List of New Zealand species extinct in the Holocene explained

This is a list of New Zealand species extinct in the Holocene that covers extinctions from the Holocene epoch, a geologic epoch that began about 11,650 years Before Present (about 9700 BCE) and continues to the present day.[1] This epoch equates with the latter third of the Haweran Stage of the Wanganui epoch in the New Zealand geologic time scale.

The North Island and South Island are the two largest islands of New Zealand. Stewart Island is the largest of the smaller islands. New Zealand proper also includes outlying islands such as the Chatham Islands, Kermadec Islands, and New Zealand Subantarctic Islands. Only New Zealand proper is represented on this list, not the Realm of New Zealand. For extinctions in the Cook Islands, Niue, or Tokelau, see the List of Oceanian animals extinct in the Holocene.

The islands of East Polynesia (including New Zealand, Hawaii, and Easter Island) were among the last habitable places on Earth colonised by humans.[2] [3] The first settlers of New Zealand migrated from Polynesia and became the Māori people.[4] According to archeological and genetic research, the ancestors of the Māori arrived in New Zealand no earlier than about 1280 CE, with at least the main settlement period between about 1320 and 1350,[5] consistent with evidence based on whakapapa (genealogical traditions).[6] [7] No credible evidence exists of pre-Māori settlement of New Zealand. In 1642, the Dutch navigator Abel Tasman became the first European explorer known to visit New Zealand.[8] In 1769, British explorer James Cook became the first European to map New Zealand and communicate with the Māori.[9] [10] From the late 18th century, the country was regularly visited by explorers and other sailors, missionaries, traders and adventurers. In 1840, the Treaty of Waitangi annexed New Zealand into the British Empire.[11] [12] [13] As a result of the influx of settlers, the population of Pākehā (European New Zealanders) grew explosively from fewer than 1,000 in 1831 to 500,000 by 1881.[14]

Numerous species have disappeared from New Zealand as part of the ongoing Holocene extinction, driven by human activity. Human contact, first by Polynesians and later by Europeans, had a significant impact on the environment. The arrival of the Māori resulted in animal extinctions due to deforestation and hunting.[15] The Māori also brought two species of land mammals, Polynesian rats (Rattus exulans) and kurī, a breed of domestic dog (Canis lupus familiaris).[16] In pre-human times, bats were the only land mammals found in New Zealand.[17] Polynesian rats definitely contributed to extinctions, and kurī might have contributed as well.[18] [19] Like the Māori settlers centuries earlier, the European settlers hunted native animals and engaged in habitat destruction. They also introduced numerous invasive species.[20] A few examples are black rats (Rattus rattus) and brown rats (Rattus norvegicus),[21] domestic cats (Felis catus),[22] stoats (Mustela erminea),[23] and common brushtail possums (Trichosurus vulpecula).[24]

This list of extinct species only includes the indigenous biota of New Zealand, not domestic animals like the kurī.

Mammals (class Mammalia)

Bats (order Chiroptera)

New Zealand short-tailed bats (family Mystacinidae)

Possibly extinct

Carnivorans (order Carnivora)

Eared seals (family Otariidae)

-- sortable-->"Common nameScientific nameRangeCommentsPictures
New Zealand sea lionPhocarctos hookeri 'NZ'Coastal North and South IslandDistinct lineage of the New Zealand sea lion once found all over the North and South Islands, as evidenced by ancient DNA studies on subfossil remains dating to just before 1280 CE.[26] It was hunted to extinction by the Maori within a few centuries. Afterwards, the subantarctic lineage of the New Zealand sea lion colonised Stewart Island and the southeastern coast of the South Island from the Auckland and Campbell Islands.[27]
Chatham Islands sea lionPhocarctos hookeri 'Chathams'Chatham IslandsDistinct lineage that became extinct due to overhunting within two centuries of Moriori settlement, in 1450-1650 CE.[28]

Birds (class Aves)

Moa (order Dinornithiformes)

Giant moa (family Dinornithidae)

Common nameScientific nameRangeCommentsPictures
North Island giant moaDinornis novaezealandiaeNorth Island and Great Barrier IslandMost recent bones in the Ruahine Range were dated to 1286–1390 CE.[29] The main cause of extinction was overhunting. Moa chicks may have also been eaten by Polynesian dogs.[30]
South Island giant moaDinornis robustusSouth Island, Stewart Island, and D'Urville IslandMost recent bones in Tumbledown Bay, Canterbury were dated to 1451–1952 CE (1558–1728 CE maximum likelihood), making this a contender for last surviving moa species.[31] They were hunted, and their bones are widespread in Māori middens, shaped into tools and ornaments. Estimates of moa remains in 1,200 open ovens and middens surveyed in the vicinity of the Waitaki River mouth during the 1930s range from 29,000 to 90,000. Moa chicks may have been eaten by Polynesian dogs. Burning of the giant moa's dry forest and shrubland habitat also likely reduced their numbers.[32]

Lesser moa (family Emeidae)

