Kayenta Formation Explained

Kayenta Formation
Type:Geological formation
Age:Sinemurian-Toarcian
~
Period:Pliensbachian
Prilithology:Sandstone
Otherlithology:Siltstone, Limestone
Namedfor:Kayenta, Arizona
Country:United States
Coordinates:37.8°N -110.6°W
Unitof:Glen Canyon Group
Underlies:Navajo Sandstone
Overlies:Wingate Sandstone
Thickness:NaNm (-2,147,483,648feet)
Extent:northern Arizona, northwest Colorado, Nevada, and Utah[1]

The Kayenta Formation is a geological formation in the Glen Canyon Group that is spread across the Colorado Plateau province of the United States, including northern Arizona, northwest Colorado, Nevada, and Utah. Traditionally has been suggested as Sinemurian-Pliensbachian, but more recent dating of detrital zircons has yielded a depositional age of 183.7 ± 2.7 Ma, thus a Pliensbachian-Toarcian age is more likely.[2] A previous depth work recovered a solid "Carixian" (Lower-Middle Pliensbachian) age from measurements done in the Tenney Canyon.[3] More recent works have provided varied datations for the layers, with samples from Colorado and Arizona suggesting 197.0±1.5-195.2±5.5 Ma (Middle Sinemurian), while the topmost section is likely Toarcian or close in age, maybe even recovering terrestrial deposits coeval with the Toarcian Oceanic Anoxic Event.[4] This last age asignation also correlated the Toarcian Vulcanism on the west Cordilleran Magmatic Arc, as the number of grains from this event correlate with the silt content in the sandstones of the upper layers.[4]

This rock formation is particularly prominent in southeastern Utah, where it is seen in the main attractions of a number of national parks and monuments. These include Zion National Park, Capitol Reef National Park, the San Rafael Swell, and Canyonlands National Park.

The Kayenta Formation frequently appears as a thinner dark broken layer below Navajo Sandstone and above Wingate Sandstone (all three formations are in the same group). Together, these three formations can result in immense vertical cliffs of 2000feet or more. Kayenta layers are typically red to brown in color, forming broken ledges.

Kayenta Formation in Utah

Southeast Utah

In most sections that include all three geologic formations of the Glen Canyon group the Kayenta is easily recognized. Even at a distance it appears as a dark-red, maroon, or lavender band of thin-bedded material between two thick, massive, cross bedded strata of buff, tan, or light-red color. Its position is also generally marked by a topographic break. Its weak beds form a bench or platform developed by stripping the Navajo sandstone back from the face of the Wingate cliffs. The Kayenta is made up of beds of sandstone, shale, and limestone, all lenticular, uneven at their tops, and discontinuous within short distances. They suggest deposits made by shifting streams of fluctuating volume. The sandstone beds, from less than 1inches to more than 10feet thick, are composed of relatively coarse, well-rounded quartz grains cemented by lime and iron. The thicker beds are indefinitely cross bedded. The shales are essentially fine-grained, very thin sandstones that include lime concretions and balls of consolidated mud. The limestone appears as solid gray-blue beds, a few inches to a few feet thick, and as lenses of limestone conglomerate. Most of the limestone lenses are less than 25feet long, but two were traced for nearly 500feet and one for 1650feet.

Viewed as a whole, the Kayenta is readily distinguished from the geologic formations above and below it. It is unlike them in composition, color, manner of bedding, and sedimentary history. Obviously the conditions of sedimentation changed in passing from the Wingate Sandstone formation to the Kayenta and from the Kayenta to the Navajo sandstone, but the nature and regional significance of the changes have not been determined. In some measured sections the transition from Wingate to Kayenta is gradual; the material in the basal Kayenta, beds seems to have been derived from the Wingate immediately below and redeposited with only the discordance characteristic of fluviatile sediments. But in many sections the contact between the two formations is unconformable; the basal Kayenta consists of conglomerate and lenticular sandstone that fills depressions eroded in the underlying beds. In Moqui Canyon near Red Cone Spring nearly 10feet of Kayenta limestone conglomerate rests in a long meandering valley cut in Wingate. Likewise, the contact between the Kayenta and the Navajo in places seems to be gradational, but generally a thin jumbled mass of sandstone and shales, chunks of shale and limestone, mud balls, and concretions of lime and iron, lies at the base of the fine-grained, cross bedded Navajo. Mud cracks, a few ripple marks, and incipient drainage channels were observed in the topmost bed of the Kayenta on Red Rock Plateau; and in west Glen Canyon, wide sand-filled cracks appear at the horizon. These features indicate that, in places at least, the Wingate and Kayenta were exposed to erosion before their overlying geologic formations were deposited, are it may be that the range in thickness of the Kayenta thus in part (is) accounted for.

