List of butterflies of Australia explained

Australia has more than 400 species of butterfly, the majority of which are continental species, and more than a dozen endemic species from remote islands administered by various Australian territorial governments. The largest butterflies in the world are endemic to the Australasian realm. They are the birdwings—Ornithoptera and other genera—of the tribe Troidini of the swallowtail butterfly family, Papilionidae.[1]

Papilionidae: swallowtails

Family: Papilionidae (swallowtails) – 18+2 species [*2 non-continental species]

Papilioninae

subfamily: Papilioninae

tribe: Leptocircini (formerly Graphiini)

genus: Protographium

P. l. leosthenes (Doubleday, 1846)

P. l. geimbia (Tindale, 1927)[3]

genus: Graphium (swordtails)

G. m. macleayanus (Leach, 1814)

G. m. moggana (Couchman, 1965)[6]

G. e. lycaon (C. & R. Felder, 1865)[8]

G. e. nyctimus (Waterhouse & Lyell, 1914)[9]

G. e. lycanoides (Rothschild, 1895)[10]

tribe: Papilionini (fluted swallowtails)

genus: Papilio (Linnaeus, 1758)

P. a. aegeus (Donovan, 1805)

P. a. ormenus (Guérin-Méneville, 1830)[14]

P. f. canopus (Westwood, 1842)[17]

P. f. capaneus (Westwood, 1843)[18]

P. f. indicatus (Butler, 1876)[19]

tribe: Troidini (Aristolochia-eating swallowtails)

genus: Cressida

C. c. cressida (Fabricius, 1775)

genus: Ornithoptera (birdwings)

O. p. poseidon (Doubleday, 1847)[23]

O. p. pronomus (Gray, 1853)

O. p. macalpinei (Moulds, 1974)[24]

genus: Pachliopta

Pieridae: whites and yellows

family: Pieridae (whites and yellows) – 35+2+1 species [†1 [[introduced species]]]

Pierinae: whites

subfamily: Pierinae (whites)

tribe: Elodinini

genus: Elodina (pearl-whites)

tribe: Leptosiaini

genus: Leptosia

tribe: Pierini

genus: Appias (albatrosses)

A. a. caria (Waterhouse & Lyell, 1914)[9]

A. a. albina (Boisduval, 1836)

A. p. ega (Boisduval, 1836)

genus: Pieris

genus: Belenois (formerly Anaphaeis)

B. j. peristhene (Boisduval, 1859)

B. j. teutonia (Fabricius, 1775)

genus: Cepora

C. p. scyllara (W. S. Macleay, 1826)

genus: Delias (Jezebels)

D. a. argenthona (Fabricius, 1793)

D. a. inferna (Butler, 1871)[32]

D. e. nigidius (Miskin, 1884)[34]

D. e. tindalii (Joicey & Talbot, 1926)[35]

D. m. mysis (Fabricius, 1775)

D. m. aestiva (Butler, 1897)[36]

D. m. onca (Fruhstorfer, 1910)[37]

D. n. nysa (Fabricius, 1775)

D. n. nivira (Waterhouse & Lyell, 1914)

Coliadinae: yellows

subfamily: Coliadinae (yellows)

genus: Catopsilia (migrants or emigrants)

C. p. crokera (W. S. Macleay, 1826)

C. s. etesia (Hewitson, 1867)[38]

C. g. gorgophone (Boisduval, 1836)

genus: Eurema (grass-yellows)

E. b. australis (Wallace, 1867)

E. b. zoraide (C. & R. Felder, 1865)

E. l. sana (Butler, 1877)

E. s. smilax (Donovan, 1805)

E. p. papuan (Butler, 1898)[40]

E. p. virgo (Wallace, 1867)

E. h. hecabe (Linnaeus, 1758)

E. h. phoebus (Butler, 1886)[42]

Riodinidae: metalmarks

family: Riodinidae (metalmarks) – 1 species

Nemeobiinae

subfamily: Nemeobiinae

genus: Praetaxila

P. s. punctaria (Fruhstorfer, 1914)[44]

Lycaenidae: gossamer-winged blues and coppers

family: Lycaenidae (gossamer-winged blues and coppers) – 142+7 species

Miletinae: harvesters

subfamily: Miletinae (harvesters)