Common nameScientific nameRangeCommentsPictures
Bush moaAnomalopteryx didiformisNorth Island, South Island, and Stewart Island[33] Most recent bones in Echo Valley, Fiordland were dated to 1310-1420 CE. The main cause of extinction was overhunting. Moa chicks may have been eaten by Polynesian dogs.[34]
Eastern moaEmeus crassusSouth IslandMost recent eggshell fragments at Wairau Bar were dated to 1320–1350 CE.[35] The main cause of extinction was overhunting. Remains are widespread in middens, along with tools used to cut up carcasses and to work bones into tools. Middens in the Marlborough district contained remains of more than 4,000 individuals and large numbers of eggs. Eastern moa was the second most abundant species recorded after the broad-billed moa. Moa chicks may have also been eaten by Polynesian dogs. Burning of the eastern moa's forest and shrubland habitat is also likely to have reduced its numbers.[36]
Broad-billed moaEuryapteryx curtusNorth, South, Stewart, and Great Barrier Island[37] Most recent bones in Ototara, North Otago were dated to 1464–1637 CE, making this a contender for last surviving moa species. The main cause of extinction was overhunting. Remains are widespread in middens, along with specialised tools used to cut up carcasses and to work bones into tools. Moa chicks may have also been eaten by Polynesian dogs. Burning of the broad-billed moa's forest and shrubland habitat is also likely to have reduced its numbers.
Heavy-footed moaPachyornis elephantopusSouth Island and Stewart IslandMost recent bones in Kawarau Valley, Central Otago were dated to 1294–1438 CE. The main cause of extinction was overhunting. Moa chicks may also have been eaten by Polynesian dogs. Burning of eastern dryland forests and shrublands is also likely to have reduced the extent of suitable habitat.[38]
Mantell's moaPachyornis geranoidesNorth IslandMost recent remains were dated to 1278–1415 CE.[39] The main cause of extinction was overhunting. Moa chicks may also have been eaten by Polynesian dogs.[40]
Crested moaPachyornis australisSouth IslandSubalpine restricted species believed extinct during the Pleistocene-Holocene extinction around 10,000 years ago, until a partial skeleton from the Bulmer Cavern of Mount Owen was dated to 1396–1442 CE. The main cause of extinction was probably overhunting. Crested moa chicks may also have been eaten by Polynesian dogs.[41]

Upland moa (family Megalapterygidae)

Landfowl (order Galliformes)

Pheasants and allies (family Phasianidae)

Waterfowl (order Anseriformes)

Ducks, geese, and swans (family Anatidae)

Common nameScientific nameRangeCommentsPictures
Chatham duckAnas chathamicaChatham IslandsRemains dated to 448–657 CE, but believed to have survived until Polynesian arrival c. 1350. The species likely became extinct through overhunting, as it was large and flightless.[46]
New Zealand musk duckBiziura delautouriNorth Island and South IslandOne bone was found in a food midden. Hunting is the most likely cause of extinction, which happened before European arrival.[47]
Finsch's duckChenonetta finschiNorth Island and South Island[48] Extinct in the 16th century due to hunting and predation by Polynesian rats.
South Island gooseCnemiornis calcitransSouth IslandHunted by early Polynesian settlers, their remains are widespread in midden deposits. Overhunting is the most likely cause of extinction.[49] The date of extinction is unknown, but probably happened in the 16th century.
North Island gooseCnemiornis gracilisNorth IslandRemains dated to the first millennium CE,[50] but believed to have survived until Polynesian arrival c. 1280. A possible historical record, describing a rufous flightless goose that was hunted with dogs, was made in 1875. Though the reasons of extinction are unknown, any flightless goose would have been extremely vulnerable to overhunting.
Chatham Islands swanCygnus sumnerensis chathamicusChatham IslandsLikely hunted to extinction by the Moriori before 1650 CE.
New Zealand swanCygnus sumnerensis sumnerensisSouth IslandBones were found in Polynesian middens,[51] and dated to 1059-1401 CE. It was probably driven to extinction through overhunting before 1450 CE. This species was considered once synonymous with the Australian black swan which was later introduced to New Zealand by Europeans, but an ancient DNA study confirmed that they were different species that separated from a common ancestor 1–2 million years ago.
Scarlett's duckMalacorhynchus scarlettiNorth Island, South Island, and Chatham Islands[52] Became extinct sometime after Polynesian arrival. It was hunted by the Māori and its nests were possibly susceptible to predation by Polynesian rats.
New Zealand merganserMergus australisAuckland Islands, North Island, South Island, Stewart IslandReduced to the Auckland Islands only by the time of European colonisation, it was last recorded in 1902. Became extinct due to hunting and predation by introduced mammals.[53]
Chatham Island merganserMergus milleneriChatham IslandOnly known from bones found in middens. Hunting is the most likely cause of extinction.[54]
New Zealand stiff-tailed duckOxyura vantetsiNorth Island and South IslandExtinct before European contact. Hunting is the most likely cause of extinction, and a bone found in the South Island may have come from a human midden.[55]
Chatham Islands shelduckTadorna cf. variegataChatham IslandsAn "as-yet unnamed shelduck species was exterminated by the first Polynesian settlers" in the Chatham Islands.[56]

Owlet-nightjars (order Aegotheliformes)

Owlet-nightjars (family Aegothelidae)

Pigeons and doves (order Columbiformes)

Pigeons and doves (family Columbidae)

Rails and cranes (order Gruiformes)

Adzebills (family Aptornithidae)

Common nameScientific nameRangeCommentsPictures
South Island adzebillAptornis defossorSouth IslandMost recent bones were dated to 1234–1445 CE. The presence of adzebill bones in middens indicates that early Polynesian settlers hunted the species, and this is the most likely cause of extinction.[59] Nests could also have been raided by Polynesian rats.
North Island adzebillAptornis otidiformisNorth IslandMost recently dated to around 1000 CE, but believed to have survived until Polynesian arrival. The presence of adzebill bones in middens indicates that early Polynesian settlers hunted them, and this is the most likely cause of extinction.[60] Nests could also have been raided by Polynesian rats.

Rails (family Rallidae)

Common nameScientific nameRangeCommentsPictures
Chatham railCabalus modestusChatham IslandsLast recorded between 1893 and 1895. Extinct due to habitat destruction, competition with and predation by introduced mammals.[61]
Snipe-railCapellirallus karamuNorth IslandExcavations at Lake Poukawa, Hawke's Bay show that snipe-rails were once common but drastically reduced in numbers after Māori colonisation and the vegetation changed from podocarp forest to bracken and scrub, likely because of fire. However, the main factor of extinction was probably predation by Polynesian rats.[62]
Hawkins's railDiaphorapteryx hawkinsiChatham IslandsLast recorded in 1895. It was hunted to extinction.[63]
Chatham cootFulica chathamensisChatham Island[64] Remains dated to 701–119 BCE, but believed to have survived until Polynesian arrival c. 1350 CE. Probably became extinct due to overhunting and predation of eggs and chicks by Polynesian rats. Its bones are common in middens, indicating that it was frequently taken for food.
New Zealand cootFulica priscaNorth Island and South IslandExtinct due to overhunting by early Māori. Its bones are common in two archaeological middens in coastal Marlborough, where some coot bones were shaped into tools.[65] Nests could also have been raided by introduced predators.
Dieffenbach's railHypotaenidia dieffenbachiiChatham IslandsLast collected in 1840. It was possibly driven to extinction by introduced predators and habitat loss due to fire.[66]
North Island takahē Porphyrio mantelliNorth IslandRestricted to high altitude grasslands, this species declined with the expansion of forests in the Holocene but survived until hunting by the Maori drove it to extinction. There is a disputed historical observation from 1894.[67]
Hodgens' waterhenTribonyx hodgenorumNorth Island and South IslandDisappeared in the 17th century as a result of hunting and predation by Polynesian rats.[68]

