S . free .
32. Barrientos-Lara, J. I.; Alvarado-Ortega, J. (2021). "A new Tithonian ophthalmosaurid ichthyosaur from Coahuila in northeastern Mexico". Alcheringa: An Australasian Journal of Palaeontology. in press. doi:10.1080/03115518.2021.1922755.
33. Jiang . D.-Y. . Schmitz . L. . Motani . R. . Hao . W.-C. . Sun . Y.-L. . 2007 . The mixosaurid ichthyosaur Phalarodon cf. P. fraasi from the Middle Triassic of Guizhou Province, China . Journal of Paleontology . 81 . 3. 602–605 . 10.1666/05092.1 . 2007JPal...81..602J . 54761545 .
34. Valentin Fischer . Nathalie Bardet . Myette Guiomar . Pascal Godefroit . amp . 2014 . High Diversity in Cretaceous Ichthyosaurs from Europe Prior to Their Extinction . PLOS ONE . 9 . 1 . e84709 . 10.1371/journal.pone.0084709 . 24465427 . 3897400. 2014PLoSO...984709F . free .
35. Campos. Lisandro. Fernández. Marta S.. Herrera. Yanina. Garrido. Alberto. 2021. Morphological disparity in the evolution of the ophthalmosaurid forefin: new clues from the Upper Jurassic of Argentina. Papers in Palaeontology. en. 7. 4. 1995–2020. 10.1002/spp2.1374. 2021PPal....7.1995C . 237770509. 2056-2802.
36. Megan L. Jacobs . David M. Martill . 2020 . A new ophthalmosaurid ichthyosaur from the Upper Jurassic (Early Tithonian) Kimmeridge Clay of Dorset, UK, with implications for Late Jurassic ichthyosaur diversity . PLOS ONE . 15 . 12 . e0241700 . 10.1371/journal.pone.0241700 . 33296370 . 7725355 . 2020PLoSO..1541700J . free .
37. Nadia B. Fröbisch . Jörg Fröbisch . P. Martin Sander . Lars Schmitz . Olivier Rieppel . amp . 2013 . Macropredatory ichthyosaur from the Middle Triassic and the origin of modern trophic networks . Proceedings of the National Academy of Sciences of the United States of America . 110. 4. 10.1073/pnas.1216750110 . 1393–1397 . 23297200 . 3557033. 2013PNAS..110.1393F . free .
38. Maisch. M. W.. Pan. X. R.. Sun. Z. Y.. Cai. T.. Zhang. D. P.. Xie. J. L.. 2006. Cranial osteology of Guizhouichthyosaurus tangae (Reptilia: Ichthyosauria) from the Upper Triassic of China. Journal of Vertebrate Paleontology. 26. 3. 588–597. 10.1671/0272-4634(2006)26[588:COOGTR]2.0.CO;2. 129117115 .
39. Shang. Q. H.. Li. C.. 2009. On the occurrence of the ichthyosaur Shastasaurus in the Guanling biota (Late Triassic), Guizhou, China. Vertebrata PalAsiatica. 47. 3. 178–193.