tribe: Liphyrini

genus: Liphyra

L. b. major (Rothschild, 1898)[46]

Theclinae: hairstreaks

subfamily: Theclinae (hairstreaks)

tribe: Luciini

genus: Lucia

genus: Paralucia

P. p. lucida (Crosby, 1951)[49]

genus: Pseudodipsas

P. e. iole (Waterhouse & Lyell, 1914)

genus: Acrodipsas (ant-blues)

genus: Hypochrysops (jewels)

H. a. apelles (Fabricius, 1775)

H. a. apollo (Miskin, 1891)

H. a. phoebus (Waterhouse, 1928)[60]

H. d. delicia (Hewitson, 1875)

H. d. duaringae (Waterhouse, 1903)[63]

H. e. elgneri (Waterhouse & Lyell, 1909)

H. e. barnardi (Waterhouse, 1934)[65]

H. h. nebulosis (Sands, 1986)[67]

H. i. ignita (Leach, 1814)

H. i. chrysonotus (Grose-Smith, 1899)[68]

H. i. erythina (Waterhouse & Lyell, 1914)

H. i. oliffi (Miskin, 1889)

H. m. miskini (Waterhouse, 1903)

H. n. narcissus (Fabricius, 1775)

H. n. sabirus (Fruhstorfer, 1908)[69]

H. p. rovena (Druce, 1891)[71]

H. p. euclides (Miskin, 1889)

H. t. medocus (Fruhstorfer, 1908)

H. t. cretatus (Sands, 1986)

genus: Philiris (moonbeams)

P. d. diana (Waterhouse & Lyell, 1914)

P. d. papuanus (Wind & Clench, 1947)

P. f. kurandae (Waterhouse, 1903)

P. n. nitens (Grose-Smith, 1898)

P. n. lucina (Waterhouse & Lyell, 1914)

tribe: Arhopalini

genus: Arhopala (oak-blues)

A. m. amphis (Waterhouse, 1942)[77]

A. w. wildei (Miskin, 1891)

tribe: Ogyrini

genus: Ogyris (azures)

tribe: Zesiini (hairstreaks)

genus: Jalmenus

J. i. inous (Hewitson, 1865)

J. i. notocrucifer (Johnson, Hay & Bollam, 1992)[88]

genus: Pseudalmenus

P. c. chlorinda (Blanchard, 1948)

P. c. conara (Couchman, 1965)

P. c. zephyrus (Waterhouse & Lyell, 1914)

P. c. myrsilus (Westwood, 1851)

P. c. chloris (Waterhouse & Lyell, 1914)

P. c. barringtonensis (Waterhouse, 1928)

tribe: Hypolycaenini

genus: Hypolycaena

H. d. turneri (Waterhouse, 1903)

H. p. phorbas (Fabricius, 1793)

H. p. ingura (Tindale, 1923)[90]

tribe: Deudorigini

genus: Deudorix

D. e. dido (Waterhouse, 1934)

D. e. diovis (Hewitson, 1863)[92]

D. e. agimar (Fruhstorfer, 1908)

D. d. democles (Miskin, 1884)

D. s. dalyensis (le Souëf & Tindale, 1970)[94]

genus: Rapala

R. v. simsoni (Miskin, 1874)

genus: Bindahara

B. p. yurgama (Couchman, 1965)

Polyommatinae: blues

subfamily: Polyommatinae (blues)

tribe: Candalidini

genus: Candalides (pencil– and dusky-blues)

C. h. helenita (Semper, 1879)

C. m. margarita (Semper, 1879)

C. c. consimilis (Waterhouse, 1942)

C. c. goodingi (Tindale, 1965)[97]

C. c. toza (Kerr, 1967)[98]

C. c. cyprotus (Olliff, 1886)

C. c. pallescens (Tite, 1963)[100]

C. h. hyacinthina (Semper, 1879)

C. h. simplexa (Tepper, 1882)[101]

C. h. gilesi (Williams & Bollam, 2001)[102]

C. e. erinus (Fabricius, 1775)

C. h. heathi (Cox, 1873)

C. h. alpina (Waterhouse, 1928)

C. h. doddi (Burns, 1948)[106]

genus: Nesolycaena (opals)

tribe: Lycaenestheni

genus: Anthene (ciliate-blues)

