List of Cyphophthalmi species explained
See main article: Cyphophthalmi. This page lists all described species of the harvestman suborder Cyphophthalmi. Unless otherwise noted, information is taken from Giribet's 2000 taxonomic catalogue[1] and from Giribet et al., 2012.[2]
Boreophthalmi
Sironidae
- Cyphophthalmus beschkovi (Mitov, 1994)
- Cyphophthalmus bithynicus (Gruber, 1969)
- Cyphophthalmus conocephalus Karaman, 2009
- Cyphophthalmus corfuanus (Kratochvíl, 1938)
- Cyphophthalmus duricorius Joseph, 1868
- Cyphophthalmus eratoae (Juberthie, 1968)
- Cyphophthalmus ere Karaman, 2008
- Cyphophthalmus gjorgjevici (Hadži, 1933)
- Cyphophthalmus gordani Karaman, 2009
- Cyphophthalmus hlavaci Karaman, 2009
- Cyphophthalmus klisurae (Hadži, 1973)
- Cyphophthalmus kratochvili Karaman, 2009
- Cyphophthalmus markoi Karaman, 2008
- Cyphophthalmus martensi Karaman, 2009
- Cyphophthalmus minutus (Kratochvíl, 1938)
- Cyphophthalmus montenegrinus (Hadzi, 1973)
- Cyphophthalmus neretvanus Karaman, 2009
- Cyphophthalmus noctiphilus (Kratochvíl, 1940)
- Cyphophthalmus nonveilleri Karaman, 2008
- Cyphophthalmus ognjenovici Karaman, 2009
- Cyphophthalmus ohridanus (Hadži, 1973)
- Cyphophthalmus paradoxus (Kratochvíl, 1958)
- Cyphophthalmus paragamiani Karaman, 2009
- Cyphophthalmus rumijae Karaman, 2009
- Cyphophthalmus serbicus (Hadži, 1973)
- Cyphophthalmus silhavyi (Kratochvíl, 1938)
- Cyphophthalmus solentiensis Dreszer, Raða & Giribet, 2015[4]
- Cyphophthalmus teyrovskyi (Kratochvíl, 1938)
- Cyphophthalmus thracicus Karaman, 2009
- Cyphophthalmus trebinjanus Karaman, 2009
- Cyphophthalmus yalovensis (Gruber, 1969)
- Cyphophthalmus zetae Karaman, 2009
- Holosiro Ewing, 1923
- Iberosiro distylos de Bivort & Giribet, 2004
- Iberosiro rosae Giribet, Merino-Sáinz & Benavides, 2017
- Neosiro Newell, 1943
- Neosiro ligiae (Giribet, 2017)
- Neosiro martensi Karaman, 2022
- Neosiro richarti (Benavides & Giribet, 2017)
- Odontosiro lusitanicus Juberthie, 1961
- Paramiopsalis Juberthie, 1962
- Paramiopsalis anadonae Giribet, Merino-Sáinz & Benavides, 2017
- Paramiopsalis eduardoi Murienne & Giribet, 2009
- Paramiopsalis ramblae Benavides & Giribet, 2017
- Paramiopsalis ramulosa Juberthie, 1962
- Siro Latreille, 1796
- Siro carpaticus Rafalski, 1956
- Siro crassus Novak & Giribet, 2006[6]
- Siro franzi Karaman & Raspotnig, 2022
- Siro ozimeci Karaman, 2022
- Siro rubens Latreille, 1804
- Siro richarti Benavides & Giribet, 2017
- Siro valleorum Chemini, 1990
- [Also †''[[Siro platypedibus]] Dunlop & Giribet, 2003 (fossil, Baltic amber)]
- Suzukielus Juberthie, 1970
- Suzukielus sauteri (Roewer, 1916)
Insetae sedis:
- Marwe coarctata Shear, 1985
Stylocellidae[7]
Fangensinae
- Fangensis cavernarus Schwendinger & Giribet, 2005
- Fangensis leclerci Rambla, 1994
- Fangensis spelaeus Schwendinger & Giribet, 2005
- Giribetia Clouse, 2012
- Giribetia insulanus (Schwendinger & Giribet, 2005)
Leptopsalidinae[8]
- Leptopsalis Thorell, 1882
- Leptopsalis beccarii Thorell, 1882
- Leptopsalis dumoga (Shear, 1993)