Southwest Utah

The red and mauve Kayenta siltstones and sandstones that form the slopes at base of the Navajo Sandstone cliffs record the record of low to moderate energy streams. Poole (1997) has shown that the streams still flowed toward the east depositing from 150 to 210 m (500 to about 700 feet) of sediment here. The sedimentary structures showing the channel and flood plain deposits of streams are well exposed on switchbacks below the tunnel in Pine Creek Canyon.

In the southeastern part of Zion National Park a stratum of cross bedded sandstone is found roughly halfway between the top and bottom of the Kayenta Formation. It is a "tongue" of sandstone that merges with the Navajo formation east of Kanab, and it shows that desert conditions occurred briefly in this area during Kayenta time. This tongue is the ledge that shades the lower portion of the Emerald Pool Trail, and it is properly called Navajo, not Kayenta.

Fossil mudcracks attest to occasional seasonal climate, and thin limestones and fossilized trails of aquatic snails or worms mark the existence of ponds and lakes. The most interesting fossils, however, are the dinosaur tracks that are relatively common in Kayenta mudstone.

These vary in size, but all seem to be the tracks of three-toed reptiles that walked upright, leaving their tracks in the muds on the flood plains. Unfortunately, so far no bone materials have been found in Washington County that would enable more specific identification.

Apparently during Kayenta time Zion was situated in a climatic belt like that of Senegal with rainy summers and dry winters at the southern edge of a great desert. The influence of the desert was about to predominate, however, as North America drifted northward into the arid desert belt.

Glen Canyon

The Kayenta Formation is approximately 400feet thick and consists of a fine-grained sandstone interbedded with layers of siltstone. The alternation of these units generally produces a series of ledges and slopes between the cliffs of the Navajo and Moenave formation. Dinosaur tracks are fairly common in the siltstone, and fresh water mussels and snails occur but are rare. The Kayenta Formation is colored pale red and adds to the splendor of the Vermilion Cliffs. It accumulated as deposits of rivers.

Fossils

Invertebrates

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Darwinula
  • D. magna
  • D.sarytirmensis
  • Gold Spring Quarry 1
Base of the "typical facies"ValvesA freshwater (lacustrine or fluvial) ostracod, type member of the family Darwinulidae. Kayenta specimens have a distinctive columnar calcitic layer. The Kayenta fauna is similar to the Sinemurian faunas of the Toutunhe Formation of Xinjiang, China.
Liratina
  • L. sp.
  • Gold Spring Quarry 1
Base of the "typical facies"Single shellA freshwater (lacustrine or fluvial) snail, incertae sedis inside Mesogastropoda. Differs considerably from the species from the Upper Jurassic Morrison Formation, Liratina jurassicum
Lymnaea[5]
  • L. hopii
  • Colorado Plateau
Base of the "typical facies"ShellsA freshwater (lacustrine or fluvial) snail, member of the family Lymnaeidae.
Scabriculocypris[6] S. n. sp
  • Gold Spring Quarry 1
Base of the "typical facies"ValvesA freshwater (lacustrine or fluvial) ostracod, incertae sedis inside Cypridacea. Differs from all other described species of the genus in being more elongate and from most in being spinose.
Unio[7]
  • U. dumblei
  • U. dockumensis
  • U. iridoides
  • South of Moab
Base of the "typical facies"ShellsA freshwater (lacustrine or fluvial) Bivalve, member of the family Unionidae.
Valvata
  • V. gregorii
  • Colorado Plateau
Base of the "typical facies"ShellsA freshwater (lacustrine or fluvial) snail, member of the family Valvatidae.

