Bromheadia Explained

Bromheadia, commonly known as reed orchids,[1] is a genus of about 29 species of orchids in the family Orchidaceae. They are evergreen terrestrial and epiphytic plants with unbranched stems, the leaves arranged in two rows along the flowering stem. The flowers appear in succession near the end of the flowering stem with the sepals and petals free from each other. The labellum is like a landing platform and has three lobes. They are native to areas from tropical Asia to northern Australia.

Description

Plants in the genus Bromheadia are evergreen, terrestrial or epiphytic, sympodial, herbs with leafy stems on a short, creeping rhizome. The leaves are arranged in two rows along an erect, unbranched stem with the flattened, leathery leaves along the upper two-thirds. The flowers are short-lived and appear singly, in succession near the ends of the flowering stem. The flowers are relatively large, resupinate and usually white, creamy yellow or reddish. The sepals are narrower but longer than petals and are often a different colour. The labellum projects forwards and has three lobes with the side lobes erect and there are two pollinia.[2] [3] [4]

Taxonomy and naming

The genus Bromheadia was first formally described in 1841 by John Lindley and the description was published in Edwards's Botanical Register.[5] The genus name (Bromheadia) honours Edward Bromhead "whose investigations of the natural affinities of plants are well known to systematic Botanists".

Distribution

Orchids in the genus Bromheadia are found in the region stretching from Myanmar and Vietnam south to Java and east to New Guinea and the Philippines, with a few species extending to Sri Lanka and Queensland.

Species

The genus is subdivided into two sections identified by the clear morphological difference in the shape of the leaves: section Bromheadia has dorso-ventrally flattened leaves whereas section Aporodes has laterally-flattened leaves.[6] While these two sections represent natural groups with clear morphological support, a 1997 phylogenetic study of the relationships within Bromheadia that used only morphological characters and placed Claderia as the outgroup concluded that while section Aporodes was monophyletic, section Bromheadia was paraphyletic. To resolve the conflict, the study's authors suggested that section Bromheadia either absorb section Aporodes to make one large monophyletic genus with no infrageneric subdivisions or to divide section Bromheadia into the three monophyletic groups they identified within it so that the genus would have four sections.[7] However, no further taxonomic work has acted on this suggestion so the genus remains with two natural sections:

Section Bromheadia
Section Aporodes
Insufficiently known

Notes and References

  1. Book: Jones. David L.. A complete guide to native orchids of Australia including the island territories. 2006. New Holland. Frenchs Forest, N.S.W.. 1877069124. 352.
  2. Web site: Genus Bromheadia . Orchids of New Guinea . 7 September 2018.
  3. Ridley . Henry N. . The genus Bromheadia . Journal of the Linnean Society, Botany . 1891 . 28 . 195 . 331 . 10.1111/j.1095-8339.1891.tb01464.x . 7 September 2018.
  4. Book: Pridgeon . Alec M. (ed.) . Cribb . Phillip J. (ed.) . Cahase . Mark W. (ed.) . Rasmussen . Finn N. (ed.) . Genera orchidacearum (volume 6) . 2014 . Oxford University Press . Oxford . 9780199646517 . 106–109 . 1st.
  5. Book: Lindley . John . Edwards's Botanical Register (Volume 4) . 1841 . James Ridgway and sons . London . 89–90 . 7 September 2018.
  6. Kruizinga, J., H.J. van Scheindelen, and E.F. de Vogel. 1997. Revision of the genus Bromheadia (Orchidaceae). Orchid Monographs, 8: 79-118, figures 29-55, plates 4b-5b.
  7. Repetur, C.P., P.C. van Welzen, and E.F. de Vogel. 1997. Phylogeny and historical biogeography of the genus Bromheadia (Orchidaceae). Systematic Botany, 22(3): 465-477.