- Leptopsalis foveolata Clouse & Schwendinger, 2012[9]
- Leptopsalis hillyardi (Shear, 1993)
- Leptopsalis javana (Thorell, 1882)
- Leptopsalis laevichelis (Roewer, 1942)
- Leptopsalis lydekkeri (Clouse & Giribet, 2007)
- Leptopsalis modesta (Hansen & Sørensen, 1904)
- Leptopsalis novaguinea (Clouse & Giribet, 2007)
- Leptopsalis pangrango (Shear, 1993)
- Leptopsalis ramblae (Giribet, 2002)
- Leptopsalis sedgwicki (Shear, 1979)
- Leptopsalis sulcata (Hansen & Sørensen, 1904)
- Leptopsalis tambusisi (Shear, 1993)
- Leptopsalis thorellii (Hansen & Sørensen, 1904)
- Leptopsalis weberii (Hansen & Sørensen, 1904)
- Miopsalis Thorell, 1890
- Miopsalis collinsi (Shear, 1993)
- Miopsalis dillyi Schmidt, Clouse & Sharma, 2020[10]
- Miopsalis globosa (Schwendinger & Giribet, 2004)
- Miopsalis gryllospeca (Shear, 1993)
- Miopsalis kinabalu (Shear, 1993)
- Miopsalis leakeyi (Shear, 1993)
- Miopsalis lionota (Pocock, 1897)
- Miopsalis mulu (Shear, 1993)
- Miopsalis pocockii (Hansen & Sørensen, 1904)
- Miopsalis pulicaria Thorell, 1890
- Miopsalis sabah (Shear, 1993)
- Miopsalis silhavyi (Rambla, 1991)
- Miopsalis tarumpitao (Shear, 1993)
Stylocellinae
- Meghalaya annandalei Giribet, Sharma, & Bastawade, 2007
- Stylocellus Westwood, 1874
- Stylocellus lornei Clouse, 2012
- Stylocellus spinifrons Roewer, 1942 (nomen dubium) [unidentifiable juvenile]
- Stylocellus sumatranus Westwood, 1874
Scopulophthalmi
Pettalidae
- Aoraki calcarobtusa (Forster, 1952)
- Aoraki crypta (Forster, 1948)
- Aoraki denticulata (Forster, 1948)
- Aoraki denticulata denticulata (Forster, 1948)
- Aoraki denticulata major (Forster, 1948)
- Aoraki granulosa (Forster, 1952)
- Aoraki healyi (Forster, 1948)
- Aoraki inerma (Forster, 1948)
- Aoraki longitarsa (Forster, 1952)
- Aoraki stephenesis (Forster, 1952)
- Aoraki tumidata (Forster, 1948)
- Aoraki westlandica (Forster, 1952)
- Archaeopurcellia Giribet, Shaw, Lord & Derkarabetian, 2022
- Archaeopurcellia eureka Giribet, Shaw, Lord & Derkarabetian, 2022
- Austropurcellia Juberthie, 1988[12]
- Austropurcellia absens Boyer & Popkin-Hall, 2015
- Austropurcellia acuta Popkin-Hall & Boyer, 2014
- Austropurcellia alata Boyer & Reuter, 2012
- Austropurcellia arcticosa (Cantrell, 1980)
- Austropurcellia barbata Popkin-Hall & Boyer, 2014
- Austropurcellia cadens Baker & Boyer, 2015
- Austropurcellia capricornia (Davies, 1977)
- Austropurcellia clousei Boyer, Baker & Popkin-Hall, 2015
- Austropurcellia culminis Boyer & Reuter, 2012
- Austropurcellia daviesae (Juberthie, 1989)
- Austropurcellis despectata Boyer & Reuter, 2012
- Austropurcellia finniganensis Popkin-Hall, Jay & Boyer, 2016
- Austropurcellia forsteri (Juberthie, 2000)
- Austropurcellia fragosa Popkin-Hall, Jay & Boyer, 2016
- Austropurcellia giribeti Boyer & Quay, 2015
- Austropurcellia megatanka Jay, Coblens & Boyer, 2016
- Austropurcellia monteithi Jay, Popkin-Hall, Coblens & Boyer, 2016
- Austropurcellia nuda Popkin-Hall, Jay & Boyer, 2016
- Austropurcellia riedeli Jay, Oberski & Boyer, 2016