Shorebirds (order Charadriiformes)

Sandpipers (family Scolopacidae)

Common nameScientific nameRangeCommentsPictures
North Island snipeCoenocorypha barrierensisLittle Barrier Island,[69] Browns Island (disputed), and North IslandSubfossil remains found across the North Island. A possible live individual was shot on Browns Island in 1820, and another was collected on Little Barrier Island in 1870. It is presumed extinct due to predation by Polynesian rats on the North Island and by feral cats introduced by Europeans on smaller islands.
Forbes's snipeCoenocorypha chathamicaChatham Island[70] and Pitt IslandPresumed extirpated by Polynesian rats on Chatham Island between 1500 and 1800 CE, and by feral cats on Pitt Island by the late 19th century.
South Island snipeCoenocorypha iredaleiSouth Island, Stewart Island, Jacky Lee Island, Big South Cape Island,[71] and possibly other offshore islandsExtirpated from the larger islands due to predation by Polynesian rats and from the smaller ones by black rats. The last population in Big South Cape Island disappeared in 1964 after a failed relocation attempt.

Albatrosses and petrels (order Procellariiformes)

Petrels and shearwaters (family Procellariidae)

Common nameScientific nameRangeComments
Imber's petrelPterodroma imberiChatham Islands[72] Probably disappeared from Chatham Island due to hunting and predation by Polynesian rats. It survived in Mangere Island until the 18th century, and in Pitt Island until the middle or late 19th century, when it was finished by feral cats.
Scarlett's shearwaterPuffinus spelaeusSouth IslandMost recently dated to 1350 CE.[73] Breeding colonies may have been overexploited directly, and their small size would have made eggs and chicks vulnerable to predation by Polynesian rats.[74]

Penguins (order Sphenisciformes)

Penguins (family Spheniscidae)

Common nameScientific nameRangeCommentsPictures
Chatham penguinEudyptes warhamiChatham Islands,[75] North Island, and South IslandLast dated after the 13th century. It was almost certainly extinct before Europeans arrived in the Chatham Islands;[76] a crested penguin captured alive in 1871 or 1872 was probably a vagrant of another species in the same genus.[77] [78]
Richdale's penguinMegadyptes antipodes richdaleiChatham Islands[79] Last dated after the 13th century. It was hunted to extinction.[80]
Waitaha penguinMegadyptes antipodes waitahaNorth Island, South Island,[81] Stewart Island, and Codfish Island / Whenua HouLast dated to 1347–1529 CE.[82] Archaeological remains indicate that early Polynesian settlers hunted the species and that this, with possible additional predation by Polynesian rats and dogs, was a probable cause of extinction. After its extinction, the subantarctic subspecies of the yellow-eyed penguin, M. a. antipodes, colonised Stewart Island and part of the South Island from the Auckland and Campbell Islands.

Boobies, cormorants, and allies (order Suliformes)

Cormorants and shags (family Phalacrocoracidae)

Pelicans, herons, and ibises (order Pelecaniformes)

Herons (family Ardeidae)

Hawks and relatives (order Accipitriformes)

Hawks, eagles, kites, harriers and Old World vultures (family Accipitridae)

Common nameScientific nameRangeCommentsPictures
Eyles's harrierCircus teauteensisNorth Island and South IslandLast dated to the first millennium CE but believed to have survived until Polynesian arrival c. 1280. There is a dubious observation from the 1870s. It is thought to have become extinct due to a combination of human-induced impacts: its forest and shrubland habitat was burned off and replaced with grassland, predation by introduced Polynesian rats probably caused a decline in its prey species, and hunting by humans is apparent from the presence of harrier bones in middens, and use of its bones to make tools. The Polynesian rat and dog may have also preyed on chicks. Unlike the closely related swamp harrier which colonised New Zealand after its extinction,[87] it was probably too heavy and small-winged to hunt over open grassland areas.[88]
Haast's eagleHieraaetus mooreiSouth Island[89] and Stewart IslandMost recent eggshell fragments at Wairau Bar were dated to 1320–1350 CE. It became extinct around the same time as all moa species. Overhunting of its moa prey was probably the main cause of its extinction. Loss of habitat due to the burning of dry mosaic forests and shrublands may also have caused declines in its prey species. Haast's eagle may also have been hunted because its bones, some of which were found in middens after being worked into tools. Nests could also have been raided by feral pigs and rats. Two claimed sightings from the 1800s are unlikely to have been Haast's eagle.

Owls (order Strigiformes)

True owls (family Strigidae)

Parrots (order Psittaciformes)

Old World parrots (family Psittaculidae)

Perching birds (order Passeriformes)

New Zealand wrens (family Acanthisittidae)

Common nameScientific nameRangeCommentsPictures
Lyall's wrenTraversia lyalliStephens Island, North Island, and South IslandDisappeared from the main islands after Maori settlement, likely because of habitat loss and predation by Polynesian rats. It lingered on Stephens Island until 1895, when it was exterminated by domestic cats.[93] [94]
BushwrenXenicus longipesNorth Island, South Island, Stewart Island, Kapiti Island, Big South Cape Island, Solomon Island and Pukeweka offshore Stewart Island[95] Extinct since 1972 due to predation by feral cats, rats, weasels, and stoats.
North Island stout-legged wrenPachyplichas jagmiNorth IslandExtinct after Māori settlement but before European contact.[96] [97]
South Island stout-legged wrenPachyplichas yaldwyniSouth IslandExtinct after Māori settlement but before European contact.
Long-billed wrenDendroscansor decurvirostrisSouth IslandExtinct shortly after Māori settlement.[98] Its flightlessness and probable ground-nesting habits would have made it easy prey for Polynesian rats.