Fishes

The "Kayenta Fish Fauna" is the last one recovered from the Glen Canyon Group sequence and it is delimited mostly to the silty facies of the Lower-Middle Part of the formation.[8] This Fauna is rather scarce and delimited to several concrete locations with proper lacustrine or fluvial deposition, and are also scarce due to preservation bias.[8] Another aspect that can explain the lack of fish fossils found is the use of different research techniques than used on the Chinle Formation.[8]

Chondrichthyes

TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
HybodontoideaIndeterminate
  • Shonto Trading Post, Segi Canyon
  • Ward Terrace
Base of the "typical facies"UCMP 136104, 136105 + ten uncatalogued specimens, teethA freshwater (lacustrine or fluvial) non-neoselachian shark, incertae sedis inside Hybodontoidea. The remains of sharks are rather rare on the formation and limited to several locations with typical lacustrine or fluvial floodplain deposition.
Toarcibatidae (= "Archaeobatidae")[9] IndeterminateGold Spring Quarry 1Base of the "typical facies"Isolated ToothA freshwater (lacustrine or fluvial) toarcibatid. Related originally with Micropristis or Libanopristis, and stated to be reworked from younger Cretaceous deposits, was found due to its asymmetrical cusp to fit within the definition of Toarcibatis, being more likely to be native of the formation.

Actinopterygii

TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
LophionotusL. kanabensis
  • Washington Dome
  • Ward Terrace
  • Warner Valley
  • Zion National Park
Base of the "typical facies"
  • Fragmentary Specimens
  • Isolated Scales
A freshwater (lacustrine or fluvial) semionotid semionotiform.
PalaeoniscidaeIndeterminate
  • Utah Route 7 near St.George
  • Warner Valley
Base of the "typical facies"
  • Utah Route 7 near St.George
  • Isolated Ganoid Scales
  • Teeth
A freshwater (lacustrine or fluvial) palaeoniscid palaeonisciform.
SemionotidaeIndeterminate
  • Utah Route 7 near St.George
  • Downtown Moab
  • Warner Valley
  • Washington Dome
  • Ward Terrace
  • Warner Valley
  • Zion National Park
  • Desert Tortoise tracksite 1
Base of the "typical facies"
  • Large (1 m) Complete specimen
  • Isolated Ganoid Scales
  • Teeth
A freshwater (lacustrine or fluvial) semionotid semionotiform, probably related to the genus Semionotus. Semionotiformes are the only properly identified bony fishes from the formation, including a large specimen exposed at the Dan O’Laurie Museum.

Sarcopterygii

Genus / TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
Ceratodus[10] C. stewarti
  • Goblin Valley State Park
Middle “Silty Facies”
  • OMNH 69332, left pterygopalatine plate
A freshwater (lacustrine or fluvial) ceratodontid dipnomorph (lungfish).
CoelacanthidaeIndeterminate
  • Tsegi Canyon
  • Warner Valley
Base of the "typical facies"Uncertain RemainsA freshwater (lacustrine or fluvial) coelacanthid Coelacanthiform. Coelacanths are quoted from this zone, but their remains have not been studied.
Potamoceratodus[11] P. guentheri
  • Tsegi Canyon?
  • Little Colorado River Valley on Ward Terrace
Middle “Silty Facies”
  • Single dipnoan tooth plate (MCZ 13865)
A freshwater (lacustrine or fluvial) ceratodontid dipnomorph (lungfish). Was described as C. felchi, know from the Upper Jurassic Morrison Formation. Other dipnoan specimens have been cited but never described.

Amphibia

Genus / TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
AnuraIndeterminateGold Spring Quarry 1Silty Facies Member
  • MCZ 9019, distal humerus
  • MCZ 9020–24, ilia
An early frog, incertae sedis relationships
Eocaecilia[12] E. micropodiaGold Spring Quarry 1Silty Facies Member
  • MNA V8066 (type)
  • MNA V8053, 8054, 8055, 8057, 8058, 8059, 8060, 8062, 8063, 8064, 8065, 8066, 8067, 8068, 8069, 8070, 8071, 8072, 8073, 9346; MCZ 9011A, 9015, 9095, 9152, 9156. 9158, 9163, 9166, 9167, 9169A, 9171A, 9173, 9231A, 9233, 9235, 9237, 9238A, 9241, 9242
A genus whose relationships are controversial, being considered one of the earliest gymnophionans as a close relative of caecilians
LissamphibiaIndeterminateGold Spring Quarry 1Silty Facies Member
  • MCZ 9025–9028, jaws; MCZ 9031, 9032, vertebrae
  • MCZ 9035, atlas vertebra; MCZ 9034 + 4 uncatalogued specimens, proximal femora
  • MCZ 9066, 9067, proximal humeri
  • MCZ 9068–9072, proximal limb bones, possibly humeri
Incertae sedis relationships
Prosalirus[13] P. bitisGold Spring Quarry 1Silty Facies MemberMNA V 8725 (type) MCZ 9324 A, MCZ 9323 AAn early frog, probably related to Notobatrachidae
UrodelaIndeterminateGold Spring Quarry 1Silty Facies MemberMCZ 9017, 9018, atlas vertebraeA possible stem-salamander, incertae sedis inside Urodela. The oldest record of an urodelan from North America