- Austropurcellia scoparia Juberthie, 1988
- Austropurcellia sharmai Boyer & Quay, 2015
- Austropurcellia superbensis Popkin-Hall & Boyer, 2014
- Austropurcellia tholei Baker & Boyer, 2015
- Austropurcellia vicina Boyer & Reuter, 2012
- Austropurcellia woodwardi (Forster, 1955)
- Chileogovea Roewer, 1961
- Chileogovea jocasta Shear, 1993
- Chileogovea oedipus Roewer, 1961
- Karripurcellia Giribet, 2003[13]
- Karripurcellia peckorum Giribet, 2003
- Karripurcellia sierwaldae Giribet, 2003
- Manangotria Shear & Gruber, 1996
- Manangotria taolanaro Shear & Gruber, 1996
- Neopurcellia Forster, 1948
- Neopurcellia salmoni Forster, 1948
- Parapurcellia Rosas Costa, 1950[14]
- Parapurcellia amatola de Bivort & Giribet, 2010
- Parapurcellia convexa de Bivort & Giribet, 2010
- Parapurcellia fissa (Lawrence, 1939)
- Parapurcellia minuta de Bivort & Giribet, 2010
- Parapurcellia monticola (Lawrence, 1939)
- Parapurcellia natalia de Bivort & Giribet, 2010
- Parapurcellia rumpiana (Lawrence, 1933)
- Parapurcellia peregrinator (Lawrence, 1963)
- Parapurcellia silvicola (Lawrence, 1939)
- Parapurcellia staregai de Bivort & Giribet, 2010
- Parapurcellia transvaalica (Lawrence, 1963)[15]
- Pettalus Thorell, 1876
- Pettalus brevicauda Pocock, 1897
- Pettalus cimiciformis (Cambridge, 1875)
- Pettalus lampetides Sharma & Giribet 2006[16]
- Pettalus thwaitesi Sharma, Karunarathna & Giribet, 2009[17]
- Purcellia Hansen & Sørensen, 1904
- Purcellia argasiformis (Lawrence, 1931)
- Purcellia griswaldi de Bivort & Giribet, 2010
- Purcellia illustrans Hansen and Sørensen, 1904
- Purcellia lawrencei de Bivort & Giribet, 2010
- Purcellia leleupi Staręga, 2008[18]
- Rakaia Hirst, 1926
- Rakaia antipodiana Hirst, 1926
- Rakaia collaria Roewer, 1942
- Rakaia dorothea Phillips and Grimmett, 1932
- Rakaia florensis (Forster, 1948)
- Rakaia isolata Forster, 1952
- Rakaia lindsayi Forster, 1952
- Rakaia macra Boyer and Giribet, 2003
- Rakaia magna Forster, 1948
- Rakaia magna australis Forster, 1952
- Rakaia magna magna Forster, 1948
- Rakaia media Forster, 1948
- Rakaia media media Forster, 1948
- Rakaia media insula Forster, 1952
- Rakaia minutissima (Forster, 1948)
- Rakaia pauli Forster, 1952
- Rakaia solitaria Forster, 1948
- Rakaia sorenseni Forster, 1952
- Rakaia sorenseni digitata Forster, 1952
- Rakaia sorenseni sorenseni Forster, 1952
- Rakaia stewartiensis Forster, 1948
- Rakaia uniloca Forster, 1952
Sternophthalmi
Neogoveidae[19]
- Brasiliogovea Martens, 1969
- Brasiliogovea aphantostylus Benavides, Hormiga, & Giribet, 2019
- Brasiliogovea chiribiqueta Benavides & Giribet, 2013
- Brasiliogovea microphaga Martens, 1969
- Brasiliogovea microstylus Benavides, Hormiga, & Giribet, 2019
- Brasiliogovea yacambuensis Benavides, Hormiga, & Giribet, 2019
- Canga DaSilva, Pinto-da-Rocha & Giribet, 2010
- Huitaca boyacanensis Benavides & Giribet, 2013
- Huitaca bitaco Benavides & Giribet, 2013
- Huitaca caldas Benavides & Giribet, 2013
- Huitaca depressa Benavides & Giribet, 2013