Old World orioles (family Oriolidae)

Common nameScientific nameRangeCommentsPictures
North Island piopioTurnagra tanagraNorth IslandLast known individual killed at Ōhura, south Waikato, in 1902. Unconfirmed sightings continued into the 1970s, mainly from forest behind Whanganui, inland Taranaki and Te Urewera. It is likely that predation by introduced black rats was the main cause of extinction, though birds were occasionally eaten by settlers, and their final disappearance coincided with the spread of stoats.[101]
South Island piopioTurnagra capensis capensisSouth Island and some Fiordland islands[102] Declined rapidly following European settlement, especially after 1870. It was extinct on Banks and Otago Peninsulas by the 1880s, and on the West Coast and Fiordland by c. 1895. This coincided with the spread of introduced black rats and later, stoats. Unconfirmed sightings persisted into the 1930s, with the last in west Otago in 1963. Suggestions to transfer South Island piopio to an offshore sanctuary such as Kapiti Island or Little Barrier Island never eventuated, as live capture was very difficult before the invention of modern mist nets.
Stephens Island piopioTurnagra capensis minorStephens IslandLast collected in 1897. Became extinct following clearance of the island's forest and introduction of cats by lighthouse keepers.

Crows and relatives (family Corvidae)

Common nameScientific nameRangeCommentsPictures
New Zealand ravenCorvus antipodumExtinct before European contact. The presence of bones in middens shows that they were eaten by humans, and they may have been impacted by the rapid extirpation of mainland seal and seabird colonies following human arrival.
Chatham ravenCorvus moriorumChatham Islands[104] Dated to 8134–7408 BC but believed to have survived until 1350 BC. It became extinct before European contact. The presence of raven bone remains in middens shows that they were eaten by humans. However, it is equally likely that the extinction was part of a trophic cascade caused by human hunting of breeding seals and seabirds in the islands.[105]

New Zealand wattlebirds (family Callaeidae)

Possibly extinct, New Zealand wattlebirds (family Callaeidae)

Grassbirds and allies (family Locustellidae)

Reptiles (class Reptilia)

Squamates (order Squamata)

Skinks (family Scincidae)

Amphibians (class Amphibia)

Frogs (order Anura)

New Zealand primitive frogs (family Leiopelmatidae)

Common nameScientific nameRangeCommentsPictures
Aurora frogLeiopelma auroraensisFiordland, South IslandKnown from subfossil bones. Probably disappeared within the past 1,000 years due to predation by the Polynesian rat.[116]
Markham's frogLeiopelma markhamiNorth Island and South IslandLast dated to 1650 CE. Probably disappeared due to predation by the Polynesian rat.
Waitomo frogLeiopelma waitomoensisNorth IslandLast dated to 220–320 CE, but believed to have survived until the past 1,000 years. Probably disappeared due to predation by the Polynesian rat.

Ray-finned fish (class Actinopterygii)

Smelts (order Osmeriformes)

Australia-New Zealand smelts and graylings (family Retropinnidae)

Insects (class Insecta)

Beetles (order Coleoptera)

Ground beetles (family Carabidae)

Clitellates (class Clitellata)

Order Opisthopora

Family Megascolecidae

Plants (kingdom Plantae)

Order Brassicales

Mustard and crucifer family (family Brassicaceae)

Common nameScientific nameRangeComments
Waitakere scurvy grassLepidium amissumCoastal Waitākere Ranges, North Island[120] Last collected in 1917. It was already uncommon then and wasn't described as a species until 2013, from herbarium specimens. The reasons for extinction are unclear, but might be related to habitat alteration.[121]
Lepidium obtusatumCoastal Waitākere Ranges and Wellington, North Island[122] [123] Last seen in Waitākere in 1917 and in Wellington in 1950. The Wellington population seems to have been eliminated through a combination of habitat destruction as a consequence of gravel extraction, weed invasion, and over collection by botanists. It is not clear why it disappeared from the Waitākere coastline.

Order Santalales

Showy mistletoes (family Loranthaceae)

Order Caryophyllales

Pink and carnation family (family Caryophyllaceae)

Order Gentianales

Family Loganiaceae

Order Boraginales

Borage and forget-me-not family (family Boraginaceae)

