Iris pallida subsp. cengialti explained

Iris pallida subsp. cengialti is a subspecies in the genus Iris, it is also in the subgenus Iris. It is a rhizomatous perennial, from Italy and (part of the former country of Yugoslavia) Slovenia. It has yellowish-green, glaucous, lanceolate or ensiform leaves, tall stem, green flushed with purple spathes, 2 short branches, 2–3 scented flowers, in shades of violet, blue-violet, deep purple, blue-purple, deep blue-purple, pale purple, deep blue, to mid-blue. It has a yellow or orange tipped beard. It was originally published as Iris cengialti but then re-classified as a subspecies of Iris pallida, and known as Iris pallida subsp. cengialti, but it is often still called Iris cengialti. It is cultivated as an ornamental garden plant in temperate regions.

Description

It has a long,[1] [2] stout,[3] fleshy, light-coloured (underground) rhizome.[1] That is 1–3 cm wide (in diameter),[1] [2] and has long secondary roots.[2] It forms creeping plants.[3]

It has yellowish-green,[3] [4] lanceolate,[1] or ensiform (sword-shaped),[3] leaves, that are glaucous.[3] [5] The basal leaves,[2] can grow up to between 15and long, and about 1.3 cm wide.[1] [3] [4] The herbaceous leaves (die in the winter),[3] sheath the stem.[1] [2]

It has a round (in section) stem,[1] [2] or peduncle, that can grow up to between 30and,[6] [7] [8] or 20- tall.[1] [9] The stems are taller than the leaves,[3] [4] and at higher levels on the mountains, the plants are shorter.[5] It is also shorter than Iris pallida.[10]

The stem has glaucous green and ensiform spathes (leaves of the flower bud).[2] They are slightly flushed with purple,[4] and before flowering, they become pale brown,[5] (scarious) membranous,[3] and papery,[4] They are 2.5 cm long,[3] and between 2and wide.[2]

It has 2 short branches (or pedicels).[3] [4] The stems (and the branches) hold between 2 and 3 flowers,[2] [3] [10] It can have up to 6 flowers,[7] but normally has 3 flowers,[4] in spring, between April and June,[2] or May,[11] to July.[9]

The small,[5] scented flowers,[2] [7] are 6- in diameter,[1] come in shades of blue, from violet,[10] blue-violet,[1] [7] [12] deep purple,[13] blue-purple,[2] [3] [14] deep blue-purple,[6] [8] [15] pale purple,[2] deep blue,[9] to mid-blue.[16]

Like other irises, it has 2 pairs of petals, 3 large sepals (outer petals), known as the 'falls' and 3 inner, smaller petals (or tepals), known as the 'standards'.[1] [2] [17] The falls are obovate or cuneate shaped,[2] [3] they are 5- long and 2.5- wide.[2] [3] In the centre of the falls, is a short,[4] (16–30 mm,[2]) thick row of small hairs (the beard), which is yellowish-white,[1] [2] [6] bright yellow,[15] or white with orange tips.[3] [4] [8] The standards are a similar size to the falls.[3]

It has 2.5 cm long style branch, that is paler than the falls and standards, and has deltoid crests.[3] It also has a six grooved, ellipsoid ovary,[4] and a 0.6–0.8 cm long perianth tube.[3] [4]

After the iris has flowered, it produces an ovoid seed capsule, that is 3- long and 1 cm wide, with 3 sections, which contain 15–20 oval grey or yellowish seeds.[1] [2]

Genetics

In 1956, a karyotype analysis was carried out on 40 species of Iris, belonging to the subgenera Eupogoniris and Pogoniris. It found that 24-chromosome tall bearded species could be divided into 3 karyotypes of Iris pallida. Iris kashmiriana has 2 pairs of median-constricted marker chromosomes, Iris illyrica, Iris cengialti, and Iris imbricata, lastly Iris variegata, Iris reginae (later classified as a synonym of Iris variegata), and Iris perrieri all have no median-constricted chromosomes.[18]