A. s. affinis (Waterhouse & RE Turner, 1905)[112]

A. l. godeffroyi (Semper, 1879)

tribe: Polyommatini

genus: Petrelaea

genus: Nacaduba

N. b. berenice (Herrich-Schäffer, 1869)

N. k. parma (Waterhouse & Lyell, 1914)

N. k. felsina (Waterhouse & Lyell, 1914)

N. b. biocellata (C. & R. Felder, 1865)

N. c. calauria (C. Felder, 1860)

genus: Erysichton

E. l. lineata (Murray, 1874)

E. p. tasmanicus (Miskin, 1890)

genus: Danis

D. d. serapis (Miskin, 1891)

D. d. syrius (Miskin, 1890)

genus: Nothodanis

genus: Psychonotis

P. c. taygetus (C. & R. Felder, 1865)

genus: Prosotas

P. d. dubiosa (Semper, 1879)

P. n. auletes (Waterhouse & Lyell, 1914)

genus: Catopyrops

C. a. mysia (Waterhouse & Lyell, 1914)

C. f. halys (Waterhouse, 1934)

C. f. estrella (Waterhouse & Lyell, 1914)

genus: Ionolyce

I. h. hyllus (Waterhouse & Lyell, 1914)

genus: Theclinesthes

T. h. hesperia (Sibatani & Grund, 1978)

T. h. littoralis (Sibatani & Grund, 1978)

T. m. miskini (T. P. Lucas, 1889)

T. m. eucalypti (Sibatani & Grund, 1978)

T. m. arnoldi (Fruhstorfer, 1916)[118]

T. o. onycha (Hewitson, 1865)

T. o. capricornia (Sibatani & Grund, 1978)

T. s. serpentata (Herrich-Schäffer, 1869)

T. s. lavara (Couchman, 1954)[119]

genus: Sahulana

genus: Neolucia (heath-blues)

N. a. agricola (Westwood, 1851)

N. a. insulana (Waterhouse & Lyell, 1914)

N. a. occidens (Waterhouse & Lyell, 1914)

N. h. hobartensis (Miskin, 1890)

N. h. monticola (Waterhouse & Lyell, 1914)

genus: Jamides (ceruleans)

J. a. coelestis (Miskin, 1891)

J. c. claudia (Waterhouse & Lyell, 1914)

genus: Catochrysops (pea-blues)

C. a. amasea (Waterhouse & Lyell, 1914)

C. p. platissa (Herrich-Schäffer, 1869)

C. p. papuana (Tite, 1959)[121]

genus: Lampides

genus: Leptotes (synonyms Tarucus and Syntarucus)

L. p. pseudocassius (Murray, 1873)[123]

genus: Zizeeria

genus: Zizina

Z. l. labradus (Godart, 1824)

Z. l. labdalon (Waterhouse & Lyell, 1914)

genus: Famegana

Black-spotted grass-blue, Famegana alsulus (Herrich-Schäffer, 1869)