See also

Further reading

External links

Notes and References

  1. Walker . Mike . Johnsen . Sigfus . Rasmussen . Sune Olander . Popp . Trevor . Steffensen . Jorgen-Peder . Gibrard . Phil . Hoek . Wim . Lowe . John . Andrews . John . Bjo Rck . Svante . Cwynar . Les C. . 2009 . Formal definition and dating of the GSSP (Global Stratotype Section and Point) for the base of the Holocene using the Greenland NGRIP ice core, and selected auxiliary records . live . . 24 . 1 . 3–17 . 2009JQS....24....3W . 10.1002/jqs.1227 . https://web.archive.org/web/20131104131948/http://www.stratigraphy.org/GSSP/Holocene.pdf . 2013-11-04 . 2022-04-24 . free . Konrad . Hughen . Peter . Kersahw . Bernd . Kromer . Thomas . Litt . David J. . Lowe . Takeshi . Nakagawa . Rewi . Newnham . Jakob . Schwander.
  2. Wilmshurst . Janet M. . Hunt . Terry L. . Lipo . Carl P. . Anderson . Atholl J. . December 27, 2010 . High-precision radiocarbon dating shows recent and rapid initial human colonization of East Polynesia . Proceedings of the National Academy of Sciences . en . 108 . 5 . 1815–1820 . 10.1073/pnas.1015876108 . 0027-8424 . 3033267 . 21187404 . free .
  3. Wilmshurst . Janet M. . Anderson . Atholl J. . Higham . Thomas F. G. . Worthy . Trevor H. . 2008-06-03 . Dating the late prehistoric dispersal of Polynesians to New Zealand using the commensal Pacific rat . Proceedings of the National Academy of Sciences . en . 105 . 22 . 7676–7680 . 10.1073/pnas.0801507105 . 0027-8424 . 2409139 . 18523023. free . 2008PNAS..105.7676W .
  4. Walters . Richard . Buckley . Hallie . Jacomb . Chris . Matisoo-Smith . Elizabeth . 7 October 2017 . Mass Migration and the Polynesian Settlement of New Zealand . Journal of World Prehistory . 30 . 4 . 351–376 . 10.1007/s10963-017-9110-y . free.
  5. Jacomb . C. . Holdaway . R. N. . Allentoft . M. E. . Bunce . M. . Oskam . C. L. . Walter . R. . Brooks . E. . 2014 . High-precision dating and ancient DNA profiling of moa (Aves: Dinornithiformes) eggshell documents a complex feature at Wairau Bar and refines the chronology of New Zealand settlement by Polynesians . Journal of Archaeological Science . 50 . 24–30 . 10.1016/j.jas.2014.05.023 . 2014JArSc..50...24J . 0305-4403.
  6. Roberton . J. B. W. . 1956 . Genealogies as a basis for Maori chronology . Journal of the Polynesian Society . 65 . 1 . 45–54.
  7. Te Hurinui . Pei . 1958 . Maori genealogies . Journal of the Polynesian Society . 67 . 2 . 162–165.
  8. Encyclopedia: European discovery of New Zealand – Abel Tasman . . New Zealand . 16 January 2010 . Wilson . John . 'On 13 December 1642 the Dutch sighted "a large land, uplifted high" – probably the Southern Alps ...'.
  9. Book: Salmond, Anne . Two worlds : first meetings between Māori and Europeans, 1642–1772 . 1991 . Viking . 0-670-83298-7 . Auckland, N.Z. . 26545658.
  10. Cook's Journal, 7 October 1769, National Library of Australia, http://southseas.nla.gov.au/journals/cook/17691007.html, visited 20120409
  11. Web site: Treaty events 1800–49 – Treaty timeline NZHistory, New Zealand history online . 26 July 2020 . nzhistory.govt.nz . Ministry for Culture and Heritage.
  12. Encyclopedia: Treaty of Waitangi – Interpretations of the Treaty of Waitangi . Te Ara: The Encyclopedia of New Zealand . 13 August 2019 . 20 June 2012 . Claudia Orange . Orange . Claudia.
  13. Encyclopedia: Honouring the treaty – 1940 to 2000s . . 26 July 2020 . 20 June 2012 . en-NZ . Orange . Claudia.
  14. James Belich, Making Peoples (1996) 278-80
  15. Web site: Yirka . Bob . Phys.org . DNA evidence suggests humans hunted moa to extinction . 2022-05-20 . phys.org . en.
  16. Web site: Taonga . New Zealand Ministry for Culture and Heritage Te Manatu . Kurī – Polynesian dogs . 2022-05-20 . teara.govt.nz . en.
  17. Web site: Bats/pekapeka . 2022-05-20 . www.doc.govt.nz . en-nz.
  18. Web site: Gone to the dogs . 2022-05-20 . New Zealand Geographic . en-NZ.
  19. Web site: The dog is in the henhouse: did the kurī (Polynesian dog) have an impact on New Zealand's wildlife? . 2022-05-20 . Sciblogs . en-US.
  20. Web site: Animal pests and threats A – Z . 2022-05-20 . www.doc.govt.nz . en-nz.
  21. Web site: Rats . 2022-05-20 . www.doc.govt.nz . en-nz.
  22. Web site: Feral cats . 2022-05-20 . www.doc.govt.nz . en-nz.
  23. Web site: Stoats . 2022-05-20 . www.doc.govt.nz . en-nz.
  24. Web site: Possums . 2022-05-20 . www.doc.govt.nz . en-nz.
  25. O'Donnell . C. . 15 October 2020 . Mystacina robusta . e.T14260A22070387 . 10.2305/IUCN.UK.2021-1.RLTS.T14260A22070387.en . 22 May 2022.
  26. Collins, C. J., Rawlence, N. J., Worthy, T. H., Scofield, R. P., Tennyson, A. J. D., Smith, I., ... & Waters, J. M. (2014). Pre-human New Zealand sea lion (Phocarctos hookeri) rookeries on mainland New Zealand. Journal of the Royal Society of New Zealand, 44(1), 1–16.
  27. Collins, C. J., Rawlence, N. J., Prost, S., Anderson, C. N., Knapp, M., Scofield, R. P., ... & Waters, J. M. (2014). Extinction and recolonization of coastal megafauna following human arrival in New Zealand. Proceedings of the Royal Society B: Biological Sciences, 281(1786), 20140097.