As most irises are diploid, having two sets of chromosomes, this can be used to identify hybrids and classification of groupings.[17] It has a chromosome count: 2n=24.[19]

Taxonomy

It is commonly known as 'Iris del Cengio',[1] 'Monte Cengio iris'.[2] [11]

In Italy, it is known as 'Giaggiolo del Cengio Alto'.[20] In Slovenia, it is known as 'Bohinj Iris',[9] [21] [22] or 'Bòcje of lof'.[1] In German, it is known as 'Rovereto Schwertlilie'.[23]

It is sometimes misspelt as Iris cengualti.[24]

The Latin specific epithet cengialti refers to a mountain or large hill 'Mount Cengialto' (also known as Monte 'Cengio Alto',[2] [25]), close to the town of Rovereto, in northern Italy. Confusingly, another mountain near Arsiero in the province of Vicenza (Veneto), is also known as Monte Cengio Alto.[2]

It was found by Francesco Ambrosi in 1854,[16] on Monte Cengialto,[3]

It was then published as Iris cengialti by Ambrosi in Flora del Tirolo Meridionale Vol.1 (Fl. Tirolo Mer.) on page 643 in 1854,[26] then fully published and described by A. Kerner in Österreichische Botanische Zeitschrift (Oesterr. Bot. Z.) Vol. 21, No.9, on pages 225–231 in September 1871.[4] [27] [28] A. Kerner was then assigned the author of the name Iris cengialti, previously ascribed to Ambrosi.[29]

Iris pallida, Iris cengialti and Iris illyrica are all considered closely related but often given different taxonomic rank (by various authors Kerner, 1871; Ambrosi, 1854; Foster, 1886; Tommasini,1875; Pampanini, 1909; Lausi, 1964; Mathew, 1981; Colasante, 1995: Terpin et al., 1996).[30]

In May 1886, Sir Michael Foster noted in Gardeners' Chronicle on pages554 and 555, that it had similarities with Iris pallida.[3] [4] William Rickatson Dykes in his book in 'Handbook of Garden Irises' in 1914, suggested that Iris cengialti and Iris pallida were connected and should be merged under Iris pallida.[5] In 1939, Brian Mathew considered it to be a subspecies of Iris pallida.[4]

Then in Giorn. Bot. Ital. Vol.130 on page575 in 1996, Iris cengialti was classified as a synonym of Iris pallida subsp. cengialti.[29]

It is normally classified as Iris pallida subsp. cengialti,[1] [16] [31] and thought to be a naturally occurring hybrid.[16]

In Slovenia, a form of the plant is called Iris cengialti vochinensis,[21] or Iris pallida subsp. cengialti f. vochinensis.[22]

probably of Asia Minor origin,[12]

It was verified as Iris pallida subsp. cengialti by United States Department of Agriculture and the Agricultural Research Service on 27 February 2002, and then updated on 1 December 2004.

It is listed in the Encyclopedia of Life,[32] and listed on the Catalogue of Life as Iris pallida subsp. cengialti.[33]

Distribution and habitat

It is native to Europe.

Range

It endemic in the alpine region,[1] [11] [24] on the Dolomites (mountain range).[34]

It is found in north east Italy,[6] [10] (within the region of Trentino,[35] and Veneto)[14] [36] and (part of the former Yugoslavia),[7] [12] in Slovenia,[6] [8] [14] (within the regions of Bohinj. and Istria).[23]

In Trentino, it has found on Monte Zugna (south of Rovereto),[37] and Mount Brione. In Veneto, it can be found on Mount Summano,[14] in the province of Belluno.[36]

Habitat

It grows on the rocky mountain sides,[2] [9] [11] on scree,[1] in meadows,[1] [11] [22] and dry grasslands.[2] Usually on karst,[1] or limestone soils.[2]

They can be found at an altitude of 200- above sea level.[1] [2] [11]