- Huitaca sharkeyi Benavides & Giribet, 2013
- Huitaca tama Benavides & Giribet, 2013
- Huitaca ventralis Shear, 1979
- Leggogovia Benavides & Giribet, 2019
- Leggogovia pabsgarnoni (Legg. 1990)
- Metagovea Rosas-Costa, 1950
- Metagovea disparunguis Rosas Costa, 1950
- Metagovea ligiae Giupponi & Kury, 2015
- Metagovea matapi Benavides, Hormiga, & Giribet, 2019
- Metagovea philipi Goodnight & Goodnight, 1980
- Metagovea planada Benavides, Hormiga, & Giribet, 2019
- Metasiro Juberthie, 1960[20]
- Metasiro americanus (Davis, 1933)
- Metasiro savannahensis Clouse & Wheeler, 2014
- Microgovia chenepau Benavides, Hormiga, & Giribet, 2019
- Microgovia oviformis Martens, 1969
- Neogovea Hinton, 1938
- Neogovea branstetteri Benavides, Hormiga, & Giribet, 2019
- Neogovea enigmatica Martens, 2019
- Neogovea hormigai Benavides & Giribet, 2013
- Neogovea immsi Hinton, 1938
- Neogovea kamakusa Shear, 1977
- Neogovea kartabo Shear, 1977
- Neogovea matawai Benavides, Hormiga, & Giribet, 2019
- Neogovea virginie Jocqué & Jocqué, 2011
- Parogovia Hansen, 1921
- Parogovia gabonica (Juberthie, 1969)
- Parogovia montealensis Benavides & Giribet, 2019
- Parogovia parasironoides Hiřman, Kotyk, Kotyková Varadínová,& Šťáhlavský, 2018
- Parogovia prietoi Benavides & Giribet, 2019
- Parogovia putnami Benavides & Giribet, 2019
- Parogovia sironoides Hansen, 1921
- Tucanogovea Karaman, 2014
- Tucanogovea schusteri Karaman, 2014
- Waiwaigovia Benavides, Hormiga, & Giribet, 2019
- Waiwaigovia shultzi Benavides, Hormiga, & Giribet, 2019
Ogoveidae[21]
- Ogovea cameroonensis Giribet & Prieto, 2003
- Ogovea grossa (Hansen & Sørensen, 1904)
- Ogovea nasuta (Hansen, 1921)
Troglosironidae[22]
- Troglosiro aelleni Juberthie, 1979
- Troglosiro brevifossa Sharma & Giribet, 2009
- Troglosiro dogny Giribet & Baker, 2021
- Troglosiro juberthiei Shear, 1993
- Troglosiro longifossa Sharma & Giribet, 2005
- Troglosiro monteithi Sharma & Giribet, 2009
- Troglosiro ninqua Shear, 1993
- Troglosiro oscitatio Sharma & Giribet, 2009
- Troglosiro pin Giribet, Baker & Sharma, 2021
- Troglosiro platnicki Shear, 1993
- Troglosiro pseudojuberthiei Giribet, Baker & Sharma, 2021
- Troglosiro raveni Shear, 1993
- Troglosiro sharmai Giribet & Baker, 2021
- Troglosiro sheari Sharma & Giribet, 2009
- Troglosiro tillierorum Shear, 1993
- Troglosiro urbanus Sharma & Giribet, 2009
- Troglosiro wilsoni Sharma & Giribet, 2009
- Ankaratra franzi Shear & Gruber, 1996
- Shearogovea Giribet, 2011
- Shearogovea mexasca (Shear, 1977)
Parasironidae Karaman, Mitov & Snegovaya, 2024
- Cimmerosiro juberthiei Karaman, Mitov & Snegovaya, 2024
- Cimmerosiro krivolutskyi Karaman, Mitov & Snegovaya, 2024
- Cimmerosiro rhodiensis Karaman, Mitov & Snegovaya, 2024
- Ebrosiro Karaman, Mitov & Snegovaya, 2024
- Tirrenosiro axeli Karaman, Mitov & Snegovaya, 2024
- Tirrenosiro minor (Juberthie, 1958)
Notes and References
- Giribet. Gonzalo. 2000. Catalogue of the Cyphophthalmi of the World (Arachnida, Opiliones). Revista Ibérica de Aracnología. 2. 49–76.