Reptilia

Rhynchocephalia

TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
Navajosphenodon[14] N. sani
  • Gold Springs Quarry
  • Silty Facies Main Quarry
  • Adeii Eechii Cliffs
Silty Facies
  • MNA.V.12442, a fully articulated skeleton, including the skull, mandibles, axial and appendicular skeleton
  • Referred MCZ VP 9098, MCZ VP 101562, MCZ VP 9099, MCZ VP 101564, MCZ VP 101575, MCZ VP 9094, MCZ VP 9102, MCZ VP 9103, MCZ VP 101569, MCZ VP 101563, MNA.V.8726, MNA.V.8727
An Advanced Sphenodont, member of Sphenodontinae. The skeleton of N. sani shows a large number of similarities with the modern tuatara S. punctatus, clustering them closely together in the morphospace of sphenodontians and early lepidosaurs.
Rhynchocephalia[15] IndeterminateAirhead WestSilty FaciesUncertain FragmentsRhynchocephalians of uncertain assignment
Sphenodontia[16] IndeterminateGold Spring Quarry 1Silty FaciesMCZ 9036 through 9040, jaw fragmentsAn indeterminate sphenodont

Testudinatans

Genus / taxonSpeciesLocationStratigraphic positionMaterialNotesImages
CryptodiraIndeterminateRed Knob Silty FaciesUncertain fragmentsCryptodirans of uncertain assignment
Kayentachelys[17]
  • K. aprix
  • K. spp.
  • Gold Spring Quarry 1
  • Gold Spring South
  • Gold Spring General
  • Gold Spring Wash
  • Hummingbird Canyon
  • Ted's Turtle Town
  • Gerald's Turtle
Silty Facies Member
  • TMM 43669-1, 2, 4
  • TMM 436701–3
  • TMM 43656-1, 4
  • TMM 45608-2
  • TMM 43687-1, 4–5, 12, 21, 25–35, 37–41, 47–49, 54
  • TMM 43687-24, 43, 54
A mesochelydian
TestudinataIndeterminate
  • Airhead West
  • Gold Spring General
  • Moenkopi Point
  • Valley of the Buttes
  • Paiute Canyon General
  • East Paiute Valley
Silty FaciesUncertain fragmentsTestudinatans of uncertain assignment

Crocodylomorphs

Genus / TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
Calsoyasuchus[18] C. vallicepsAdeii Eechii Cliffs, Navajo NationSilty FaciesTMM 43631-1 (holotype), partial skullA terrestrial member of the Hsisosuchidae. Alternatively can be a relative of Thalattosuchia
Crocodylomorpha
  • "Undescribed new genus"
  • Indeterminate
  • Utah Route 7 near St.George
  • Airhead West
  • Moenkopi Point, Pumpkin Patch
  • Blue layer, Silty Facies
  • Gold Spring Quarry 1
Silty Facies
  • Teeth (+30)
  • UCMP 97639, 97640
  • MCZ 9044, dermal armor fragment
  • MCZ 9199, fragment of pseudosuchian dermal scute
  • MCZ 9200, dermal scute, probably Eopneumatosuchus
  • UCMP 136102, fragment of lower jaw
Indeterminate crocodylomorphs. Includes a new taxon with skull similar to Orthosuchus stormbergi.
Eopneumatosuchus[19] E. colbertiBlue layer, Silty FaciesSilty Facies
  • MNA P1.2460, partial skull
  • Isolated Jaw
An early terrestrial or semiterrestrial protosuchid crocodylomorph
Kayentasuchus[20] [21] K. walkeri
  • Warner Valley
  • Willow Springs 13
Silty Facies
  • UCMP 131830, nearly complete skeleton
  • UMNH VP 21923
An early terrestrial or semiterrestrial crocodylomorph
Protosuchidae[22]
  • "Gomphosuchus wellesi"
  • "Edentosuchus-like taxon"
  • Moenkopi Point, Pumpkin Patch
  • Blue layer, Silty Facies
Silty Facies
  • MCZ 8816, mandible
  • UCMP 97638, A skull and articulated mandible
  • UCMP 125395: A cranium.
  • UCMP 125871: Skull with mandibles and one epibranchial lacking the dorsal part of the braincase, articulated with the atlas, axis, and 2 cervical vertebrae.
  • UCMP 125358
  • UCMP 125359: An eroded compressed braincase.
  • UCMP 125872: A right jugal and maxilla in articulation in a large block of unprepared material.
  • UCMP 125870: A very well-preserved braincase
  • UCMP 130082
Early terrestrial or semi-terrestrial herbivorous Crocodylomorphs. Includes two taxa similar to the Cretaceous Edentosuchus tienshanensis, one that has been referred to informally as "Gomphosuchus" (including UCMP 97638 and UCMP 125871) and another unnamed taxon (including UCMP 130082).[23] [24] Previously considered one taxon (the 'Kayenta form') in older literature.

