Kota Formation Explained

Kota Formation
Type:Geological formation
Age:Jurassic
~
Period:Jurassic
Prilithology:Mudstone, sandstone
Otherlithology:Limestone
Namedfor:Kota Village
Region:Telangana
Country: India
Coordinates:18.9°N 80°W
Paleocoordinates:-31.6°N 31.9°W
Subunits:Lower & Upper members
Overlies:Dharmaram Formation
Thickness:550-
Extent:Pranhita-Godavari Basin

The Kota Formation is a geological formation in India. The age of the Kota Formation is uncertain; it is commonly considered to date to the Early Jurassic, but some studies have suggested it may extend into the Middle Jurassic or even later. It conformably overlies the Lower Jurassic Upper Dharmaram Formation and is unconformably overlain by the Lower Cretaceous Gangapur Formation. It is split into a Lower Member and Upper Member. The Lower Member is approximately 100 m thick while the Upper Member is 490 m thick. Both subunits primarily consist of mudstone and sandstone, but near the base of the upper unit there is a 20-30 metre thick succession of limestone deposited in a freshwater setting.[1]

Stratigraphy

The lower boundary of the Kota Formation is made of pebbly sandstone, covering the topmost clay seen in the Dharmaram Formation.[2] The Kota Formation has been traditionally divided into 2 main members, the Lower and Upper members, yet more recent work have redivided it into 3.[3] The Lower member can be seen at locations such as Adamilli, Kamavarapukota and Sudikonda, being made of sandstones, with clay clasts, with greater or lower stratification.[4] The Middle Member is well developed along the Continental Gondwana basin, specially towards the northwestern part, and is made of medium to fine white sandstone with clay and concretionary limestone, suggesting the development of paleosols associated with alluvial floodplains.[3] The last member is mostly made of broad sandstone sheets with large clay casts associated with fluvial channels, and has an extension that can be easuly seen on several continuous kilometers.[4] [5] The Uppermost section of the unit is mostly made of limestones and is overlain on an angular unconformity by the Gangapur Formation.[3]

Age

The age of the Kota Formation is controversial. There are no magmatic rocks or volcanic ash beds associated with the Kota Formation, which means that its age cannot be determined directly through radiometric dating.[6] [7] The maximum age of the Kota Formation is constrained by the underlying Upper Dharmaram Formation, which is Early Jurassic, probably Hettangian or Sinemurian, in age.[8] [7] Various researchers have attempted to date the Kota Formation using biostratigraphy. Krishnan (1968), Jain (1973), and Yadagiri and Prasad (1977) favored an Early Jurassic age based on the fish fauna. Govindan (1975) suggested a Middle Jurassic age based on ostracods. In 2006, Bandyopadhyay and Sengupta argued that the fish fauna suggested a Toarcian age for the Upper Kota Formation, possibly extending into the Aalenian, and in turn estimated the Lower Kota to be Sinemurian to Pliensbachian in age.[9] Guntupalli V. R. Prasad, along with various coauthors, has argued for a younger age. In 2001, Vijaya and Prasad proposed based on palynological evidence that the Kota Formation was deposited between the Callovian age of the Middle Jurassic and the Barremian age of the Early Cretaceous.[10] In 2002, Prasad and Manhas argued that the mammal genus Dyskritodon, known only from the Kota Formation and the Early Cretaceous of Morocco, provides evidence for a young age for the Kota Formation. In 2020, Prasad and Parmar argued that the similarity of the dinosaur fauna of the Kota Formation to that of the Middle Jurassic of the United Kingdom supported a Middle Jurassic age for the Kota Formation.

