Celyphidae Explained
The Celyphidae, commonly known as beetle flies or beetle-backed flies, are a family of flies (order Diptera). About 115 species in about 9 genera are known chiefly from the Oriental and Afrotropic biogeographic regions with one lineage in the New World.
Description
Celyphidae are small to medium-sized and easily recognised. The scutellum is enlarged, and forming a protective shell over the abdomen, giving them a beetle-like appearance. Also, like many beetles, Celyphidae are often shiny or metallic in color. The head has few bristles. The wings, when at rest, are folded beneath the scutellar "shell". The arista of the antenna is often flattened and leaf-like at the base. The family name is derived from the Greek word κέλυφος for pod or shell. Male celyphids lack an aedeagus and instead have paired gonapophyses that are used in copulation and are of key taxonomic value.[1] [2]
Biology
The biology of the family is poorly known. Adults are found along streams and rivers, and in wet, grassy areas. Larvae are saprophagous.
Relatives
The family Celyphidae is considered by most authors to be the sister taxon of the Lauxaniidae (e.g., Griffiths 1972), and this has been supported by some molecular studies which suggest the (Chamaemyiidae + (Lauxaniidae + Celyphidae)) within the Lauxanoiodea.[3] In the past they have occasionally been considered a specialized lineage within the Lauxaniidae.[4]
Classification
- Genus Acelyphus Malloch, 1929[5] - Asia[6]
- Genus Atopocelyphus, Gaimari, 2017 - French Guiana[7]
- Genus Celyphus Dalman, 1818 - Africa and Asia[6]
- Subgenus Celyphus Dalman, 1818 - Asia
- Subgenus Hemiglobus Frey, 1941 - Africa
- Subgenus Paracelyphus Bigot, 1859 - Asia
- Genus Chamaecelyphus Frey, 1941[8] - Africa
- Genus Idiocelyphus Malloch, 1929[5] - Asia
- Genus Oocelyphus Chen, 1949[9] - southern China
- Genus Spaniocelyphus Hendel, 1914[10] - Africa and Asia
Further reading
- Frey . R . 1941 . Die Gattungen und Arten der Dipterenfamilie Celyphidae . Notul. Entomol . 21 . 3–16 . Keys genera, species.
- Vanschuytbroeck . P . 1959 . Celyphidae (Diptera Acalyptratae) . Explor. Parc Natl. Garamba Mission H. De Saeger . 13 . 27–44 . Keys genera, species.
- Tenorio . J.M. . 1972 . A revision of the Celyphidae (Diptera) of the Oriental Region . Trans. R. Entomol. Soc. Lond. . 123 . 4. 359–453 . 10.1111/j.1365-2311.1972.tb00847.x .
- Paap, L. Celyphidae in Soós, Á, Paap, L. (Eds.) 1984 - 1992. Catalogue of Palaearctic Diptera 9 . Akadémiai Kiadó, Budapest - Elsevier, Amsterdam: 63-66..
External links
Notes and References
- Tenorio. Joann M.. 2009-04-24. A revision of the Celyphidae (Diptera) of the Oriental Region*. Transactions of the Royal Entomological Society of London. en. 123. 4. 359–453. 10.1111/j.1365-2311.1972.tb00847.x. 0035-8894.
- Tenorio, JoAnn M.. 1969. A revision of the Celyphidae (Diptera) from the Philippine Islands. Pacific Insects. 11. 3–4. 579–611.
- Li. Xuankun . Li. Wenliang . Ding. Shuangmei . Cameron. Stephen L. . Mao. Meng . Shi. Li . Yang. Ding. 2017-04-14. Mitochondrial Genomes Provide Insights into the Phylogeny of Lauxanioidea (Diptera: Cyclorrhapha) . International Journal of Molecular Sciences . 18 . 4 . 773 . 10.3390/ijms18040773 . 5412357 . 28420076. free .
- http://www.tolweb.org/Lauxanioidea/10537 Lauxanioidea
- Malloch. J.R.. Notes on some Oriental sapromyzid flies (Diptera), with particular reference to the Philippine species. Proceedings of the United States National Museum. 1929. 74. 6. 1–97. 26 August 2017. 10.5479/si.00963801.74-2751.1.
- Tenorio. JoAnn M. . 1969. A Revision of the Celyphidae (DIPTERA) From the Philippine Islands. Pacific Insects. 11. 3–4. 579–611. Hawaii Biological Survey. 6 November 2012.
- Gaimari. Stephen D.. 2017-10-23. The dipteran family Celyphidae in the New World, with discussion of and key to world genera (Insecta, Diptera) . ZooKeys . en. 711. 113–130. 10.3897/zookeys.711.20840. 1313-2970. 5674188. 29134031. free.
- Frey. R.. Die Gattungen und Arten der Dipteren familie Celyphidae. Notul. Entomol.. 1941. 21. 3–16. 1 pl.
- Chen. S. H.. Records of Chinese Diopsidae and Celyphidae (Diptera). Sinensia. 1949. 10. 1–6. 1–6.
- Hendel. F. . H.. Sauter's Formosa-Ausbeute. Acalyptrate Musciden (Dipt.) III. Supplementa Entomologica. 1914. 3. 90–117.