Helicostyla Explained

Helicostyla is a genus of small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Helicostylinae of the family Camaenidae. Helicostyla is the type genus of the subfamily Helicostylinae.^[1]

Distribution

This genus is endemic to the Philippines.

Species

Species within the genus Helicostyla include:^{[2] [3]}

• Helicostyla albina (Grateloup, 1840)
• Helicostyla amagaensis de Chavez, Kendrich, Fontanilla, Batomalaque & Chiba, 2015
• Helicostyla amaliae (Möllendorff, 1890)
• Helicostyla aplomorpha (Jonas, 1843)
• Helicostyla bicolorata (I. Lea, 1840)
• Helicostyla boettgeriana (Möllendorff, 1888)
• Helicostyla bullula (Broderip, 1841)
• Helicostyla buschi (L. Pfeiffer, 1846)
• Helicostyla bustoi (Hidalgo, 1887)
• Helicostyla caesar (L. Pfeiffer, 1855)
• Helicostyla calamianica (Quadras Moellendorff, 1894)^[4]
• Helicostyla canonizadoi (Bartsch, 1932)
• Helicostyla chocolatina (Poppe, Tagaro & Sarino, 2015)
• Helicostyla cinerosa (L. Pfeiffer, 1855)
• Helicostyla collodes (G. B. Sowerby I, 1841)
• Helicostyla cunctator (Reeve, 1849)
• Helicostyla curta (G. B. Sowerby I, 1841)
• Helicostyla daphnis (Broderip, 1841)
• Helicostyla decorata (Férussac, 1821)
• Helicostyla delicata Fulton, 1903
• Helicostyla dimera (Jonas, 1846)
• Helicostyla domingoi (Bartsch, 1919)
• Helicostyla dimera (Jonas, 1846)
• Helicostyla effusa (Pfeiffer, 1842)
• Helicostyla euconica (Bartsch, 1932)
• Helicostyla fastidiosa (Bartsch, 1932)
• Helicostyla faunus (Broderip, 1841)
• Helicostyla frater (Férussac, 1821)
• Helicostyla fulgens (G. B. Sowerby I, 1841)
• Helicostyla fulgetra (Broderip, 1841)
• Helicostyla gigas (Hidalgo, 1901)
• Helicostyla gilva (Pfeiffer, 1845)
• Helicostyla glaucophthalma (L. Pfeiffer, 1851)
• Helicostyla halichlora (Semper, 1866)
• Helicostyla hololeuca (L. Pfeiffer, 1855)
• Helicostyla huberorum Thach, 2021
• Helicostyla imperator (Pfeiffer, 1848)
• Helicostyla indusiata (L. Pfeiffer, 1871)
• Helicostyla jonasi (L. Pfeiffer, 1846)
• Helicostyla juglans (L. Pfeiffer, 1842)
• Helicostyla lacerata (Semper, 1877) (nomen nudum)
• Helicostyla leai (L. Pfeiffer, 1846)
• Helicostyla leopardus (L. Pfeiffer, 1845)
• Helicostyla leucophaea (G. B. Sowerby I, 1841)
• Helicostyla librosa (L. Pfeiffer, 1857)
• Helicostyla lignaria (L. Pfeiffer, 1842)
• Helicostyla macrostoma (L. Pfeiffer, 1843)
• Helicostyla marinduquensis (Hidalgo, 1887)
• Helicostyla mateoi (Bartsch, 1932)
• Helicostyla mearnsi Bartsch, 1905
• Helicostyla mirabilis (Férussac, 1821)
• Helicostyla montana (C. Semper, 1877)
• Helicostyla mus (Delessert, 1841)
• Helicostyla nobilis (Reeve, 1848)
• Helicostyla nux (C. Semper, 1877)
• Helicostyla okadai Kuroda, 1932
• Helicostyla orbitula (Sowerby, 1841)
• Helicostyla palavanensis (L. Pfeiffer, 1857)
• Helicostyla persimilis (Deshayes, 1850)
• Helicostyla philippinensis (L. Pfeiffer, 1846)
• Helicostyla pictor (Broderip, 1841)
• Helicostyla pithogaster (Ferussac, 1821)
• Helicostyla polita Reeve, 1848
• Helicostyla portii (L. Pfeiffer, 1861)
• Helicostyla rufogaster (Lesson, 1832)
• Helicostyla saranganica (Hidalgo, 1887)
• Helicostyla satyrus (Broderip, 1841)
• Helicostyla schadenbergi (Möllendorff, 1890)
• Helicostyla seckendorffiana (L. Pfeiffer, 1847)
• Helicostyla simonei Thach & F. Huber, 2021
• Helicostyla simplex (Jonas, 1843)
• Helicostyla smaragdina (Reeve, 1842)
• Helicostyla solida (L. Pfeiffer, 1843)
• Helicostyla solivaga (Reeve, 1849)
• Helicostyla speciosa (Jay, 1839)
• Helicostyla tephrodes (L. Pfeiffer, 1843)
• Helicostyla thomsoni (L. Pfeiffer, 1871)
• Helicostyla ticaonica (Broderip, 1841)
• Helicostyla turbo (Pfeiffer, 1845)
• Helicostyla ventricosa (Bruguière, 1792)
• Helicostyla vidali (Hidalgo, 1887)
• Helicostyla villari (Hidalgo, 1887)
• Helicostyla volubilis (Reeve, 1851)
• Helicostyla weberi (Bartsch, 1919)
• Helicostyla woodiana (I. Lea, 1840)
• Helicostyla worcesteri Bartsch, 1909