Paleoenvironment

The Kota Formation represents mostly a Continental succession related to a continental rift basin, the Pranhita-Godavari Gondwana Basin of peninsular India. The associated facies of sandstone and limestones are likely related to playa-type lake, with nearby fluvial currents, part of low gradient hanging wall alluvial fans, being deposited on it´s margin. There have been records of freshwater lue green algal stromatolites and oncolites, suggested to be deposited on low energy and low bathymetry lacustrine settings.[11] More recent works have proven the basin hosted in the Early Jurassic a freshwater carbonate wetland marked by the presence of limestones.[1] The environmental model proposed include a depositional cycle marked by several facies types, A for the sublittoral zones of shallow water bodies, followed by palustrine environments, including surfaces with abundance of influence of both plants and animals, specially rhizobrecciation indicating active colonization of the margins by plants, having a similar deposition to the modern Las Tablas de Daimiel wetlands.[1] Associated with the lacustrine facies have recovered microbial bioherms and lacustrine spring mounds, shallow ephemeral ponds with carbonated mud and Phyllopods, pedogenic calcrete under arid seasons and short-lived distributary channels. The depositional setting may have been partially sheltered from the input of siliciclastic materials, except on flooding seasons. Microbial biomats likely developed on shallow waters, while rhizoliths increased it´s presence of abandoned channel fills and pedogenic facies indicate drought seasons.

Fossil content

Ostracoda

Ostracods
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
ClinocyprisClinocypris sp.
  • Daroghapalli
  • Upper Member
Six carapacesA freshwater ostracodan of the Family Pontocyprididae.
CypredeaCypredea sp.
  • Daroghapalli
  • Upper Member
Fourteen incomplete carapacesA freshwater ostracodan of the Family Palaeocytheridae.
Darwinula[12] [13] Darwinula cf.sarytirmenensis
  • Aklapalli
  • Daroghapalli
  • Kanchelli
  • Metpalli
  • Potepalli
  • Lower Member
  • Upper Member
More than 200 carapaces and valvesA freshwater ostracodan of the Family Darwinulidae. The most dominant genus locally and the main indicator of both fluvial and lacustrine settings
Darwinula kingi
  • Aklapalli
  • Daroghapalli
  • Kanchelli
  • Metpalli
  • Potepalli
  • Lower Member
  • Upper Member
Around 120 carapaces and valvesA freshwater ostracodan of the Family Darwinulidae.
Darwinula spp.
  • Daroghapalli
  • Lower Member
  • Upper Member
Nineteen CarapacesA freshwater ostracodan of the Family Darwinulidae.
EucandonaEucandona sp.
  • Daroghapalli
  • Upper Member
Eight incomplete carapacesA freshwater ostracodan of the Family Candoninae.
LimnocythereLimnocythere spp.
  • Daroghapalli
  • Lower Member
Three complete carapacesA freshwater ostracodan of the family Limnocytheridae.
Stenocypris?Stenocypris sp.
  • Daroghapalli
  • Upper Member
Single incomplete carapaceA freshwater ostracodan of the family Cyprididae.
TimiriaseviaTimiriasevia digitalis
  • Daroghapalli
  • Lower Member
Twenty complete carapaces and thirty-six partly broken carapaces.A freshwater ostracodan of the family Limnocytheridae.

Phyllopoda

Phyllopods
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Estheriina[14] Estheriina alibadadensis
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Estheriininae. The most abundant Estheriid in the region and the key element of the Estheriina biozone
Estheriina indijurassica
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Estheriininae.
Estheriina bullata
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Estheriininae.
Estheriina pranhitaensis
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Estheriininae.
Lioestheria[15] Lioestheria kotaensis
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Lioestheriidae. The second key element of the Estheriina biozone
Lioestheria crustabundis
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Lioestheriidae.
Lioestheria ssp.
  • Kota limestone ridge
  • Lower Member
ValvesA freshwater clam shrimp of the family Lioestheriidae.
PaleolimnadiaPaleolimnadia spp.
  • Kota limestone ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Estheriininae.
PseudeasmussiataPseudeasmussiata andhrapradeshia
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
ValvesA freshwater clam shrimp of the family Lioestheriidae.

