Shorea is a large genus in family Dipterocarpaceae, and has been subject to substantial revision over the decades. Prior to 2022 the genus included nearly 200 genera.
Based on phylogenomic analyses showing that genus Shorea sensu Ashton (1982) was polyphyletic, in 2022 Ashton and Heckenhauer proposed reviving the former genera Doona Thwaites, Pentacme A.DC., and Richetia F.Heim, raising Shorea sections Anthoshorea and Neohopea to genus rank, and combining Shorea sections Brachyptera, Mutica, Ovalis, Pachycarpae, and Rubella into the new genus Rubroshorea, with two sections, Shorea and Barbata, remaining in genus Shorea.[1] Plants of the World Online currently recognizes Anthoshorea, Doona, Neohopea, Pentacme, Richetia, and Rubroshorea as separate genera.[2]
Plants of the World Online currently accepts 47 species.[2] Ashton et al. (2022) accept two sections, Barbata and Shorea.[1]
This is a complete listing of Shorea species accepted by Plants of the World Online in July 2019. The subgeneric classification follows Ashton (2004) and covers only species native to northern Borneo, with some Sri Lankan species added.[3] The timber groups classified according to the system proposed by Symington in the early 1940s.
| Species | Ashton (2004) | Groups | Accepted names per Plants of the World Online, October 2024 | |
|---|---|---|---|---|
| sections | subsections | |||
| Shorea affinis | Doona | Doona affinis | ||
| Shorea congestiflora | Doona | Doona congestiflora | ||
| Shorea cordifolia | Doona | Doona cordifolia | ||
| Shorea disticha | Doona | Doona disticha | ||
| Shorea megistophylla | Doona | Doona macrophylla | ||
| Shorea trapezifolia | Doona | Doona trapezifolia | ||
| Shorea zeylanica | Doona | Doona zeylanica | ||
| Shorea contorta | Pentacme | Pentacme contorta | ||
| Shorea siamensis | Pentacme | Pentacme siamensis | ||
| Shorea agami | Anthoshorea | white meranti group | Anthoshorea agami | |
| Shorea bracteolata | Anthoshorea | white meranti group | Anthoshorea bracteolata | |
| Shorea confusa | Anthoshorea | white meranti group | Anthoshorea confusa | |
| Shorea cordata | Anthoshorea | white meranti group | Anthoshorea cordata | |
| Shorea dealbata | Anthoshorea | white meranti group | Anthoshorea dealbata | |
| Shorea gratissima | Anthoshorea | white meranti group | Anthoshorea gratissima | |
| Shorea lamellata | Anthoshorea | white meranti group | Anthoshorea lamellata | |
| Shorea ochracea | Anthoshorea | white meranti group | Anthoshorea ochracea | |
| Shorea resinosa | Anthoshorea | white meranti group | Anthoshorea resinosa | |
| Shorea symingtonii | Anthoshorea | white meranti group | Anthoshorea symingtonii | |
| Shorea virescens | Anthoshorea | white meranti group | Anthoshorea virescens | |
| Shorea isoptera | Neohopea | selangan batu group | Neohopea isoptera | |
| Shorea asahi | Shorea | Barbata | selangan batu group | Shorea asahi |
| Shorea biawak | Shorea | Barbata | selangan batu group | Shorea biawak |
| Shorea ladiana | Shorea | Barbata | selangan batu group | Shorea ladiana |
| Shorea laevis | Shorea | Barbata | selangan batu group | Shorea laevis |
| Shorea maxwelliana | Shorea | Barbata | selangan batu group | Shorea maxwelliana |
| Shorea micans | Shorea | Barbata | selangan batu group | Shorea micans |
| Shorea atrinervosa | Shorea | Shorea | selangan batu group | Shorea atrinervosa |
| Shorea brunnescens | Shorea | Shorea | selangan batu group | Shorea brunnescens |
| Shorea calcicola | Shorea | Shorea | selangan batu group | Shorea calcicola |
| Shorea crassa | Shorea | Shorea | selangan batu group | Shorea crassa |
| Shorea domatiosa | Shorea | Shorea | selangan batu group | Shorea domatiosa |