- Giribet. Gonzalo. Sharma. Prashant. Benavides. Ligia. Boyer. Sarah. Clouse. Ronald. de Bivort. Benjamin. Dimitrov. Dimitar. Kawauchi. Gisele. Murienne. Jerome. Schwendinger. Peter. 3. 2012. Evolutionary and biogeographical history of an ancient and global group of arachnids (Arachnida: Opiliones: Cyphophthalmi) with a new taxonomic arrangement. Biological Journal of the Linnean Society. 105. 1. 92–130. 10.1111/j.1095-8312.2011.01774.x. free.
- Karaman. Ivo M.. 2009-06-01. The taxonomical status and diversity of Balkan sironids (Opiliones, Cyphophthalmi) with descriptions of twelve new species. Zoological Journal of the Linnean Society . 156. 2. 260–318. 10.1111/j.1096-3642.2009.00446.x. 0024-4082. free.
- Giribet. Gonzalo. Raða. Tonći. Dreszer. Taras B.. 2015. Cyphophthalmus solentiensis sp. nov. (Cyphophthalmi, Sironidae), a New Endogean Mite Harvestman Species from Croatia, with an Application of Confocal Laser Microscopy to Illustrate Genitalia in Opiliones. Breviora. 543. 1. 1–15. 10.3099/MCZ18.1. 0006-9698.
- Merino-Sáinz. Izaskun. Benavides. Ligia R.. Giribet. Gonzalo. 2017-08-30. The systematics and biogeography of the mite harvestman family Sironidae (Arachnida : Opiliones : Cyphophthalmi) with the description of five new species. Invertebrate Systematics. en. 31. 4. 456–491. 10.1071/IS16086. 1447-2600.
- (2006): A new species of Cyphophthalmi (Arachnida, Opiliones, Sironidae) from Eastern Slovenia. Zootaxa 1330: 27-42. PDF
- Clouse. Ronald. 2012. The lineages of Stylocellidae (Arachnida: Opiliones: Cyphophthalmi). Zootaxa. 3595. 3595. 1–34. 10.11646/zootaxa.3595.1.1. 1175-5334.
- Kury. Adriano. 2018. The correct spelling of Leptopsalidinae (Arachnida: Opiliones: Cyphophthalmi).. Arachnida – Rivista Aracnologica Italiana. 18. 40–43.
- Clouse. Ronald. Schwendinger. Peter. 2012. Leptopsalis foveolata sp. n., a new species of Stylocellidae from Thailand that displays a novel morphological feature in the suborder Cyphophthalmi (Arachnida, Opiliones). Revue suisse de Zoologie. 119. 529–546. 10.5962/bhl.part.150204.
- Schmidt. Stephanie. Clouse. Ronald. Sharma. Prashant. 2020. A new Miopsalis from Mindanao supports a biogeographic umbilicus between Borneo and the southern Philippines (Arachnida: Opiliones: Cyphophthalmi: Stylocellidae). Zootaxa. 4779. 3. 379–390. 10.11646/zootaxa.4779.3.6.
- Boyer. Sarah. Giribet. Gonzalo. 2007. A new model Gondwanan taxon: systematics and biogeography of the harvestman family Pettalidae (Arachnida, Opiliones, Cyphophthalmi), with a taxonomic revision of genera from Australia and New Zealand. Cladistics. 23. 4. 337–361. 10.1111/j.1096-0031.2007.00149.x.