Insecta

Insects
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Archimesoblatta[16] A. shivaTasch outcrop K-2 bed 8
  • Upper Member
Right forewing tegmenA mesoblattinid cockroach
Coleopteron[17] C. sp.Kota limestone ridge
  • Lower Member
Isolated wingsAn Indeterminate Coleopteran.
Kotaphialtites[18] K. frankmortoniSirpur Taluka, Tasch's K1 outcrop bed 2(A)
  • Upper Member
MCZ 11909, Isolated wingAn ephialtitid hymenopteran.
Protogryllus[19] P. lakshmi Kota Formation outcrop K-2
  • Upper Member
MCZ 3046, Isolated wing
Taschigatra[20] T. bharatajaSirpur Taluka, Tasch's K1 outcrop bed 3(A)
  • Upper Member
No.2013(3013), part and counterpart of well preserved wingA rhagionid dipteran.
T. tulyabhijanaSirpur Taluka, Tasch's K1 outcrop bed 2(A)
  • Upper Member
No. 5034, well preserved wingA rhagionid dipteran.
XyelulaX. alexandriKota Formation outcrop K-2
  • Upper Member
MCZ 11831, well preserved wingA sepulcid hymenopteran.

Fish

Fish
Taxon SpeciesLocation Stratigraphic position Material Notes Images
Indocoelacanthus[21] Indocoelacanthus robustus
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
  • ISI P. 39. Impression of the caudal fin
  • ISI P. 40. Nearly complete head and trunk squamation
  • ISI P. 41-45. Fragments of fin-rays
  • ISI P. 46. Neural arches, isolated scales and girdle elements
A robust freshwater coelacanth of the family Latimeriidae. Represents the largest member of the local freshwater fauna, measuring up to 70 cm.
Lepidotes[22] [23] Lepidotes deccanensis
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Complete Specimen
  • Isolated Remains
A freshwater neopterygian of the family Lepisosteiformes.
Lepidotes spp.
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
Isolated remainsA freshwater neopterygian of the family Lepisosteiformes.
Lonchidion[24] [25] Lonchidion indicus
  • Paikasigudem village
  • Yamanpalli bonebed
  • Upper Member
  • GSI.TI.2, tooth
  • Isolated Teeth
A freshwater elasmobranch of the family Lonchidiidae.
Paradapedium[26] Paradapedium egertoni
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Near Ankisha
  • Lower Member
  • Upper Member
  • BMNHP12146, Partial specimen
  • BMNHP12147, Almost complete specimen
  • BMNHP12148, Partial specimen
  • ISIP.32 Nearly complete specimen
  • ISIP.33 Nearly complete specimen
  • ISIP.34 Nearly complete specimen
  • ISIP.35 specimen lacking tail and head
A freshwater neopterygian of the family Dapediidae.
Pholidophorus[27] Pholidophorus kingi
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Several complete and incomplete specimens
A freshwater neopterygian of the family Pholidophoridae.
Pholidophorus indicus
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Several complete and incomplete specimens
A freshwater neopterygian of the family Pholidophoridae.
Polyacrodus[28] Polyacrodus? sp.
  • Paikasigudem village
  • Upper Member
  • Tooth
A freshwater elasmobranch of the family Polyacrodontidae.
PycnodontidaeIndeterminate
  • Paikasigudem village
  • Upper Member
  • About 60 specimens of Pharyngeal Teeth, including GSITL3-6
A freshwater neopterygian of the family Pycnodontidae, originally classified as Perciformes, yet suggested to be very similar to the Cretaceous pycnodont Stephanodus.
SemionotiformesIndeterminate
  • Paikasigudem village
  • Upper Member
  • Teeth
A freshwater neopterygian of the family Semionotiformes.
TetragonolepisTetragonolepis oldhami
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
  • GSI2145, an impression including most of the body
  • ISP36, fragment of trunk and skull
  • ISP37, nearly complete specimen
A freshwater neopterygian of the family Dapediidae.

Amphibia

Amphibians
Taxon SpeciesLocation Stratigraphic position Material Notes Images
Indeterminate
  • Paikasigudem village
  • Upper Member
  • GSI.TL.19, Mandible
  • GSI.TL.20, right illium
Indeterminate frog remains, originally referred to Pelobatidae due to be compared with younger Creteaceous Indian frog material
Indeterminate
  • Paikasigudem village
  • Upper Member
  • GSI.TI.10, mandible with one teeth
  • GSI.TI.16, fragment of mandible
  • GSI.TI.17, madible with 4 teeth
Indeterminate caudatan remains, originally referred to Sirenidae due to be compared with younger Creteaceous sirenid material