  28. Rawlence, N. J., Collins, C. J., Anderson, C. N., Maxwell, J. J., Smith, I. W., Robertson, B. C., ... & Waters, J. M. (2016). Human‐mediated extirpation of the unique Chatham Islands sea lion and implications for the conservation management of remaining New Zealand sea lion populations. Molecular ecology, 25(16), 3950–3961.
  29. Wood, J. R., and J. M. Wilmshurst. "Age of North Island giant moa (Dinornis novaezealandiae) bones found on the forest floor in the Ruahine Range." Journal of the Royal Society of New Zealand 43.4 (2013): 250–255.
  30. Web site: North Island giant moa New Zealand Birds Online . 2022-05-17 . www.nzbirdsonline.org.nz.
  31. Rawlence, N. J., and A. Cooper. "Youngest reported radiocarbon age of a moa (Aves: Dinornithiformes) dated from a natural site in New Zealand." Journal of the Royal Society of New Zealand 43.2 (2013): 100–107.
  32. Web site: South Island giant moa New Zealand Birds Online . 2022-05-17 . nzbirdsonline.org.nz.
  33. Book: . Checklist of the Birds of New Zealand, Norfolk and Macquarie Islands, and the Ross Dependency, Antarctica . 4th . 2010 . Wellington, New Zealand . Te Papa Press in association with the Ornithological Society of New Zealand . 978-1-877385-59-9 . New Zealand Birds Online . 17 May 2022.
  34. Web site: Little bush moa New Zealand Birds Online . 2022-05-18 . www.nzbirdsonline.org.nz.
  35. Jacomb, Chris, et al. "High-precision dating and ancient DNA profiling of moa (Aves: Dinornithiformes) eggshell documents a complex feature at Wairau Bar and refines the chronology of New Zealand settlement by Polynesians." Journal of Archaeological Science 50 (2014): 24–30
  36. Web site: Eastern moa New Zealand Birds Online . 2022-05-18 . nzbirdsonline.org.nz.
  37. Web site: Stout-legged moa New Zealand Birds Online . 2022-05-18 . www.nzbirdsonline.org.nz.
  38. Web site: Heavy-footed moa New Zealand Birds Online . 2022-05-18 . nzbirdsonline.org.nz.
  39. Book: Turvey, Sam. Holocene extinctions. 2009. Oxford University Press. 978-0-19-953509-5. 29 February 2012.
  40. Web site: Mantell's moa New Zealand Birds Online . 2022-05-18 . www.nzbirdsonline.org.nz.
  41. Web site: Crested moa New Zealand Birds Online . 2022-05-18 . nzbirdsonline.org.nz.
  42. Web site: Upland moa New Zealand Birds Online . 2022-05-18 . www.nzbirdsonline.org.nz.
  43. Hume, J.P. (2017) Extinct Birds. Bloomsbury Publishing, 560 pages.
  44. https://recentlyextinctspecies.com/galliformes-francolins-junglefowl-mound-builders-pheasants-turkeys-etc/megapodius-sp-nov-raoul-island Recently Extinct Species
  45. Coturnix novaezelandiae. e.T22678955A92795779. BirdLife International. 2016. 2016. 17 May 2022. 10.2305/IUCN.UK.2016-3.RLTS.T22678955A92795779.en.
  46. Web site: Chatham Island duck New Zealand Birds Online . 2022-05-18 . www.nzbirdsonline.org.nz.
  47. Web site: New Zealand musk duck New Zealand Birds Online . 2022-05-18 . www.nzbirdsonline.org.nz.
  48. BirdLife International . 2017 . Chenonetta finschi . 2017 . e.T62239833A119205634 . 10.2305/IUCN.UK.2017-3.RLTS.T62239833A119205634.en . 17 May 2022.
  49. Web site: South Island goose New Zealand Birds Online . 2022-05-18 . nzbirdsonline.org.nz.
  50. Wood, J.R., Scofield, R.P., Hamel, J., Lalas, C., & Wilmshurst, J.M. (2017). Bone stable isotopes indicate a high trophic position for New Zealand's extinct South Island adzebill (Aptornis defossor) (Gruiformes: Aptornithidae). New Zealand Journal of Ecology, 41(2), 240–244.
  51. Rawlence . Nicolas J. . Kardamaki . Afroditi . Easton . Luke J. . Tennyson . Alan J. D. . Scofield . R. Paul . Waters . Jonathan M. . 2017 . Ancient DNA and morphometric analysis reveal extinction and replacement of New Zealand's unique black swans . Proceedings of the Royal Society B: Biological Sciences . en . 284 . 1859 . 20170876 . 10.1098/rspb.2017.0876 . 0962-8452 . 5543223 . 28747476.
  52. Web site: Scarlett's duck New Zealand Birds Online . 2022-05-17 . nzbirdsonline.org.nz.
  53. BirdLife International . 2016 . Mergus australis . 2016 . e.T22680496A92864737 . 10.2305/IUCN.UK.2016-3.RLTS.T22680496A92864737.en . 17 May 2022.
  54. Web site: Chatham Island merganser New Zealand Birds Online . 2022-05-17 . nzbirdsonline.org.nz.
  55. Web site: New Zealand blue-billed duck New Zealand Birds Online . 2022-05-18 . nzbirdsonline.org.nz.
  56. Web site: Paradise shelduck Pūtangitangi New Zealand Birds Online . 2023-03-30 . www.nzbirdsonline.org.nz.
  57. Holdaway, R.N., Jones, M.D., & Athfield, N.R.B. (2002) Late Holocene extinction of the New Zealand owlet‐nightjar Aegotheles novaezealandiae. Journal of The Royal Society of New Zealand, 32(4), 653–667.
  58. Web site: New Zealand owlet-nightjar New Zealand Birds Online . 2022-05-17 . www.nzbirdsonline.org.nz.
  59. Web site: South Island adzebill New Zealand Birds Online . 2022-05-17 . www.nzbirdsonline.org.nz.
  60. Web site: North Island adzebill New Zealand Birds Online . 2022-05-17 . nzbirdsonline.org.nz.
  61. BirdLife International . 2016 . Cabalus modestus . 2016 . e.T22728873A94999473 . 10.2305/IUCN.UK.2016-3.RLTS.T22728873A94999473.en . 11 November 2021.
  62. Web site: Snipe-rail New Zealand Birds Online . 2022-05-17 . nzbirdsonline.org.nz.