Species brought into synonymy:

• Helicostyla aegle (Broderip, 1841): synonym of Cochlostyla aegle (Broderip, 1841)
• Helicostyla annulata (G.B. Sowerby I, 1841): synonym of Pachysphaera annulata (G. B. Sowerby I, 1841) (unaccepted combination)
• Helicostyla balteata (G.B. Sowerby I, 1841): synonym of Pachysphaera balteata (G. B. Sowerby I, 1841) (unaccepted combination)
• Helicostyla bembicodes (Pfeiffer, 1851): synonym of Helicobulinus bembicodes (L. Pfeiffer, 1851) (unaccepted combination)
• Helicostyla butleri (Pfeiffer, 1842): synonym of Dryocochlias butleri (L. Pfeiffer, 1842) (unaccepted combination)
• Helicostyla camelopardalis (Broderip, 1841): synonym of Hypselostyla camelopardalis (Broderip, 1841) (unaccepted combination)
• Helicostyla carinata (Lea, 1840): synonym of Hypselostyla carinata (I. Lea, 1840) (superseded combination)
• Helicostyla cincinna (G.B. Sowerby I, 1840): synonym of Hypselostyla cincinna (G. B. Sowerby I, 1841) (unaccepted combination)
• Helicostyla cincinniformis (G.B. Sowerb Iy, 1841): synonym of Hypselostyla cincinniformis (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla collodes (G.B. Sowerby I, 1841): synonym of Cochlostyla collodes (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla concinna (G.B. Sowerby I, 1841): synonym of Hypselostyla concinna (G. B. Sowerby I, 1841) (unaccepted combination)
• Helicostyla curta (G.B. Sowerby I, 1841) : synonym of Cochlostyla curta (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla dactylus (Broderip, 1841): synonym of Hypselostyla dactylus (Broderip, 1841) (superseded combination)
• Helicostyla daphnis (Broderip, 1841): synonym of Cochlostyla daphnis (Broderip, 1841) (superseded combination)
• Helicostyla decora (A. Adams & Reeve, 1850): synonym of Cochlodryas decora (A. Adams & Reeve, 1850) (superseded combination)
• Helicostyla denticulata (Jay, 1839): synonym of Calocochlea denticulata (Jay, 1839) (superseded combination)
• Helicostyla dimera (Jonas, 1846): synonym of Cochlostyla dimera (Jonas, 1846) (superseded combination)
• Helicostyla diana (Broderip, 1841): synonym of Hypselostyla diana (Broderip, 1841) (unaccepted combination)
• Helicostyla dubiosa (Pfeiffer, 1846): synonym of Cochlostyla dubiosa (L. Pfeiffer, 1846) (superseded combination
• Helicostyla euconica (Bartsch, 1932): synonym of Cochlostyla effusa (L. Pfeiffer, 1843)
• Helicostyla evanescens (Broderip, 1841): synonym of Hypselostyla evanescens (Broderip, 1841) (unaccepted combination)
• Helicostyla fenestrata (Sowerby I, 1841): synonym of Pachysphaera fenestrata (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla florida (Sowerby I, 1841): synonym of Cochlodryas florida (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla fulgens (Broderip, 1841): synonym of Cochlostyla fulgens (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla fulgetra (Broderip, 1841): synonym of Cochlostyla fulgetra (Broderip, 1841)
• Helicostyla grandis (Pfeiffer, 1845): synonym of Helicobulinus grandis (L. Pfeiffer, 1845) (unaccepted combination)
• Helicostyla hydrophana G.B. Sowerby I, 1841: synonym of Dryocochlias metaformis (Férussac, 1821) (junior synonym)
• Helicostyla ignobilis Sowerby I, 1841: synonym of Cochlodryas ignobilis (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla iloconensis (Sowerby I, 1841): synonym of Pachysphaera iloconensis (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
• Helicostyla jonasi (Pfeiffer, 1845): synonym of Cochlostyla jonasi (L. Pfeiffer, 1846) (superseded combination)
• Helicostyla juglans (Pfeiffer, 1842): synonym of Cochlostyla juglans (L. Pfeiffer, 1842) (superseded combination)