Dinosauria

Indeterminate ornithischian remains located in Arizona, USA. Ornithischian tracks located in Arizona, USA.[25] Indeterminate theropod remains located in Arizona, US. Theropod tracks located in Arizona and Utah, US. Possible theropod tracks located in Arizona, Colorado, and Utah, US.

Genus / TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
Coelophysidae[26] Unnamed, informally known as the "Shake-N-Bake" coelophysid
  • Rock Head (Bowl Area)
  • Shake-N-Bake
Silty Facies
  • MCZ 8817b dorsal vertebra, partial sacrum, partial pelvis
  • MCZ 8817c two partial mid cervical vertebrae
  • MCZ 8817d incomplete posterior cervical vertebra
  • MCZ 8817e proximal caudal vertebra
  • MCZ 8817f) partial tooth, partial caudal centrum
  • MCZ 8817g) anterior dorsal vertebra
  • MCZ 8817h) incomplete anterior dorsal vertebra
  • MCZ 8817i) partial sacrum
  • MCZ 8817j) proximal femur
  • MCZ 8817k) (robust) proximal femur
  • MCZ 8817l) (gracile) proximal femur
  • MCZ 8817m) distal tibiotarsus
  • MCZ 8817n) distal tibiotarsus, distal fibula
  • MCZ 8817o) distal tibiotarsus, distal fibula
  • MCZ 8817p) distal tibiotarsus
  • MCZ 8817q) distal tibia, partial astragalus
  • MCZ 8817r) partial scapulocoracoid
  • MCZ 8817t) partial scapulocoracoid
  • MCZ 9442; = 8817a) (adult) sacrum, partial ilia, proximal pubes, proximal ischia
  • MCZ 9463; = 8817m-p?) (adult) distal tibiotarsus
  • TMM 43689-4) (adult) proximal tarsometatarsus
  • MNA V3181, pubis
A coelophysid neotheropod.
Coelophysis[27] [28] C. kayentakatae
  • Rock Head (Bowl Area)
  • Gold Spring Wash
  • Shake-N-Bake
Silty Facies
  • MNI; MNA V2623
  • TMM 43669-3
  • MNA V100, V140
A coelophysid neotheropod. Referred to as Syntarsus by Weishampel et al. Formerly known as Megapnosaurus.
Dilophosaurus[29] [30] D. wetherilli
  • Tuba City, Silty Facies
  • Dilophosaurus Quarry
  • Gold Spring East
  • Gold Spring General
  • Rock Head (Bowl Area)
  • Moenkopi Point, Pumpkin Patch
Silty Facies
  • UCMP 37302 (holotype), nearly complete skeleton
  • UCMP 37303, partial skeleton; third skeleton eroded and not collected
  • MNA V3145, disal end of R femur
  • TMM 43687-52
  • UCMP 77270
  • UCMP 130053
  • TMM 43646-0, 1
  • TMM 47006-1
An advanced neotheropod, type member of the family Dilophosauridae. Dilophosaurus is the main identified dinosaur from the formation, being both the most known and studied. It was among the largest theropods present locally, and very likely an active hunter, rather than a fisher.
Heterodontosauridae[31] [32] Heterodontosauridae indeterminateGold SpringsSilty FaciesMCZ 9092, Partial Skull & Associated Remains (complete upper and lower dentitions, many other portions of the skull, vertebrae from all portions of the axial column, and portions of fore and hind girdles and limbs)A heterodontosaurid of uncertain placement. Its presence in North America suggest early dispersal of the family.
Kayentavenator[33] K. elysiaeWillow SpringsSilty FaciesUCMP V128659, six proximal caudal centra, three centra, two partial neural arches, fragmentary ilium, proximal pubes, pubic shaft fragments, incomplete femora, proximal tibiae, proximal fibula, fragmentsA neotheropod of uncertain relationships, probably a coelophysoid. Originally referred to M. kayentakatae by Rowe.
Ornithischia[34] [35]
  • Utah Route 7 near St.George
  • Gold Spring Quarry
Silty Facies
  • MNA.V.109, a large left femur
  • Isolated Teeth (+20)
A uncertain placement large ornithischian and teeth from diverse type of genera. The femur was assigned to Dilophosaurus wetherilli. The femur resembles that of the early neornithischian Lesothosaurus.
Sarahsaurus[36] S. aurifontanalis
  • Gold Springs
  • Rock Head
Silty Facies
  • TMM 43646–2 partially articulated skeleton
  • TMM 43646–3, partial skeleton
  • MCZ 8893, articulated skull with fragmentary postcranial elements
A sauropodomorph, a member of the family Massospondylidae. Originally thought to be Massospondylus
Scelidosaurus[37] [38] S. sp. (S. "arizonensis")Valley of the ButtesSilty FaciesUCMP 130056, scutesA controversial thyreophoran, resembles the osteoderms of S. harrisonii.
Scutellosaurus[39] [40] S. lawleri
  • Paiute North 1
  • West Moenkopi Plateau
  • Paiute Canyon General
  • East Paiute Valley
  • Southwest Paiute Canyon
  • Rock Head
  • Willow Spring General
  • Gold Spring General
  • Gold Spring Quarry
  • Gold Spring South
  • Gerald's Turtle
  • Ted's Turtle Town
  • Hummingbird Canyon
Silty Facies
  • MNA P1.175, almost complete skeleton (holotype)
  • MNA P1.1752, partial skeleton (paratype)
  • TMM 43669-5, 6
  • TMM 43661-1
  • TMM 43691-18
  • TMM 43691-20
  • TMM 43648-13
  • TMM 43663-1
  • TMM 43664-1, 2
  • TMM 47001-1
  • TMM 43690-6
  • TMM 43687-13, 117, 123
  • TMM 43687-9, 16, 17, 22, 42, 50, 57, 75, 81, 96, 112, 114–116, 121–122, 124
  • TMM 45609-10, 11
  • TMM 45609-4, 6
  • TMM 45609-5, 7–9
  • TMM 43669-11
  • TMM 45608-1, 3
  • TMM 43647-7
  • TMM 43656-5
  • TMM 43670-5, 7–8
A basal thyreophoran, the most abundant dinosaur of the formation
Theropoda[41] Indeterminate
  • Utah Route 7 near St.George
  • Airhead West
  • Gold Spring General
  • Gold Spring Wash
Silty Facies
  • Isolated Teeth (+50)
  • TMM 43669-10
  • TMM 43687-10, 58, 60, 71, 85, 91, 98, 102, 105, 119
Incertae sedis within Theropoda, probably Neotheropoda

Pterosauria

Genus / TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
PterosauriaIndeterminate
  • Utah Route 7 near St.George
Silty Facies
  • Teeth
Possible pterosaur teeth
Rhamphinion[42] R. jenkinsi
  • Airhead West
  • MCZ 23/78A, Foxtrot Mesa
Silty Facies, Ward Mesa
  • MNA V 4500 (holotype), skull fragments
  • UCMP 128227, left fourth wing metacarpal
A pterosaur, considered a member of the family Dimorphodontidae. Was originally classified as a "rhamphorhynchoid", represents the only major pterosaur identified from the formation and one of the oldest from North America.