| Shorea exelliptica | Shorea | Shorea | selangan batu group | Shorea exelliptica |
| Shorea falcifera | Shorea | Shorea | selangan batu group | Shorea falcifera |
| Shorea falciferoides | Shorea | Shorea | selangan batu group | Shorea falciferoides |
| Shorea foxworthyi | Shorea | Shorea | selangan batu group | Shorea foxworthyi |
| Shorea geniculata | Shorea | Shorea | selangan batu group | Shorea geniculata |
| Shorea guiso | Shorea | Shorea | selangan batu group | Shorea guiso |
| Shorea havilandii | Shorea | Shorea | selangan batu group | Shorea havilandii |
| Shorea hypoleuca | Shorea | Shorea | selangan batu group | Shorea hypoleuca |
| Shorea inappendiculata | Shorea | Shorea | selangan batu group | Shorea inappendiculata |
| Shorea lunduensis | Shorea | Shorea | selangan batu group | Shorea lunduensis |
| Shorea materialis | Shorea | Shorea | selangan batu group | Shorea materialis |
| Shorea obscura | Shorea | Shorea | selangan batu group | Shorea obscura |
| Shorea scrobiculata | Shorea | Shorea | selangan batu group | Shorea scrobiculata |
| Shorea seminis | Shorea | Shorea | selangan batu group | Shorea seminis |
| Shorea superba | Shorea | Shorea | selangan batu group | Shorea superba |
| Shorea almon | Brachypterae | red meranti group | Rubroshorea almon | |
| Shorea andulensis | Brachypterae | red meranti group | Rubroshorea andulensis | |
| Shorea bullata | Brachypterae | red meranti group | Rubroshorea bullata | |
| Shorea carapae | Brachypterae | red meranti group | Rubroshorea carapae | |
| Shorea coriacea | Brachypterae | red meranti group | Rubroshorea coriacea | |
| Shorea fallax | Brachypterae | red meranti group | Rubroshorea fallax | |
| Shorea flaviflora | Brachypterae | red meranti group | Rubroshorea flaviflora | |
| Shorea flemmichii | Brachypterae | red meranti group | Rubroshorea flemmichii | |
| Shorea inaequilateralis | Brachypterae | red meranti group | Rubroshorea inaequilateralis | |
| Shorea johorensis | Brachypterae | red meranti group | Rubroshorea johorensis | |
| Shorea kunstleri | Brachypterae | red meranti group | Rubroshorea kunstleri | |
| Shorea monticola | Brachypterae | red meranti group | Rubroshorea monticola | |
| Shorea pachyphylla | Brachypterae | red meranti group | Rubroshorea pachyphylla | |
| Shorea palembanica | Brachypterae | red meranti group | Rubroshorea palembanica | |
| Shorea parvistipulata | Brachypterae | red meranti group | Rubroshorea parvistipulata | |
| Shorea pauciflora | Brachypterae | red meranti group | Rubroshorea pauciflora | |
| Shorea platyclados | Brachypterae | red meranti group | Rubroshorea platyclados | |
| Shorea pubistyla | Brachypterae | red meranti group | Rubroshorea pubistyla | |
| Shorea scaberrima | Brachypterae | red meranti group | Rubroshorea scaberrima | |
| Shorea smithiana | Brachypterae | red meranti group | Rubroshorea smithiana | |
| Shorea venulosa | Brachypterae | red meranti group | Rubroshorea venulosa | |
| Shorea waltoni | Brachypterae | red meranti group | Rubroshorea waltoni | |
| Shorea acuta | Mutica | Auriculatae | red meranti group | Rubroshorea acuta |
| Shorea ferruginea | Mutica | Auriculatae | red meranti group | Rubroshorea ferruginea |
| Shorea macroptera | Mutica | Auriculatae | red meranti group | Rubroshorea macroptera |
| Shorea myrionerva | Mutica | Auriculatae | red meranti group | Rubroshorea myrionerva |
| Shorea sagittata | Mutica | Auriculatae | red meranti group | Rubroshorea sagittata |
| Shorea slootenii | Mutica | Auriculatae | red meranti group | Rubroshorea slootenii |
| Shorea argentifolia | Mutica | Mutica | red meranti group | Rubroshorea argentifolia |
| Shorea curtisii | Mutica | Mutica | red meranti group | Rubroshorea curtisii |