• Helicostyla leopardus (Pfeiffer, 1845): synonym of Cochlostyla leopardus (L. Pfeiffer, 1845) (superseded combination)
• Helicostyla leucophaea (Sowerby I, 1841): synonym of Cochlostyla leucophaea (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla lignaria (Pfeiffer, 1842): synonym of Cochlostyla lignaria (L. Pfeiffer, 1842) (suspected synonym)
• Helicostyla macrostoma (Pfeiffer, 1843): synonym of Cochlostyla macrostoma (L. Pfeiffer, 1843) (superseded combination)
• Helicostyla marinduquensis (Hidalgo, 1887): synonym of Cochlostyla marinduquensis Hidalgo, 1887 (superseded combination)
• Helicostyla metaformis (Ferussac, 1821): synonym of Dryocochlias metaformis (Férussac, 1821) (misspelling of original name, Corasia, Albers, 1850)
• Helicostyla mindoroensis W. J. Broderip, 1892 : synonym of Chrysallis mindoroensis (Broderip, 1841) (unaccepted generic combination)
• Helicostyla mirabilis (Ferussac, 1821): synonym of Cochlostyla mirabilis (Férussac, 1821) (superseded combination)
• Helicostyla montana Semper, 1891: synonym of Cochlostyla montana C. Semper, 1877 (superseded combination)
• Helicostyla monticula (G.B. Sowerby I, 1841): synonym of Orustia monticula (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
• Helicostyla nimbosa (Broderip, 1841): synonym of Hypselostyla nimbosa (Broderip, 1841) (unaccepted combination)
• Helicostyla olanivanensis (Bartsch, 1913): synonym of Cochlostyla olanivanensis Bartsch, 1913
• Helicostyla opalina (G.B. Sowerby I, 1841): synonym of Phengus opalinus (G. B. Sowerby I, 1841)
• Helicostyla phaeostyla (Pfeiffer, 1856): synonym of Cochlostyla phaeostyla (L. Pfeiffer, 1857)
• Helicostyla pictor (Broderip, 1841): synonym of Cochlostyla pictor (Broderip, 1841) (superseded combination)
• Helicostyla polillensis (Pfeiffer, 1861): synonym of Cochlostyla polillensis (L. Pfeiffer, 1861) accepted as Calocochlea polillensis (L. Pfeiffer, 1861)
• Helicostyla pudibunda Semper, 1891: synonym of Pfeifferia pudibunda (C. Semper, 1877) (unaccepted combination)
• Helicostyla rehbeini (L. Pfeiffer, 1852): synonym of Dryocochlias rehbeini (L. Pfeiffer, 1852) (unaccepted combination)
• Helicostyla rufogaster (Lesson, 1831): synonym of Cochlostyla rufogaster (Lesson, 1832) (superseded combination)
• Helicostyla rustica (Mousson, 1849): synonym of Dryocochlias metaformis rustica (Mousson, 1849)
• Helicostyla sarcinosa (Ferussac, 1821): synonym of Helicobulinus sarcinosa (Férussac, 1821)
• Helicostyla satyra (Broderip, 1841): synonym of Cochlostyla satyrus (Broderip, 1841) (superseded combination)
• Helicostyla simplex (Jonas, 1843): synonym of Cochlostyla simplex (Jonas, 1843) (superseded combination)
• Helicostyla solivaga (Reeve, 1849): synonym of Cochlostyla solivaga (Reeve, 1849) (superseded combination)
• Helicostyla sphaerica (Sowerby I, 1841): synonym of Pachysphaera sphaerica (G. B. Sowerby I, 1841)
• Helicostyla submirabilis Möllendorff, 1897: synonym of Cochlostyla submirabilis Möllendorff, 1897 (unaccepted combination)
• * Helicostyla turbinoides (Broderip, 1841): synonym of Helicobulinus turbinoides (Broderip, 1841) (superseded combination)
• Helicostyla velata (Broderip, 1841): synonym of Hypselostyla velata (W.J. Broderip, 1841)
• Helicostyla ventricosa (Bruguière, 1792): synonym of Cochlostyla ventricosa (Bruguière, 1792) (superseded combination)
• Helicostyla viridostriata (Lea, 1840): synonym of Cochlodryas viridostriata (I. Lea, 1840) (superseded combination)
• Helicostyla woodiana (Lea, 1840): synonym of Cochlostyla woodiana (I. Lea, 1840) (superseded combination)