Synapsida

Genus / TaxonSpeciesLocationStratigraphic positionMaterialNotesImages
Dinnebitodon[43]
  • D. amarali
  • D. spp.
  • Dinnebito Wash
  • Hummingbird Canyon
  • Gold Spring General
Silty Facies
  • MNA V3222 (Type, partial skull and associated postcrania); MNA V3223, partial skull and scapula
  • MCZ 8831 includes two left dentaries
  • MCZ 8830, Crushed snout
  • TMM 43647-3, 4
  • TMM 43687-7
A relatively large and common tritylodont
DinnetheriumD. nezorum
  • Gold Spring Quarry 1
Silty FaciesMNA V3221; MCZ 20870-20877A mammaliaform, member of the family Megazostrodontidae
HaramiyidaeIndeterminate
  • Gold Spring Quarry 1
Silty FaciesMCZ 20879Incertae sedis, a possible haramiyid
Kayentatherium[44]
  • K. wellesi
  • K. spp.
  • MCZ, The Landmark
  • Hummingbird Canyon
  • Gold Spring General
  • Gold Spring wash
  • Willow Spring General
  • Rock Head, general area
Silty Facies
  • MCZ 8811
  • TMM 43669-9
  • MCZ 8842
  • TMM 43647-9, 10
  • TMM 43687-111
A Large tritylodont, with a suggested semiaquatic mode of life. A specimen has been recovered with several associated perinates.[45]
Morganucodon[46] M. sp.
  • Gold Spring Quarry 1
Silty FaciesMCZ 20878A mammaliaform, member of the family Morganucodontidae
Oligokyphus[47]
  • O. sp.
  • cf. O. sp.
  • Gold Spring General
  • Gold Spring Quarry 1
Silty Facies
  • TMM 43687-86
  • > 42 specimens
A tritylodont, also present on coeval deposits from Asia and Europe.
TritylodontidaeIndeterminate
  • Airhead West
  • MCZ, The Landmark
  • Rock Head, general area
  • Willow springs
  • Gold Spring General
  • Gold Spring 1
Silty FaciesTeethIndeterminate tritylodontid remains