F. a. alsulus (Herrich-Schäffer, 1869)

genus: Zizula

Z. h. attenuata (T. P. Lucas, 1890)[127]

genus: Everes

E. l. australis (Couchman, 1962)[128]

genus: Pithecops

P. d. dionisius (Boisduval, 1832)

genus: Neopithecops

N. l. heria (Fruhstorfer, 1919)[129]

genus: Megisba

M. s. nigra (Miskin, 1890)

genus: Udara

genus: Euchrysops

E. c. cnidus (Waterhouse & Lyell, 1914)

genus: Freyeria

F. p. putli (Kollar, 1844)

Nymphalidae: brush– or four-footed

family: Nymphalidae (brush– or four-footed) – 81+6 species

Morphinae

subfamily: Morphinae

tribe: Amathusiini

genus Taenaris

T. a. jamesi (Butler, 1876)

Satyrinae

subfamily: Satyrinae

tribe: Biini

subtribe: Melanititi

genus: Melanitis

M. l. bankia (Fabricius, 1775)

tribe: Elymniini

subtribe: Elymniiti

genus: Elymnias

E. a. australiana (Fruhstorfer, 1900)[133]

subtribe: Mycalesiti

O. m. moira (Waterhouse & Lyell, 1914)

M. p. perseus (Fabricius, 1775)

M. s. sirius (Fabricius, 1775)

M. t. terminus (Fabricius, 1775)

tribe: Satyrini

subtribe: Ypthimiti

genus: Ypthima

Y. a. arctoa (Fabricius, 1775)

subtribe: Hypocystiti

genus: Hypocysta

H. a. angustata (Waterhouse & Lyell, 1914)

genus: Nesoxenicaendemic to Tasmania

N. l. leprea (Hewitson, 1864)

N. l. elia (Waterhouse & Lyell, 1914)

genus: Argynnina

genus: Oreixenica

genus: Geitoneura

genus: Heteronympha

genus: Tisiphone

Charaxinae: leafwings

subfamily: Charaxinae (leafwings)

genus: Charaxes

genus: Polyura

Polyura pyrrhus sempronius (Fabricius, 1793)

Apaturinae

subfamily: Apaturinae

genus: Apaturina

A. e. papuana (Ribbe, 1884)[148]

Heliconiinae: longwings

subfamily: Heliconiinae (longwings)

tribe: Acraeini

genus: Acraea

A. a. andromacha (Fabricius, 1775)

tribe: Heliconiini

genus: Cethosia

C. c. chrysippe (Fabricius, 1775)

C. p. paksha (Fruhstorfer, 1905)[149]

tribe: Vagrantini

genus: Vindula

V. a. ada (Butler, 1874)[150]

genus: Cupha

C. p. prosope (Fabricius, 1775)

genus: Vagrans

V. e. propinqua (Miskin, 1884)

genus: Phalanta

P. p. araca (Waterhouse & Lyell, 1914)

tribe: Argynnini

genus: Argynnis

A. h. inconstans Butler, 1873[151]

Nymphalinae

subfamily: Nymphalinae

genus: Doleschallia

D. b. australis (C. & R. Felder, 1867)[152]

genus: Hypolimnas

H. a. lamina (Fruhstorfer, 1903)[153]

H. a. darwinensis (Waterhouse & Lyell, 1914)

H. a. albula (Wallace, 1869)

H. b. bolina (Linnaeus, 1758)

H. b. nerina (Fabricius, 1775)

genus: Yoma

Yoma sabina parva (Butler, 1876)

genus: Junonia

Junonia hedonia zelima (Fabricius, 1775)

Junonia orithya albicincta (Butler, 1875)

Junonia villida calybe (Godart, 1819)[156]

genus: Vanessa

genus: Mynes

M. g. guerini (Wallace, 1869)

Biblidinae

subfamily: Biblidinae (planes)

genus: Pantoporia

P. c. consimilis (Boisduval, 1832)

P. v. moorei (W. J. Macleay, 1866)[158]

genus: Neptis

N. p. staudingereana (de Nicéville, 1898)[159]

genus: Phaedyma

P. s. shepherdi (Moore, 1858)

genus: Lexias

Libytheinae

subfamily: Libytheinae

genus: Libythea

L. g. genia (Waterhouse, 1938)[160]

L. g. nicevillei (Olliff, 1891)[161]

Danainae: milkweed butterflies

subfamily: Danainae (milkweed butterflies)

tribe: Danaini (tigers and crows)

genus Tirumala

T. h. hamata (W. S. Macleay, 1826)

genus Danaus

D. a. affinis (Fabricius, 1775)

D. a. alexis (Waterhouse & Lyell, 1914)

D. a. gelanor (Waterhouse & Lyell, 1914)

conspecific with:[162] D. philene (Stoll, 1782)