They can found in the Dolomites, with other plants including; alyssum ovirense, androsace villosa, centaurea haynaldii, crepis froelichiana subsp. dinarica, eritrichium nanum, euphorbia kerneri, geranium argenteum, leontodon berinii, lilium carniolicum, potentilla incana and thlaspi minimum.[34] They also grow in the Belluno forests of Acer pseudoplatanus (Mount Maple) and Tilia platyphyllos (Lime tree) with other geophytes and Leontodon tenuiflorus (Dandelion insubrico), Knautia persicina, Globularia nudicaulis and Paeonia officinalis (Wild Peony).[36] In Slovenia, in Bohinj, it grows in meadows with another rare plant, Linnaea borealis (twinflower).[22]

Conservation

It is a rare plant,[14] although locally it is abundant on the mountains,[11] growing in masses, hanging onto relatively steep slopes.

In Italy, it is listed as a protected species of regional interest (Annex B, LR 9/2007, Art. 96), and would be classed as 'vulnerable' on the IUCN Red List.[1] [2]

Cultivation

It is hardy to 0 °C.[15]

It prefers could grown in rich (in fertile loan,[15]), well drained soil in full sun.[10] [15]

It can be grown the rock garden.[10]

It is generally not attacked by pests and diseases.[10]

A herbarium specimen can be found at the Museum National d'Histoire Naturelle in Paris.[35]

Propagation

Irises can generally be propagated by division.[38] They should be lifted every three or four years, after the flowers are over. The tuber can be divided, with the dead roots trimmed and removed. The remains can then be replanted in enriched soil (with added compost). The leaves should be cut back to half their length, to prevent wind rock.[10]

Hybrids and cultivars

It has been used in hybridizing (or breeding programmes), and the hybrids that have Iris cengialti as a parent, can have multiple flowers, on average between 5 and 8. The branches are generally short on slender stems.[39]

Toxicity

Like many other irises, most parts of the plant are poisonous (rhizome and leaves), if mistakenly ingested can cause stomach pains and vomiting. Also handling the plant may cause a skin irritation or an allergic reaction.[40]