  63. BirdLife International . 2017 . Diaphorapteryx hawkinsi . 2017 . e.T22733394A119260892 . 10.2305/IUCN.UK.2017-3.RLTS.T22733394A119260892.en . 17 May 2022.
  64. Web site: Chatham Island coot New Zealand Birds Online . 2022-05-17 . nzbirdsonline.org.nz.
  65. Web site: New Zealand coot New Zealand Birds Online . 2022-05-17 . nzbirdsonline.org.nz.
  66. BirdLife International . 2016 . Hypotaenidia dieffenbachii . 2016 . e.T22692455A93354540 . 10.2305/IUCN.UK.2016-3.RLTS.T22692455A93354540.en . 16 November 2021.
  67. BirdLife International . 2016 . Porphyrio mantelli . 2016 . e.T22728833A94998264 . 10.2305/IUCN.UK.2016-3.RLTS.T22728833A94998264.en . 17 May 2022.
  68. BirdLife International . 2017 . Tribonyx hodgenorum . 2017 . e.T62274163A119207810 . 10.2305/IUCN.UK.2017-3.RLTS.T62274163A119207810.en . 17 May 2022.
  69. BirdLife International . 2016 . Coenocorypha barrierensis . 2016 . e.T62178398A95191768 . 10.2305/IUCN.UK.2016-3.RLTS.T62178398A95191768.en . 18 May 2022.
  70. Web site: Forbes' snipe New Zealand Birds Online . 2022-05-19 . www.nzbirdsonline.org.nz.
  71. Web site: South Island snipe New Zealand Birds Online . 2022-05-19 . www.nzbirdsonline.org.nz.
  72. Web site: Imber's petrel New Zealand Birds Online . 2022-05-20 . www.nzbirdsonline.org.nz.
  73. Rando, J.C., & Alcover, J.A. (2008) Evidence for a second western Palaearctic seabird extinction during the last Millennium: the Lava Shearwater Puffinus olsoni. Ibis, 150(1), 188–192.
  74. Web site: Scarlett's shearwater New Zealand Birds Online . 2022-05-20 . www.nzbirdsonline.org.nz.
  75. Cole. Theresa L.. Ksepka. Daniel T.. Mitchell. Kieren J.. Tennyson. Alan J. D.. Thomas. Daniel B.. Pan. Hailin. Zhang. Guojie. Rawlence. Nicolas J.. Wood. Jamie R.. Bover. Pere. Bouzat. Juan L.. Cooper. Alan. Fiddamanl. Steven. Hart. Tom. Miller. Gary. Ryan. Peter G.. Shepherd. Lara D.. Wilmshurst. Janet M.. Waters. Jonathan M.. 2019. Mitogenomes uncover extinct penguin taxa and reveal island formation as a key driver of speciation. Molecular Biology and Evolution. en. 36. 4. 784–797. 10.1093/molbev/msz017. 30722030. free.
  76. Millener . P. R. . 1999 . The history of the Chatham Islands' bird fauna of the last 7000 years – a chronicle of change and extinction. Proceedings of the 4th International meeting of the Society of Avian Paleontology and Evolution (Washington, D.C., June 1996) . Smithsonian Contributions to Paleobiology . 89 . 85–109 .
  77. Tennyson. A.J.D.. Millener. P.R.. 1994. Bird extinctions and fossil bones from Mangere Island, Chatham Islands. Notornis. 41. supplement. 165–178.
  78. Miskelly. Colin M.. Bell. Mike. 2004. An unusual influx of Snares crested penguins (Eudyptes robustus) on the Chatham Islands, with a review of other crested penguin records from the islands. Notornis. 51. 4. 235–237.
  79. Miskelly, Colin & Forsdick, Natalie & Gill, Brian & Palma, Ricardo & Rawlence, Nicolas & Tennyson, Alan. (2022). CHECKLIST OF THE BIRDS OF NEW ZEALAND. https://www.researchgate.net/publication/361824003_CHECKLIST_OF_THE_BIRDS_OF_NEW_ZEALAND
  80. Cole, Theresa L., et al. "Mitogenomes uncover extinct penguin taxa and reveal island formation as a key driver of speciation." Molecular biology and evolution 36.4 (2019): 784–797.
  81. Web site: Waitaha penguin New Zealand Birds Online . 2022-05-19 . www.nzbirdsonline.org.nz.
  82. Rawlence, Nicolas J., et al. "Radiocarbon-dating and ancient DNA reveal rapid replacement of extinct prehistoric penguins." Quaternary Science Reviews 112 (2015): 59–65.
  83. Web site: Kohatu shag Kawau kōhatu New Zealand Birds Online . 2023-03-30 . nzbirdsonline.org.nz . en.
  84. Rawlence . Nicolas J. . Till . Charlotte E. . Easton . Luke J. . Spencer . Hamish G. . Schuckard . Rob . Melville . David S. . Scofield . R. Paul . Tennyson . Alan J.D. . Rayner . Matt J. . Waters . Jonathan M. . Kennedy . Martyn . 2017 . Speciation, range contraction and extinction in the endemic New Zealand King Shag complex . Molecular Phylogenetics and Evolution . 115 . 197–209 . 10.1016/j.ympev.2017.07.011 . 28803756. 2017MolPE.115..197R .
  85. Ixobrychus novaezelandiae. e.T22697307A93607264. BirdLife International. 2016. 2016. 20 May 2022. 10.2305/IUCN.UK.2016-3.RLTS.T22697307A93607264.en.
  86. Web site: New Zealand little bittern New Zealand Birds Online . 2022-05-21 . www.nzbirdsonline.org.nz.
  87. Worthy, T. H. and R. N. Holdaway. (2002): The lost world of the Moa: Prehistoric Life of New Zealand. Indiana University Press, Bloomington.
  88. Web site: Eyles' harrier New Zealand Birds Online . 2022-05-21 . www.nzbirdsonline.org.nz.
  89. Web site: Haast's eagle New Zealand Birds Online . 2022-05-21 . nzbirdsonline.org.nz.
  90. Web site: Laughing owl New Zealand Birds Online . 2022-05-21 . www.nzbirdsonline.org.nz.
  91. BirdLife International . 2016 . Sceloglaux albifacies . 2016 . e.T22689496A93232295 . 10.2305/IUCN.UK.2016-3.RLTS.T22689496A93232295.en . 18 November 2021.