D. g. alexis (Waterhouse & Lyell, 1914)

D. c. petilia (Stoll, 1790)[163]

D. c. cratippus (C. Felder, 1860)

genus Euploea

E. a. eichhorni (Staudinger, 1884)[164]

E. a. enastri (Fenner, 1991)[165]

E. a. monilifera (Moore, 1883)[166]

E. c. corinna (W. S. Macleay, 1826)

E. a. violetta (Butler, 1876)

E. b. resarta (Butler, 1876)

E. c. macleari (Butler, 1887)[167]

E. d. darchia (W. S. Macleay, 1826)

E. d. niveata (Butler, 1875)

E. s. sylvester (Fabricius, 1793)

E. s. pelor (Doubleday, 1847)

E. t. tulliolus (Fabricius, 1793)

tribe: Tellervini

genus Tellervo

Cairns hamadryad, Tellervo zoilus zoilus (Fabricius, 1775)

Cape York hamadryad, Tellervo zoilus gelo (Waterhouse & Lyell, 1914)

Hesperiidae: skippers

family: Hesperiidae (skippers)[170]121+1 species

Pyrginae: spread-winged skippers

subfamily: Pyrginae (spread-winged skippers) (Burmeister, 1878)[171]

genus: dusk-flats, Chaetocneme (C. Felder, 1860)

C. c. sphinterifera (Fruhstorfer, 1910)

genus: Euschemon (Doubleday, 1846)

E. r. rafflesia (W. S. Macleay, 1826)

E. r. alba (Mabille, 1903)[174]

genus: Exometoeca

genus: Netrocoryne

N. r. repanda (C. & R. Felder, 1867)

N. r. expansa (Waterhouse, 1932)[176]

genus: Tagiades

T. j. janetta (Butler, 1870)[177]

Coeliadinae: awls, awlets and policemen

subfamily: Coeliadinae (awls, awlets and policemen)

genus: Allora

genus: Badamia

genus: Hasora

H. d. mastusia (Fruhstorfer, 1911)[180]

H. c. chromus (Cramer, 1780)

H. k. haslia (Swinhoe, 1899)[182]

Trapezitinae: Australian skippers

subfamily: Trapezitinae (Australian skippers)

genus: Antipodia

genus: Croitana

genus: Herimosa (Atkins, 1994);[188] previously Anisynta (Lower, 1911)[189]

A. c. cynone (Hewitson, 1874)

A. c. gunneda (Couchman, 1954)

genus: Hesperilla

genus: Mesodina

genus: Motasingha

genus: Neohesperilla

genus: Oreisplanus

genus: Signeta

genus: Toxidia

genus: Trapezites

Hesperinae: grass skippers

subfamily: Hesperinae (grass skippers)

genus: Notocrypta

N. w. proserpina (Butler, 1883)

genus: Taractrocera

T. i. ilia (Waterhouse, 1903)

genus: Ocybadistes

O. a. ardea (Bethune-Baker, 1906)

O. a. heterobathra (Lower, 1908)

genus: Suniana

genus: Arrhenes

A. d. iris (Waterhouse, 1903)

A. m. affinis (Waterhouse & Lyell, 1912)

genus: Telicota

T. a. krefftii (W. J. Macleay, 1866)

T. c. argeus (Plötz, 1883)[215]

T. m. mesoptis (Lower, 1911)

genus: Cephrenes

C. a. sperthias (C. Felder, 1862)

genus: Sabera

genus: Pelopidas

genus: Parnara

genus: Borbo

B. i. lavinia (Waterhouse, 1932)

B. i. tetragaphus (Mabille, 1891)

Remote islands species

See also: List of islands of Australia.

Australia has 8,222 islands within her maritime borders. This is small in comparison with her northern neighbour Indonesia, with about 18,300 islands (high geological activity is constantly adding and removing Indonesian islands). The British Isles include more than 6,000 islands over a much smaller area. The Greek islands include about as many islands depending on the minimum size to take into account, but in an even smaller area.