Ichnofossils

GenusSpeciesLocationMaterialTypeOriginNotesImages
Anomoepus
  • A. scambus
  • A. shingi
  • A. moabensis
  • A. isp.
  • Moenkopi Wash
  • Lisbon Valley Oilfield tracksite
  • Hamblin tracksite
  • Warner Valley tracksite
  • Utah Route 7 near St.George
  • Poison Spider Mesa tracksite
FootprintsMoving Tracks
  • Ornithischians
Ornithischian Footprints of the ichnofamily Moyenisauropodidae.
Batrachopus[48]
  • B. isp.
  • Desert Tortoise tracksite
TracksMoving Tracks
  • Crocodrylomorphs
Pseudosuchia Footprints of the ichnofamily Batrachopodidae.
Characichnos
  • C. isp.
  • Utah Route 7 near St.George
TracksMoving Trails
  • Dinosaurs
Dinosaur Traces left while swimming
Dilophosauripus
  • D. williamsi
  • D. isp.
  • Moenkopi Wash 4 tracksite
  • Moenave Road Tracksite
  • Goldtooth Spring tracksite
  • Cameron tracksite
FootprintsMoving Tracks
  • Theropods (Dilophosaurus?)
Theropod Footprints of the ichnofamily Grallatoridae.
Dinosauropedida[49] [50] Indeterminate
  • North Creek tracksite
  • Parunuweap-West Temple tracksite
  • Tenmile Canyon tracksite
FootprintsMoving Tracks
  • Dinosaurs
Possible Dinosaur Footprints, non assigned to any concrete ichnogenus
Eubrontes[51] [52]
  • E. giganteus
  • E. isp.
  • Washington City Water Tank tracksite 1
  • Flat Iron Mesa tracksite
  • Trout Water Canyon tracksite
  • Grapevine Pass Wash Tracksite
  • Warner Valley tracksite
  • Desert Tortoise tracksite
  • Flag Point I tracksite
  • Rainbow Bridge tracksite
  • Explorer's Canyon tracksite
  • Mike's Mesa tracksite (Kayenta)
  • Utah Route 7 near St.George
  • Cactus Park track site
FootprintsMoving Tracks
  • Theropods (Dilophosaurus?)
Theropod Footprints of the ichnofamily Grallatoridae. This type of tracks match with Dilophosaurus pes
Grallator[53]
  • G. tenuis
  • G. isp.
  • Lisbon Valley Oilfield tracksite
  • Washington City Water Tank tracksite 1
  • Exit 13 North tracksite
  • Exit 13 South tracksite
  • Desert Tortoise site
  • Warner Valley tracksite
  • Desert Tortoise tracksite
  • Flag Point II tracksite
  • Utah Route 7 near St.George
  • Flat Iron Mesa tracksite
  • Cactus Park track site
FootprintsMoving Tracks
  • Theropods (Coelophysoids?)
Theropod Footprints of the ichnofamily Grallatoridae. Likely from smaller local theropods
Kayentapus[54] [55]
  • K. hopii[56]
  • K. soltykovensis
  • K. isp.
  • Moenkopi Wash
  • Trout Water Canyon tracksite
  • Desert Tortoise tracksite
  • Flag Point I tracksite
  • Flag Point II tracksite
  • Flat Iron Mesa tracksite
FootprintsMoving Tracks
  • Theropods
Theropod Footprints of the ichnofamily Grallatoridae.
Limulidae[57] Indeterminate
  • Lisbon Valley Oilfield tracksite
TrackwaysMoving Tracks
  • Limuloids
  • Insects?
Saltwater/Blackish-linked tracks with resemblance with extant Xiphosuran traces
Moyenisauropus
  • M. isp.
  • Utah Route 7 near St.George
FootprintsMoving Tracks
  • Thyreophorans
Ornithischian Footprints of the ichnofamily Moyenisauropodidae.
Otozoum[58] [59]
  • O. isp.
  • Flat Iron Mesa tracksite