Mammaliaforms

Mammaliaforms
Taxon SpeciesLocationStratigraphic positionMaterialNotesImages
Australosphenida[29] IndeterminatePaikasigudem village
  • Upper Member
Isolated lower molar A mammal of the group Australosphenida, resembling the south american genus Asfaltomylos
Dyskritodon?[30] Dyskritodon? indicusPaikasigudem village
  • Upper Member
VPL/JU/KM/13, lower left molarA dubious mammal of the group Eutriconodonta. This Genus is known from the Early Cretaceous of Morocco, what has been used to suggest a minimum Berrasian age for the Upper Kota Formation
GondtheriumGondtherium dattaiPaikasigudem village
  • Upper Member
VPL/JU/KM 12 right lower molarA mammal of the family Docodontidae
Indotherium[31] Indotherium pranhitai5 km west of Yamanapalli
  • Lower Member
GSI20795, right upper molarA mammaliform of the family Morganucodontidae. Includes the informally named "Indozostrodon simpsoni".[32]
Indobaatar[33] Indobaatar zofiae Paikasigudem village
  • Upper Member
VPL/JU/KM/20, a left upper premolar A mammal described as an eobaatarid multituberculate, but this interpretation has been challenged.[34]
Kotatherium[35] Kotatherium haldanei5 km west of Yamanapalli
  • Lower Member
GSI19634, right upper molar A mammaliform of the family Kuehneotheriidae
Nakunodon[36] Nakunodon paikasiensisPaikasigudem village
  • Upper Member
GSI.SR/PAL/12, right upper molar A mammal of the family Amphidontidae
Paikasigudodon[37] Paikasigudodon yadagiriiPaikasigudem village
  • Upper Member
VPL/JU/KM/10, right upper molarA mammaliaform of the family Morganucodontidae, originally known as "Kotatherium yadagirii"
TrishulotheriumTrishulotherium kotaensis Paikasigudem village
  • Upper Member
GSISR/PAL/10, left lower molarA mammal of the order Symmetrodonta

Lepidosauromorpha

Lepidosauromorphs
Taxon SpeciesLocation Stratigraphic position Material Notes Images
Bharatagama[38] Bharatagama rebbanensisPaikasigudem village
  • Upper Member
  • VPL/JU/KR 66, dentary
  • VPL/JU/KR 88, anterior region of a left maxilla
  • VPL/JU/KR 91, left maxilla
  • VPL/JU/KR 90-92, posterior ends of right maxillae
  • VPL/JU/KR 67-80-103, anterior symphysial region
  • JU/KR 79, 98, 100, dentaries mid-region
  • VPL/JU/KR 69, 81-84, 87, dentary with hatchling dentition
  • VPL/JU/KR 68, 85-86, dentary
  • VPL/JU/KR 70, 71, 76,78, 93, 94, 97, 104, 105, dentary
A lepidosauromorph originally described as an Iguanian lizard. May actually be a sphenodontian rather than a lizard.[39]
Godavarisaurus[40] Godavarisaurus lateefi
  • Gorlapalli village
  • Paikasigudem village
  • Upper Member
  • VP4Ju/KR47, a partial right dentary
  • VP4Ju/KR44, 51, 52, right maxilla
  • VP4Ju/KR40, 41, 43, 49, 50, left maxilla
  • VP4Ju/KR37, 48, 65, right dentary
  • VP4Ju/KR39, fragment of left dentary
  • VP4Ju/KR46, left palatine
A small sphenodontian, with a skull estimated to measure less than 20 mm
PaikasisaurusPaikasisaurus indicusPaikasigudem village
  • Upper Member
  • GSI.TI.14, left dentary with teeth
  • GSI.TI.15, left dentary with one tooth
An indeterminate and dubious lepidosauromorph, originally suggested to be a varanoid lizard
RebbanasaurusRebbanasaurus jainiPaikasigudem village
  • Upper Member
  • About 50 specimens, including premaxillae, maxillae, dentaries, and a palatine
A small sphenodont
Squamata?IndeterminatePaikasigudem village
  • Upper Member
  • VPL/JU/KR 62, fragmentary right maxilla
  • VPL/JU/KR 61, partial maxilla
Distinct from Bharatagama rebbanensis; may include material formerly assigned to the dubious Kota squamate Paikasisaurus indicus.
SphenodontidaeIndeterminatePaikasigudem village
  • Upper Member
  • GSI.TI.8-12, 17, maxilla
Indeterminate Sphenodontidae remains