Christmas Island (105°E)

Torres Strait islands (142–144°E)

Dauan Island (Dauan, 142.5°E)
Saibai Island (Saibai, 142.7°E)
Yorke Island (Masig, 143.4°E)
Darnley Island (Erub, 143.7°E)
Murray Island (Mer, 144.0°E)

Norfolk Island (168°E)

Glossary

Major collections

Butterflies (Papilionoidea and Hesperioidea)
in principal collections
CollectionSpecimensAmateurPercent
ANIC, Canberra115,00092,00080%
Australian Museum, Sydney65,00063,90098%
Museum of Victoria24,00018,00075%
Queensland Museum9,0006,30070%
South Australian Museum36,00019,80055%
Total249,000200,00080%

Museums outside of Australia with significant Australian butterfly collections

See also

collections
butterfly lists
Australian animal lists

Bibliography

[Reviewed in Australian Journal of Entomology 40 (2001): 202–204.]

Taxonomic authorities

External links

Notes and References

  1. Book: Collins . N. Mark . Morris . Michael G. . Threatened Swallowtail Butterflies of the World: The IUCN Red Data Book . 1985 . . Gland & Cambridge . 978-2-88032-603-6 . Biodiversity Heritage Library.
  2. [Edward Doubleday]
  3. [Norman Barnett Tindale]
  4. [Pieter Cramer]
  5. [William Elford Leach]
  6. [Leonard Edgar Couchman]
  7. [Carl Linnaeus|Caroli Linnæi]
  8. [Baron Cajetan von Felder|Cajetan]
  9. [Gustavus Athol Waterhouse]
  10. [Walter Rothschild, 2nd Baron Rothschild|Walter Rothschild]
  11. [Arthur Gardiner Butler]
  12. [William Sharp Macleay]
  13. [Edward Donovan]
  14. [Félix Édouard Guérin-Méneville]
  15. [Jean Baptiste Boisduval|Jean Baptiste Alphonse de Chauffour de Boisduval]
  16. Goeze, 1779.
  17. [John Obadiah Westwood]
  18. [John Obadiah Westwood]
  19. [Arthur Gardiner Butler]
  20. [Jean Baptiste Boisduval|Jean Baptiste Alphonse de Chauffour de Boisduval]
  21. [Johan Christian Fabricius]
  22. Gray, 1853.
  23. [Edward Doubleday]
  24. [Max S. Moulds]
  25. [Carl Linnaeus]
  26. [Hippolyte Lucas]
  27. M. de Baar and DL Hancock (1993), "The Australian species of Elodina C. & R. Felder (Lepidoptera: Pieridae), Australian Entomological Magazine 20(1): 25–43.M. de Baar and DL Hancock (1993), "Further Notes on the Australian Species of Elodina C. & R. Felder (Lepidoptera: Pieridae)", Entomological Society of Queensland News Bulletin 21(8): 131–135.
  28. [William Chapman Hewitson]
  29. [William Henry Miskin]
  30. Swinhoe (1890).
  31. [Johan Christian Fabricius|Johannes Christophorus Fabricius]
  32. [Arthur Gardiner Butler]
  33. [Alfred Russel Wallace]
  34. [William Henry Miskin]
  35. Joicey & Talbot, 1926.
  36. [Arthur Gardiner Butler]
  37. [Hans Fruhstorfer]
  38. [William Chapman Hewitson]
  39. [Jean Baptiste Boisduval|Jean Baptiste Alphonse de Chauffour de Boisduval]
  40. [Arthur Gardiner Butler]
  41. [Baron Cajetan von Felder|Cajetan]
  42. [Arthur Gardiner Butler]
  43. [William Chapman Hewitson]
  44. [Hans Fruhstorfer|Fruhstorfer, Hans]
  45. Westwood, 1864.
  46. [Walter Rothschild, 2nd Baron Rothschild|Walter Rothschild]
  47. W Swainson (1833).