| Shorea dasyphylla | Mutica | Mutica | red meranti group | Rubroshorea dasyphylla |
| Shorea foraminifera | Mutica | Mutica | red meranti group | Rubroshorea foraminifera |
| Shorea hemsleyana | Mutica | Mutica | red meranti group | Rubroshorea hemsleyana |
| Shorea leprosula | Mutica | Mutica | red meranti group | Rubroshorea leprosula |
| Shorea macrantha | Mutica | Mutica | red meranti group | Rubroshorea macrantha |
| Shorea ovata | Mutica | Mutica | red meranti group | Rubroshorea ovata |
| Shorea pallidifolia | Mutica | Mutica | red meranti group | Rubroshorea pallidifolia |
| Shorea parvifolia | Mutica | Mutica | red meranti group | Rubroshorea parvifolia |
| Shorea platycarpa | Mutica | Mutica | red meranti group | Rubroshorea platycarpa |
| Shorea quadrinervis | Mutica | Mutica | red meranti group | Rubroshorea quadrinervis |
| Shorea retusa | Mutica | Mutica | red meranti group | Rubroshorea retusa |
| Shorea revoluta | Mutica | Mutica | red meranti group | Rubroshorea revoluta |
| Shorea rubra | Mutica | Mutica | red meranti group | Rubroshorea rubra |
| Shorea rugosa | Mutica | Mutica | red meranti group | Rubroshorea rugosa |
| Shorea scabrida | Mutica | Mutica | red meranti group | Rubroshorea scabrida |
| Shorea teysmanniana | Mutica | Mutica | red meranti group | Rubroshorea teysmanniana |
| Shorea uliginosa | Mutica | Mutica | red meranti group | Rubroshorea uliginosa |
| Shorea ovalis | Ovalis | red meranti group | Rubroshorea ovalis | |
| Shorea amplexicaulis | Pachycarpae | red meranti group | Rubroshorea amplexicaulis | |
| Shorea beccariana | Pachycarpae | red meranti group | Rubroshorea beccariana | |
| Shorea macrophylla | Pachycarpae | red meranti group | Rubroshorea macrophylla | |
| Shorea mecistopteryx | Pachycarpae | red meranti group | Rubroshorea mecistopteryx | |
| Shorea pilosa | Pachycarpae | red meranti group | Rubroshorea pilosa | |
| Shorea pinanga | Pachycarpae | red meranti group | Rubroshorea pinanga | |
| Shorea praestans | Pachycarpae | red meranti group | Rubroshorea praestans | |
| Shorea rotundifolia | Pachycarpae | red meranti group | Rubroshorea rotundifolia | |
| Shorea splendida | Pachycarpae | red meranti group | Rubroshorea splendida | |
| Shorea stenoptera | Pachycarpae | red meranti group | Rubroshorea stenoptera | |
| Shorea woodii | Pachycarpae | red meranti group | Rubroshorea woodii | |
| Shorea albida | Rubella | red meranti group | Rubroshorea albida | |
| Shorea dispar | Rubella | red meranti group | Rubroshorea dispar | |
| Shorea elliptica | Rubella | red meranti group | Rubroshorea elliptica | |
| Shorea rubella | Rubella | red meranti group | Rubroshorea rubella | |
| Shorea acuminatissima | Richetioides | yellow meranti group | Richetia acuminatissima | |
| Shorea alutacea | Richetioides | yellow meranti group | Richetia alutacea | |
| Shorea angustifolia | Richetioides | yellow meranti group | Richetia angustifolia | |
| Shorea bakoensis | Richetioides | yellow meranti group | Richetia bakoensis | |
| Shorea balanocarpoides | Richetioides | yellow meranti group | Richetia balanocarpoides | |
| Shorea chaiana | Richetioides | yellow meranti group | Richetia chaiana | |
| Shorea collaris | Richetioides | yellow meranti group | Richetia collaris | |
| Shorea cuspidata | Richetioides | yellow meranti group | Richetia cuspidata | |
| Shorea faguetiana | Richetioides | yellow meranti group | Richetia faguetiana | |
| Shorea faguetioides | Richetioides | yellow meranti group | Richetia faguetioides | |
| Shorea gibbosa | Richetioides | yellow meranti group | Richetia gibbosa | |
| Shorea hopeifolia | Richetioides | yellow meranti group | Richetia hopeifolia | |