Testudinata

Turtles
Taxon SpeciesLocation Stratigraphic position Material Notes Images
Indochelys[41] [42] Indochelys spatulata
  • 3 km NNE of Kistapur village, Kota
  • Near Kota village, north of Sironcha
  • Upper Member
  • GSI 20380, a partial shell
  • ISI R176, a partial shell
  • ISI R177, the anterior half of a shell
Mesochelydian stem-turtle, suggested to be related with Condorchelys
Testudines[43] Indeterminate 1 km south of Bodepalli
  • Lower Member
Carapace fragmentsIndeterminate turtle remains

Crocodylomorpha

Atoposaurid crocodiles are known from the unit, yet is not clear from what locality.[7]

Crocodylomorpha
TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
Crocodylomorpha[44] Indeterminate1 km south of BodepalliLower MemberMaxillae, dentaries, teethIndeterminate crocodylomorph remains, previously mixed with thyreophoran material and part of the chimaeric "Andhrasaurus"
Teleosauridae?[45] IndeterminateKota limestone ridge
  • Lower Member
Dermal scutes, with a femur and some fragments of other bonesIndeterminate material referred to crocodylomorphs similar to Teleosaurus

Pterosauria

Pterosaurs
TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
Campylognathoides[46] Campylognathoides indicusKota limestone ridge
  • Lower Member
  • ISI R38, holotype, fragment of skull and upper jaw
The holotype of Campylognathoides indicus, a pair of premaxillae, may represent a fish rather than a pterosaur.[47]
Pterosauria[48] Indeterminate
  • Boraigudem
  • Chitur
  • Lower Member
  • GSI17868, radius and ulna, wing metacarpal, first wing-phalanx and a clawed toe
  • ISIR49. fragment displaying dissociated bones from post-cranial region
Indeterminate pterosaur remains

Dinosaurs

Dinosaurs
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
"Andhrasaurus"[49] "Andhrasaurus indicus"1 km south of BodepalliLower MemberSacral vertebra, vertebral centra, dorsal vertebrae, caudal vertebrae, parts of scapula and ilium, osteodermsA chimaera of thyreophoran postcranial material and Crocodylomorph skull pieces. The armor was later suggested to be Ankylosauria indet. And other latter works pointed out it likely belongs to an indeterminate basal thyreophoran.[50]
Barapasaurus[51] Barapasaurus tagorei
  • North of Krishnapur
  • Pochampalli village
  • Lower Member
  • Roughly 300 bones, representing the disarticulated remains of at least six individuals
A sauropod dinosaur, either a Eusauropoda or more likely a Gravisauria. Represents the best-known Early Jurassic sauropod
Dandakosaurus[52] Dandakosaurus indicusYamanpalli bonebedLower Member
  • Lateral tooth, proximal ischium
  • Dorsal vertebra, proximal caudal vertebra (Likely of Kotasaurus)
A chimaera of large theropod bones, including and ischium and tooth, probably belonging to a carnosaur, and sauropod bones (two Kotasaurus vertebrae)
Dromaeosauridae?IndeterminatePaikasigudem village
  • Upper Member
Isolated TeethFive distinct morphotypes have been identified, mostly resemble coelurosaurs or dromaeosauroids.
Hypsilophodontidae[53] IndeterminateGorlapalli Village
  • Upper Member
  • Single Tooth
Identified originally as a member of Hypsilophodontidae, probably represents a tooth of a basal neornithischian.
Kotasaurus[54] Kotasaurus yamanpalliensisYamanpalli bonebed
  • Lower Member
Disarticulated remains of at least 12 individualsA basal sauropod
Ornithischia[55] Indeterminate
  • 1 km south of Bodepalli
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • End of left humerus
  • Isolated Teeth
Indeterminate ornithischian material. Among the teeth, at least five distinct morphotypes have been identified.
Richardoestesia?Richardoestesia? spp.Paikasigudem village
  • Upper Member
Isolated TeethTeeth similar to those of the problematic taxon Richardoestesia, of supposed coelurosaur affinities
Thyreophora[56] IndeterminatePaikasigudem village
  • Upper Member
Scute and associated fragmentary limb bones.Indeterminate scelidosaurid material.
SauropodaIndeterminate.Yamanpalli bonebed.• Lower MemberTeeth.Found in the same bonebed as Kotasaurus, resembles Amygdalodon.
CarnosauriaIndeterminate.Yamanapalli bonebed.• Lower MemberTeeth.Found in the same bonebed as Kotasaurus, referred to as'carnosaurs'