  48. CE Blanchard, 1848.
  49. Crosby, 1951.
  50. ED Edwards & Common, 1978.
  51. [William Chapman Hewitson]
  52. [Baron Cajetan von Felder|Cajetan]
  53. CG Miller & ED Edwards, 1978.
  54. Don PA Sands, 1997.
  55. Don PA Sands, 1965.
  56. Don PA Sands, 1980.
  57. Don PA Sands, CG Miller & JFR Kerr, 1997.
  58. [Gustavus Athol Waterhouse]
  59. [William Henry Miskin]
  60. [Gustavus Athol Waterhouse]
  61. Grose-Smith, 1900.
  62. [William Chapman Hewitson]
  63. [Gustavus Athol Waterhouse]
  64. [Gustavus Athol Waterhouse]
  65. [Gustavus Athol Waterhouse]
  66. [William Henry Miskin]
  67. Don PA Sands, 1986.
  68. Grose-Smith, 1899.
  69. [Hans Fruhstorfer]
  70. JFR Kerr, Jack Macqueen & Don PA Sands, 1969.
  71. [Herbert Druce|Herbert H Druce]
  72. RG Wind and Harry Kendon Clench, 1947.
  73. Grose-Smith & Kirby, 1897.
  74. [William Henry Miskin]
  75. Grose-Smith, 1898.
  76. [William Henry Miskin]
  77. [Gustavus Athol Waterhouse]
  78. [John Obadiah Westwood]
  79. [Gustavus Athol Waterhouse]
  80. [William Chapman Hewitson]
  81. [Gustavus Athol Waterhouse]
  82. Field (1999).
  83. Graham & Moulds, 1988.
  84. [Herbert Druce|Herbert H Druce]
  85. [Georg Semper]
  86. Staudinger (1888).
  87. [William Chapman Hewitson]
  88. Johnson, RW Hay & Bollam, 1992.
  89. JFR Kerr & Jack Macqueen, 1967.
  90. [Norman Barnett Tindale]
  91. Moore, 1858.
  92. [William Chapman Hewitson]
  93. [Baron Cajetan von Felder|Cajetan Felder]
  94. JC le Souef & Norman Barnett Tindale, 1970.
  95. Horsfield, 1829.
  96. [Baron Cajetan von Felder|Cajetan Felder]
  97. [Norman Barnett Tindale]
  98. JFR Kerr, 1967.
  99. [Arthur Sydney Olliff]
  100. Tite, 1963.
  101. JGO Tepper (1882).
  102. Williams & Bollam, 2001.
  103. ED Ewdards & Kerr, 1978.
  104. Cox, 1873.
  105. [Jacob Hübner]
  106. Burns, 1948.
  107. Michael F. Braby & Fabian Douglas (2004), "The taxonomy, ecology and conservation status of the Golden-rayed Blue: a threatened butterfly endemic to western Victoria", Australia Biological Journal of the Linnean Society 81(2): 275–299.
  108. Michael F. Braby, 1996.
  109. [Norman Barnett Tindale]
  110. John WC d'Apice and CG Miller (1992), "The genus Nesolycaena Waterhouse and Turner (Lepidoptera: Lycaenidae) with a description of a new species", Australian Entomological Magazine 19(3): 75–80 [75].
  111. Röber, 1886.
  112. [Gustavus Athol Waterhouse]
  113. Herrich-Schäffer, 1869.
  114. Murray, 1874.
  115. Eschscholtz, 1821.
  116. Sibatani & Grund, 1978.
  117. [Thomas Pennington Lucas]
  118. [Hans Fruhstorfer]
  119. [Leonard Edgar Couchman]
  120. Mathew, 1889.
  121. Tite, 1959.
  122. [Carl Linnaeus]
  123. Murray, 1873.
  124. Moore, 1865.
  125. Godart, 1824.
  126. [Johan Christian Fabricius]
  127. [Thomas Pennington Lucas]
  128. [Leonard Edgar Couchman]
  129. [Hans Fruhstorfer]
  130. [Johan Christian Fabricius]
  131. Kollar, 1844.
  132. Snellen, 1860.
  133. [Hans Fruhstorfer]
  134. [Jacob Hübner]
  135. [Arthur Gardiner Butler]