Sources

External links

Notes and References

  1. Web site: Iris cengialti Ambrosi former A.Kern. subsp. illyrica (Asch. & Graebn.) Poldini . flora.uniud.it . 17 November 2015.
  2. Web site: Silvano . Radivo . Iris cengialti . 24 June 2008 . it . actaplanatarum.org . 19 November 2015 . 16 March 2015 . https://web.archive.org/web/20150316022439/http://www.actaplantarum.org/floraitaliae/viewtopic.php?t=5242 . dead .
  3. Richard Lynch
  4. Web site: Alain . Franco . (SPEC) Iris cengialti Ambrosi . 30 November 2014 . wiki.irises.org (American Iris Society) . 6 November 2015.
  5. Web site: William . Dykes . William Rickatson Dykes . Handbook of Garden Irises . 2009 . beardlessiris.org (The Group for Beardless Irises) . 1 November 2014.
  6. Web site: Iris summary . 14 April 2014 . pacificbulbsociety.org . 23 November 2014.
  7. Book: Cassidy . George E.. Linnegar . Sidney . 1987 . Revised . 121 . Growing Irises . Bromley . Christopher Helm . 978-0-88192-089-5.
  8. Web site: D. . Kramb . Iris pallida subsp. cengialtii . 30 September 2004 . signa.org (Species Iris Group of North America) . 19 November 2015.
  9. Web site: Bohinj Iris (Iris cengialti f. Vochinensis) . hribi.net . sl . 6 November 2015.
  10. Web site: Bob . Mitchell . Plant of the Month June 2015 Iris pallida (Orris root) . June 2015 . st-andrews.ac.uk . 19 November 2015.
  11. Web site: Iris of Mount Cengio . it . dolimitipark.it . 19 November 2015.
  12. Bezzi . A. . Aiello . N. . Villa . S. . Bicchi . C. . Rubiolo . P. . 1993 . Productivity And Quality Of Rhizomes Of Some Different Types Of Iris Sp. . Acta Horticulturae . 344 . 344. 98–109 . 10.17660/ActaHortic.1993.344.11 .
  13. Web site: Dalmatian iris . mediteranka.com . 16 November 2015.
  14. Web site: Le Tre Venezie – THIENE . letrevenezie.net . 17 November 2015.
  15. Web site: Iris cengialti . rareplants.co.uk . 6 November 2015.
  16. Web site: Cengialti . historiciris.org . 6 November 2015 . 17 April 2015 . https://web.archive.org/web/20150417031025/http://historiciris.org/photos/cengialti-lm.html . dead .
  17. Book: Austin, Claire . Irises; A Garden Encyclopedia . 2005 . Timber Press . 978-0881927306 .
  18. Mitra . Jyotirmay . 1956 . Karyotype analysis of bearded iris . Bot Gaz . 117. 4. 265–293 . 10.1086/335916. 85117341 . 12 September 2015.
  19. Löve . Áskell . November 1971 . IOPB Chromosome Number Reports XXXIV . 1218605 . Taxon . 20 . 5 . 785–797 . 10.1002/j.1996-8175.1971.tb03208.x.
  20. Web site: Giaggiolo del Monte Cengio Alto (Iris pallida ssp. cengialti) . fiorialpini.ch . 18 November 2015.
  21. Web site: Let's Walk Together to the Iris of Bohinj (Rudnica) . sl . bohinj.si . 17 November 2015.
  22. Web site: Bohinj Slovenia . bohinj.si . sl . 18 November 2015.
  23. Web site: Paul . Tout . Istria 8th–16th May 2013 . 2013 . 18 November 2015.
  24. Jevremović . Slađana . Subotić . Angelina . Trifunović . Milana . Nikolić . Marija . 2009 . Plant Regeneration Of Southern Adriatic Iris By Somatic Embryogenesis . Arch. Biol. Sci. . 61 . 3 . 413–418 . 10.2298/ABS0903413J . 19 November 2015. free .
  25. Web site: Cengio Alto . it.geoview.info . 19 November 2015.
  26. Web site: Iridaceae Iris cengialti Ambrosi . ipni.org (International Plant Names Index) . 19 November 2014.
  27. Web site: Iridaceae Iris cengialti Ambrosi ex A.Kern. . ipni.org . 19 November 2014.
  28. Kerner . A. . September 1871 . Ueber Iris Cengialli Ambrosi . 43333505 . Österreichische Botanische Zeitschrift . 21 . 9 . 225–231 . 10.1007/BF01615257. 12472346 .
  29. Terpina . Katia . Nardib . Enio . Garbaria . Fabio . 1996 . Author and type of the name Iris cengialti (Iridaceae) . Giornale Botanico Italiano . 130 . 2 . 575–578 . 10.1080/11263509609430326 .
  30. Colasante . M. . Vosa . C.G. . 2000 . Iris:Allocyclic segments as Chromosome markers? . Annali di Botanica . 58 . 127–134 . 28 October 2015 .
  31. Elaine Nowick
  32. Web site: Iris pallida subsp. cengialti . eol.org . 17 November 2015.
  33. Web site: Infraspecific taxon details : Iris pallida subsp. cengialti (Ambrosi ex A.Kern.) Foster . catalogueoflife.org . 17 November 2015.
  34. Erika Pignatti and Sandro Pignatti
  35. Web site: Iris cengialti (MNHN/P/P02158427) . lesherbonautes.mnhn.fr . 6 November 2015.
  36. Web site: Cengia Prada . arpa.veneto.it . it . 19 November 2015 . 12 April 2022 . https://web.archive.org/web/20220412003834/https://www.arpa.veneto.it/rete-ea/retedamb_area.php?id=286 . dead .
  37. Web site: [ftp://ftp.minambiente.it/PNM/Natura2000/TrasmissioneCE_2014/schede_mappe/Trento/ZSC_schede/Site_IT3120114.pdf NATURA 2000 – STANDARD DATA FORM ]. September 2013 . minambinente.it .
  38. Web site: How to divide iris rhizomes . gardenersworld.com . 12 October 2015.
  39. Benjamin Yoe Morrison
  40. David G Spoerke and Susan C. Smolinske