Megaflora

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Agathoxylon[57] Agathoxylon kotaense Near Kota village
  • Lower Member
Fossil woodAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Agathoxylon chandrapurensisNear Kota village
  • Lower Member
Fossil woodAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Agathoxylon santacruzense[58] Near Kota village
  • Lower Member
Fossil woodAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Agathoxylon pranhitaensis[59] Near Kota village
  • Lower Member
Fossil woodAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Agathoxylon santalenseNear Kota village
  • Lower Member
Fossil woodAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Agathoxylon spp.
  • Bodepalli
  • Near Kota village
  • Paikasigudem village
  • Lower Member
  • Upper Member
Fossil woodAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
AraucaritesAraucarites minutus
  • Dronadula
  • Lower Member
Branched shootsAffinities with Araucariaceae inside Pinales.
Araucarites sp.
  • Near Kota village
  • Lower Member
Branched shootsAffinities with Araucariaceae inside Pinales.
BrachyphyllumBrachyphyllum expansum
  • Near Kota village
  • Lower Member
Branched shootsAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
ElatocladusElatocladus conferta Near Kota village
  • Lower Member
Branched shootsAffinities with Cupressaceae inside Pinales.
Elatocladus tenerrimusNear Kota village
  • Lower Member
Branched shootsAffinities with Cupressaceae inside Pinales.
Elatocladus jabalpurensis Near Kota village
  • Lower Member
Branched shootsAffinities with Cupressaceae inside Pinales.
Elatocladus plana Near Kota village
  • Lower Member
Branched shootsAffinities with Cupressaceae inside Pinales.
Elatocladus sp.Near Kota village
  • Lower Member
Branched shootsAffinities with Cupressaceae inside Pinales.
CladophlebisCladophlebis denticulata
  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member
Isolated frondsAffinities with Osmundaceae in the Osmundales.
Cladophlebis indica
  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member
Isolated frondsAffinities with Osmundaceae in the Osmundales
Cladophlebis reversa
  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member
Isolated frondsAffinities with Osmundaceae in the Osmundales
Cladophlebis spp.
  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member
Isolated frondsAffinities with Osmundaceae in the Osmundales
Circoporoxylon[60] Circoporoxylon kotaense Near Kota village
  • Lower Member
Fossil woodAffinities with Podocarpaceae inside Pinales.
ConiopterisConiopteris hymenophylloides
  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member
Isolated frondsAffinities with Polypodiales in the Polypodiidae. Common cosmopolitan Mesozoic fern genus. Recent research has reinterpreted it a stem group of the Polypodiales (closely related to the extant genera Dennstaedtia, Lindsaea, and Odontosoria)
Coniopteris sp.
  • Near Kota village
  • Lower Member
Isolated frondsAffinities with Polypodiales in the Polypodiidae.
CupressinoxylonCupressinoxylon kotaense Near Kota village
  • Lower Member
Fossil woodAffinities with Cupressaceae inside Pinales.
DictyozamitesDictyozamites falcatus
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
Dictyozamites kotaense
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
EquisetumEquisetum rajmahalensis
  • Chitur village
  • Near Kota village
  • Lower Member
  • Upper Member
Isolated StemsAffinities with Equisetaceae inside Equisetales.
GinkgoitesGinkgoites lobata
  • Near Kota village
  • Lower Member
LeafsAffinities with Ginkgoaceae inside Ginkgoopsida.
Ginkgoxylon[61] Ginkgoxylon dixii Near Kota village
  • Lower Member
Fossil woodAffinities with Ginkgoaceae inside Ginkgoopsida.