  136. [William Chapman Hewitson]
  137. [Arthur Sydney Olliff]
  138. [Edward Meyrick]
  139. [Leonard Edgar Couchman]
  140. Geyer, 1832.
  141. [Arthur Gardiner Butler]
  142. [Oswald Bertram Lower]
  143. [Gustavus Athol Waterhouse]
  144. [Gustavus Athol Waterhouse]
  145. [Arthur Sydney Olliff]
  146. [Arthur Gardiner Butler]
  147. [Arthur Gardiner Butler]
  148. Ribbe, 1884.
  149. [Hans Fruhstorfer]
  150. [Arthur Gardiner Butler]
  151. [Arthur Gardiner Butler]
  152. [Baron Cajetan von Felder|Cajetan]
  153. [Hans Fruhstorfer]
  154. Wallace, 1869.
  155. [Carl Linnaeus|Carolus Linnaeus]
  156. Godart, 1819.
  157. McCoy, 1868.
  158. [William John Macleay]
  159. de Nicéville, 1898.
  160. [Gustavus Athol Waterhouse]
  161. [Arthur Sydney Olliff]
  162. JA Edgar (1982), "Pyrrolizidine alkaloids sequestered by Solomon Island Danaine butterflies: the feeding preferences of the Danainae and Ithomiinae", Journal of the Zoological Society of London 196: 385–399 [393].
  163. Stoll (1790).
  164. Staudinger (1884).
  165. Fenner (1991).
  166. Moore, 1883.
  167. [Arthur Gardiner Butler]
  168. [Johann Friedrich Gmelin]
  169. Snellen, 1889.
  170. [Pierre André Latreille]
  171. [Hermann Burmeister|Karl Hermann Konrad Burmeister]
  172. [Félix Édouard Guérin-Méneville]
  173. [Edward Meyrick]
  174. [Jules Paul Mabille]
  175. [Edward Meyrick]
  176. [Gustavus Athol Waterhouse]
  177. [Arthur Gardiner Butler]
  178. [Walter Rothschild, 2nd Baron Rothschild|Walter Rothschild]
  179. [Baron Cajetan von Felder|Cajetan]
  180. [Hans Fruhstorfer]
  181. [Jules Paul Mabille]
  182. Swinhoe (1899).
  183. [Arthur Gardiner Butler]
  184. [William Chapman Hewitson]
  185. [Oswald Bertram Lower]
  186. Moore, 1878.
  187. ED Edwards, 1979.
  188. A. Atkins, "A new genus Herimosa (Lepidoptera: Hesperiidae: Trapezitinae) and its relationship to the Proeidosa group of endemic Australian skippers", Australian Entomologist 21 (1994): 143–152.
  189. [Oswald Bertram Lower]
  190. [Gustavus Athol Waterhouse]
  191. [Carl Plötz]
  192. [Gustavus Athol Waterhouse]
  193. Don PA Sands & JFR Kerr, 1973.
  194. Andrew F. Atkins, 1978.
  195. [Edward Meyrick]
  196. ED Edwards, 1987.
  197. ED Edwards & Graham, 1995.
  198. [William Chapman Hewitson]
  199. WF Kirby, 1893.
  200. [Arthur Gardiner Butler]
  201. [Jules Paul Mabille]
  202. A. A. E. Williams, M. R. Williams & R. W. Hay 1998.
  203. Andrew F. Atkins, 1997.
  204. JFR Kerr & Don PA Sands, 1970.
  205. [William Chapman Hewitson]
  206. [Jacob Hübner]
  207. Mayo & Atkins, 1992.
  208. [Carl Plötz]
  209. [George Thomas Bethune-Baker]
  210. [Pierre André Latreille]
  211. Lambkin & Donaldson, 1994.
  212. Heron, 1894.
  213. Rosenstock, 1885.
  214. [Carl Plötz]
  215. [Carl Plötz]
  216. [William Chapman Hewitson]
  217. Evans, 1866.
  218. Wallace, 1866.
  219. [Jules Paul Mabille]
  220. http://www.parksaustralia.gov.au/christmas/pub/butterfly-brochure.pdf Christmas Island National Park