  • Poison Spider Mesa tracksite
FootprintsMoving Tracks
  • Sauropodomorphs
Theropod Footprints of the ichnofamily Otozoidae. Includes tracks referable to bipedal Sauropodomorphs
Planolites[60]
  • P. isp.
  • Near St. George, Washington County
Cylindrical burrows Pascichnia
  • Annelids
Burrow-like ichnofossils. It is referred to vermiform deposit-feeders. It is controversial, since is considered a strictly a junior synonym of Palaeophycus.[61]
Skolithos
  • S. isp.
  • Near St. George, Washington County
Cylindrical to subcylindrical Burrows Domichnia
  • Annelids
  • Crustaceans
  • Fishes
Burrow-like ichnofossils. Ichnofossils done by organisms advancing along the bottom surface. Very narrow, vertical or subvertical, slightly winding unlined shafts filled with mud. Interpreted as dwelling structures of vermiform animals, more concretely the Domichnion of a suspension-feeding Worm or Phoronidan, with certain Skolithos representing entrance shafts to more complicated burrows.
SynapsidipediaIndeterminate
  • Lisbon Valley Oilfield tracksite
TracksMoving Tracks
  • Mammaliformes
  • Tritylodonts
Possible Synapsid Footprints, non assigned to any concrete ichnogenus
Taenidium
  • T. isp
  • Near St. George, Washington County
Unlined meniscate burrowsFodinichnia
  • Deposit-feeding Sipuncula
  • Annelids
Saltwater/Blackish burrow-like ichnofossils. Taenidium is a meniscate backfill structure, usually considered to be produced by an animal progressing axially through the sediment and depositing alternating packets of differently constituted sediment behind it as it moves forward.
Theropodipedia[62] [63] Indeterminate
  • Lisbon Valley Oilfield tracksite
  • Hamblin tracksite
  • South Gate tracksite
  • Long Canyon tracksite
  • Washington City Water Tank tracksite 2
  • Lion's Back tracksite
FootprintsMoving Tracks
  • Theropods
Possible Theropod Footprints, non assigned to any concrete ichnogenus
Undichna
  • U. isp.
  • Utah Route 7 near St.George
TrailsMoving Trails
  • Bony Fishes
Fish-swimming fossil trail left as a fossil impression on a substrate

Plants

GenusSpeciesStratigraphic positionMaterialNotesImages
Clathropteris
  • C. sp.
  • Utah Route 7 near St.George
  • Leaflets
Affinities with Dipteridaceae inside Polypodiales.
CycadidaeIndeterminate
  • Utah Route 7 near St.George
  • Leaflets
Affinities with Cycadidae inside Cycadopsida.
Otozamites
  • O. sp.
  • Utah Route 7 near St.George
  • Leaflets
Affinities with Williamsoniaceae inside Bennettitales.
PinopsidaIndeterminate
  • Utah Route 7 near St.George
  • Leave Compressions
  • Isolated Cones
Affinities with Pinopsida inside Pinaceae.
Zamites
  • Z. powellii
  • Utah Route 7 near St.George
  • Leaflets
A member of Williamsoniaceae inside Bennettitales. It has been interpreted as a cycad in the family Cycadaceae or a Bennettitalean plant, and also a late surviving member of Noeggerathiales.

See also

References

External links

Notes and References

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