- Boyer. Sarah. Sharma. Prashant. Oberski. Jill. Coblens. Michelle. Popkin-Hall. Zachary. Jay. Katya. 2016-04-05. New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia's Wet Tropics biodiversity hotspot. ZooKeys. en. 586. 37–93. 10.3897/zookeys.586.6774. 27199608. 4857030 . 1313-2970. free.
- Giribet. Gonzalo. 2003. Karripurcellia, a new pettalid genus (Arachnida: Opiliones: Cyphophthalmi) from Western Australia, with a cladistic analysis of the family Pettalidae. Invertebrate Systematics. 17. 3. 387–406. 10.1071/IS02014.
- de Bivort. Benjamin. Giribet. Gonzalo. 2010. A systematic revision of the South African Pettalidae (Arachnida:Opiliones:Cyphophthalmi) based on a combined analysis of discrete and continuous morphological characters with the description of seven new species. https://web.archive.org/web/20190226103610/http://pdfs.semanticscholar.org/7d11/88f81d4e9920293581a68e6f1c14f85555c3.pdf. dead. 2019-02-26. Invertebrate Systematics. 24. 4. 371–406. 10.1071/IS10015. 32628735 .
- Giribet. Gonzalo. Boyer. Sarah. Baker. Caitlin. Fernandéz. Rosa. Sharma. Prashant. de Bivort. Benjamin. Daniels. Savel. Harvey. Mark. Griswold. Charles. 3. 2016. A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society. 178. 3. 523–545. 10.1111/zoj.12419.
- Sharma. Prashant. Giribet. Gonzalo. 2006. A new Pettalus species (Opiliones, Cyphophthalmi, Pettalidae) from Sri Lanka with a discussion on the evolution of eyes in Cyphophthalmi. The Journal of Arachnology. 34. 2. 331–341. 10.1636/H05-33.1. 2019-09-10. 2020-10-24. https://web.archive.org/web/20201024203226/http://www.americanarachnology.org/JoA_free/JoA_v34_n2/arac-34-2-331.pdf. dead.
- Sharma. Prashant. Karunarathna. Indika. Giribet. Gonzalo. 2009. On the endemic Sri Lankan genus Pettalus (Opiliones, Cyphophthalmi, Pettalidae) with a description of a new species and a discussion of its diversity. Journal of Arachnology. 37. 1. 60–7. 10.1636/H08-56.1. 2019-09-10. 2021-04-17. https://web.archive.org/web/20210417124405/http://www.americanarachnology.org/JoA_free/JoA_v37_n1/arac-37-01-60.pdf. dead.
- Staręga. Wojciech. 2008. The second species of Purcellia Hansen et Sørensen, 1904 (Arachnida: Opiliones: Pettalidae) from South Africa. Polish Journal of Entomology. 77. 1. 51–56.
- Benavides. Ligia. Hormiga. Gustavo. Giribet. Gonzalo. 2019. Phylogeny, evolution and systematic revision of the mite harvestman family Neogoveidae (Opiliones Cyphophthalmi). Invertebrate Systematics. 33. 1. 101–180. 10.1071/IS18018.
- Clouse. Ronald. Wheeler. Ward. 2014. Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates . Zootaxa. 3814. 2. 177–201. 10.11646/zootaxa.3814.2.2. 24943422. 1175-5334.
- Giribet. Gonzalo. Prieto. Carlos E.. 2003-12-31. A new Afrotropical Ogovea (Opiliones, Cyphophthalmi) from Cameroon, with a discussion on the taxonomic characters in the family Ogoveidae. Zootaxa. 329. 1–18. 10.5281/zenodo.156214.
- Book: Zoologia Neocaledonica. Sharma. Prashant. Giribet. Gonzalo. Mémoires du Muséum national d'Histoire naturelle. 2009. 9782856536186. 7. 205–245. The family Troglosironidae (Opiliones: Cyphophthalmi) of New Caledonia. https://www.researchgate.net/publication/232643583.