| Shorea iliasii | Richetioides | yellow meranti group | Richetia iliasii | |
| Shorea induplicata | Richetioides | yellow meranti group | Richetia induplicata | |
| Shorea kudatensis | Richetioides | yellow meranti group | Richetia kudatensis | |
| Shorea laxa | Richetioides | yellow meranti group | Richetia laxa | |
| Shorea longiflora | Richetioides | yellow meranti group | Richetia longiflora | |
| Shorea longisperma | Richetioides | yellow meranti group | Richetia longisperma | |
| Shorea macrobalanos | Richetioides | yellow meranti group | Richetia macrobalanos | |
| Shorea mujongensis | Richetioides | yellow meranti group | Richetia mujongensis | |
| Shorea multiflora | Richetioides | yellow meranti group | Richetia multiflora | |
| Shorea obovoidea | Richetioides | yellow meranti group | Richetia obovoidea | |
| Shorea patoiensis | Richetioides | yellow meranti group | Richetia patoiensis | |
| Shorea peltata | Richetioides | yellow meranti group | Richetia peltata | |
| Shorea polyandra | Richetioides | yellow meranti group | Richetia polyandra | |
| Shorea richetia | Richetioides | yellow meranti group | Richetia coriacea | |
| Shorea subcylindrica | Richetioides | yellow meranti group | Richetia subcylindrica | |
| Shorea tenuiramulosa | Richetioides | yellow meranti group | Richetia tenuiramulosa | |
| Shorea xanthophylla | Richetioides | yellow meranti group | Richetia xanthophylla | |
| Shorea acuminata | Rubroshorea acuminata | |||
| Shorea altopoensis | Shorea altopoensis | |||
| Shorea arsorianoi | Shorea arsorianoi | |||
| Shorea assamica | Anthoshorea assamica | |||
| Shorea astylosa | Shorea astylosa | |||
| Shorea bentongensis | Anthoshorea bentongensis | |||
| Shorea blumutensis | Richetia blumutensis | |||
| Shorea ciliata | Shorea ciliata | |||
| Shorea collina | Shorea collina | |||
| Shorea conica | Richetia conica | |||
| Shorea dyeri | Shorea dyeri | |||
| Shorea falcata | Shorea falcata | |||
| Shorea farinosa | Anthoshorea farinosa | |||
| Shorea gardneri | Doona gardneri | |||
| Shorea glauca | Shorea glauca | |||
| Shorea henryana | Anthoshorea henryana | |||
| Shorea hulanidda | Anthoshorea hulanidda | |||
| Shorea hypochra | Anthoshorea hypochra | |||
| Shorea javanica | Anthoshorea javanica | |||
| Shorea kuantanensis | Richetia kuantanensis | |||
| Shorea lepidota | Rubroshorea lepidota | |||
| Shorea leptoderma | Shorea leptoderma | |||
| Shorea lissophylla | Shorea lissophylla | |||
| Shorea lumutensis | Shorea lumutensis | |||
| Shorea malibato | Shorea malibato | |||
| Shorea maxima | Richetia maxima | |||
| Shorea montigena | Anthoshorea montigena | |||
| Shorea negrosensis | Rubroshorea negrosensis | |||
| Shorea oblongifolia | Shorea oblongifolia | |||
| Shorea obtusa | Shorea obtusa | |||
| Shorea ochrophloia | Shorea ochrophloia | |||
| Shorea ovalifolia | Doona ovalifolia | |||
| Shorea pallescens | Shorea pallescens | |||
| Shorea palosapis | Rubroshorea palosapis | |||
| Shorea polita | Anthoshorea polita | |||
| Shorea polysperma | Rubroshorea polysperma | |||
| Shorea retinodes | Anthoshorea retinodes | |||
| Shorea robusta | Shorea robusta | |||
| Shorea roxburghii | Anthoshorea roxburghii | |||
| Shorea selanica | Rubroshorea selanica | |||
| Shorea singkawang | Rubroshorea singkawang | |||
| Shorea squamata | Rubroshorea palosapis | |||
| Shorea stipularis | Anthoshorea stipularis | |||
| Shorea submontana | Shorea submontana | |||
| Shorea sumatrana | Shorea sumatrana | |||
| Shorea thorelii | Shorea thorelii | |||
| Shorea tumbuggaia | Shorea tumbuggaia | |||
| Shorea wangtianshuea | Parashorea chinensis | |||
| Shorea worthingtonii | Doona venulosa | |||