References

• Bank, R. A. (2017). Classification of the Recent terrestrial Gastropoda of the World. Last update: 16 July 2017

External links

• • Férussac, A.E.J.P.F. d'Audebard de. (1821–1822). Tableaux systématiques des animaux mollusques classés en familles naturelles, dans lesquels on a établi la concordance de tous les systèmes; suivis d'un Prodrome général pour tous les mollusques ou fluviatiles, vivantes ou fossiles. Paris, 1821 et 1822.
• Pilsbry, H. A. (1896). On the names of certain subgenera of Helicostyla. The Nautilus. 9(9): 108.
• Albers, J. C. (1850). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet. Berlin: Enslin. 262 pp
• Kennard, A. S. (1942). The Histoire and Prodrome of Férussac. Part III. The divisional names. Proceedings of the Malacological Society of London. 25 (3): 111–118. London
• Pilsbry, H. A. (1903). The use of the generic name Helicostyla. The Nautilus. 17(5): 58
• Albers, J. C.; Martens, E. von. (1860). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet von Joh. Christ. Albers. Ed. 2. Pp. i–xviii, 1–359. Leipzig: Engelman

Notes and References

1. MolluscaBase eds. (2021). MolluscaBase. Helicostyla Férussac, 1821. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=993127 on 2 February 2021
2. http://malacos.chez.com/htm/J99.HTM Bradybaenidae
3. https://www.biolib.cz/cz/taxon/id18127/ Biolib
4. Springsteen, F.J. & Leobrera, F.M. 1986. Shells of the Philippines. Carfel Seashell Museum, Philippines: 1–377.