HausmanniaHausmannia cf. buchii Near Kota village
  • Lower Member
Isolated pinnaeAffinities with Dipteridaceae in the Polypodiales.
OtozamitesOtozamites vemavarmensis
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
PagiophyllumPagiophyllum peregrinum Near Kota village
  • Lower Member
Branched shootsAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Pagiophyllum cf.peregrinum Near Kota village
  • Lower Member
Branched shootsAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Pagiophyllum spp. Near Kota village
  • Lower Member
Branched shootsAffinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
PachypterisPachypteris indica Near Kota village
  • Lower Member
Isolated pinnaeAffinities with Umkomasiaceae in the Pteridospermatophyta.
PlanoxylonPlanoxylon mahabalei Near Kota village
  • Lower Member
Fossil woodAffinities with Protopinaceae inside Pinales.
PodocarpoxylonPodocarpoxylon chandrapurensis Near Kota village
  • Lower Member
Fossil woodAffinities with Podocarpaceae inside Pinales.
Podocarpoxylon chiturensisChitur village
  • Upper Member
Fossil woodAffinities with Podocarpaceae inside Pinales.
Podocarpoxylon krauseliiNear Kota village
  • Lower Member
Fossil woodAffinities with Podocarpaceae inside Pinales.
Podocarpoxylon rajmahalenseNear Kota village
  • Lower Member
Fossil woodAffinities with Podocarpaceae inside Pinales.
Podocarpoxyion sewardiiNear Kota village
  • Lower Member
Fossil woodAffinities with Podocarpaceae inside Pinales.
Podocarpoxylon sp.Near Kota village
  • Lower Member
Fossil woodAffinities with Podocarpaceae inside Pinales.
PodozamitesPodozamites sp.
  • Near Kota village
  • Lower Member
LeafletsBroad conifer leaves
PseudoctenisPseudoctenis cf. frngilis
  • Near Kota village
  • Lower Member
LeafletsAffinities with Cycadales in the Cycadopsida.
PtilophyllumPtilophyllum fissum
  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
Ptilophyllum acutifolium
  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
Ptilophyllum cutchense
  • Chitur village
  • Near Kota village
  • Lower Member
  • Upper Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
Ptilophyllum cf.sahnii
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
Ptilophyllum cf.institacallum
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
Ptilophyllum sp.
  • Near Kota village
  • Lower Member
LeafletsAffinities with Williamsoniaceae in the Bennettitales.
Protaxodioxylon[62] Protaxodioxylon sahnii Chitur village
  • Upper Member
Fossil woodAffinities with Cupressaceae inside Pinales.
Prototaxoxylon liassicum[63] Near Kota village
  • Lower Member
Fossil woodAffinities with Cupressaceae inside Pinales.
SphenopterisSphenopteris kotaensis Near Kota village
  • Lower Member
Isolated FrondsAffinities with Dicksoniaceae in the Cyatheales.
TaxaceoxylonTaxaceoxylon sahnii Near Kota village
  • Lower Member
Fossil woodAffinities with Cupressaceae inside Pinales.
Taxaceoxylon biradarii[64] Chitur village
  • Lower Member
Fossil woodAffinities with Cupressaceae inside Pinales.
Taxaceoxylon antiquum Near Kota village
  • Lower Member
Fossil woodAffinities with Cupressaceae inside Pinales.
Taxaceoxylon sp.Near Kota village
  • Upper Member
Fossil woodAffinities with Cupressaceae inside Pinales.
TorreyitesTorreyites constricte
  • Chitur village
  • Upper Member
Branched shootsAffinities with Taxaceae in the Pinales.

See also

Notes and References

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  6. 10.7306/vj.17.1. 17. 1–16. Chinnappa. Chopparapu. Rajanikanth. Annamraju. Pauline Sabina. Kavali. Palaeofloras from the Kota Formation, India: palaeodiversity and ecological implications. Volumina Jurassica. 2019. 31 January 2024.
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