  92. An extinct nestorid parrot (Aves, Psittaciformes, Nestoridae) from the Chatham Islands, New Zealand. Zoological Journal of the Linnean Society . 2014. J.R. . Wood . Mitchell K. J. . Scofield R. P. . Tennyson A. J. D. . Fidler A. E. . Wilmshurst J. M. . Llamas B. . Cooper A. . 172. 185–199. 10.1111/zoj.12164 . free .
  93. Web site: Lyall's wren New Zealand Birds Online . 2022-05-22 . www.nzbirdsonline.org.nz.
  94. BirdLife International . 2016 . Traversia lyalli . 2016 . e.T22698593A93691279 . 10.2305/IUCN.UK.2016-3.RLTS.T22698593A93691279.en . 18 November 2021.
  95. Web site: Bush wren New Zealand Birds Online . 2022-05-22 . www.nzbirdsonline.org.nz.
  96. Web site: North Island stout-legged wren New Zealand Birds Online . 2022-05-22 . www.nzbirdsonline.org.nz.
  97. Web site: South Island stout-legged wren New Zealand Birds Online . 2022-05-22 . nzbirdsonline.org.nz.
  98. Web site: Long-billed wren New Zealand Birds Online . 2022-05-22 . nzbirdsonline.org.nz.
  99. BirdLife International . 2016 . Anthornis melanocephala . 2016 . e.T22728814A94997726 . 10.2305/IUCN.UK.2016-3.RLTS.T22728814A94997726.en . 18 November 2021.
  100. Web site: Chatham Island bellbird New Zealand Birds Online . 2022-05-22 . www.nzbirdsonline.org.nz.
  101. Web site: North Island piopio New Zealand Birds Online . 2022-05-23 . nzbirdsonline.org.nz.
  102. Web site: South Island piopio New Zealand Birds Online . 2022-05-23 . www.nzbirdsonline.org.nz.
  103. Web site: New Zealand raven New Zealand Birds Online . 2022-05-23 . www.nzbirdsonline.org.nz.
  104. Web site: Chatham Island raven New Zealand Birds Online . 2022-05-23 . www.nzbirdsonline.org.nz.
  105. Web site: Chatham Island raven New Zealand Birds Online . 2022-05-23 . www.nzbirdsonline.org.nz.
  106. Web site: Huia New Zealand Birds Online . 2022-05-23 . www.nzbirdsonline.org.nz.
  107. Callaeas cinereus. e.T103730380A156237250. BirdLife International. 2019. 2019. 22 May 2022. 10.2305/IUCN.UK.2019-3.RLTS.T103730380A156237250.en.
  108. Web site: South Island kokako New Zealand Birds Online . 2022-05-23 . nzbirdsonline.org.nz.
  109. Web site: Newman . Tim . 2018-11-09 . Sightings spark hope in the search for New Zealand's most wanted bird . 2022-05-23 . Stuff . en.
  110. Web site: 2022-01-13 . South Island kōkako: Recording raises hopes of spotting elusive bird . 2022-05-23 . RNZ . en-nz.
  111. Web site: Ridout . Amy . 2022-01-11 . Bird call raises hopes of finding the elusive kōkako . 2022-05-23 . Stuff . en.
  112. Web site: Chatham Island fernbird New Zealand Birds Online . 2022-05-23 . nzbirdsonline.org.nz.
  113. Web site: Oligosoma northlandi (Worthy, 1991) COL . 2022-05-23 . www.catalogueoflife.org.
  114. Worthy . T.H. . December 1991 . Fossil skink bones from Northland, New Zealand, and description of a new species of Cyclodina, Scincidae . Journal of the Royal Society of New Zealand . 21 . 4 . 329–348 . 10.1080/03036758.1991.10420831 . 1991JRSNZ..21..329W .
  115. Book: Hitchmough . Rod . Conservation Status of New Zealand Reptiles, 2021 . Barr . Ben . Knox . Carey . Lettink . Marieke . Monks . Joanne M. . Patterson . Geoff B. . Reardon . James T. . van Winkel . Dylan . Rolfe . Jeremy . 2021 . . New Zealand Threat Classification Series . 35 . 3–6 . species:Geoff B. Patterson . 5 November 2023 . Michel . Pascale.
  116. Worthy . Trevor H. . 1987-12-01 . Palaeoecological information concerning members of the frog genus Leiopelma: Leiopelmatidae in New Zealand . Journal of the Royal Society of New Zealand . 17 . 4 . 409–420 . 10.1080/03036758.1987.10426482 . 0303-6758. free . 1987JRSNZ..17..409W .
  117. West, D. . David, B. . Ling, N. . 2014 . Prototroctes oxyrhynchus . 2014 . e.T18384A20887241 . 10.2305/IUCN.UK.2014-3.RLTS.T18384A20887241.en . 23 May 2022.
  118. Mecodema punctellum. e.T12922A3398669. World Conservation Monitoring Centre. 1996. 1996. 23 May 2022. World Conservation Monitoring Centre. 10.2305/IUCN.UK.1996.RLTS.T12922A3398669.en.
  119. Tokea orthostichon. e.T103192401A103193535. Blakemore, R. J.. 2017. 2017. 23 May 2022. 10.2305/IUCN.UK.2017-2.RLTS.T103192401A103193535.en.
  120. Web site: Lepidium amissum . 2022-05-26 . New Zealand Plant Conservation Network . en.
  121. de Lange, P. . 2014. Lepidium amissum . e.T62798275A62798278 . 10.2305/IUCN.UK.2014-2.RLTS.T62798275A62798278.en. 1 June 2024.
  122. Lepidium obtusatum. e.T62798292A62798296. de Lange. P.. 2013. 2013. 25 May 2022. 10.2305/IUCN.UK.2014-2.RLTS.T62798292A62798296.en.
  123. Web site: Lepidium obtusatum . 2022-05-26 . New Zealand Plant Conservation Network . en.
  124. Adams Mistletoe. e.T62798347A62798350. de Lange. P.. 2013. 2013. 25 May 2022. World Conservation Monitoring Centre. 10.2305/IUCN.UK.2014-2.RLTS.T62798347A62798350.en.
  125. Stellaria elatinoides. e.T62798337A62798340. de Lange. P.. 2013. 2013. 25 May 2022. 10.2305/IUCN.UK.2014-2.RLTS.T62798337A62798340.en.
  126. Web site: Stellaria multiflora subsp. multiflora . 2022-05-26 . New Zealand Plant Conservation Network . en.
  127. de Lange, P. . 2014. Logania depressa . e.T62798304A62798308. 10.2305/IUCN.UK.2014-2.RLTS.T62798304A62798308.en. 1 June 2024.
  128. de Lange, P. . 2014. Myosotis laingii . e.T62798327A62798331 . 10.2305/IUCN.UK.2014-2.RLTS.T62798327A62798331.en . 1 June 2024.