Barbodes Explained

Barbodes is a genus of small to medium-sized cyprinid fish native to tropical Asia. The majority of the species are from Southeast Asia. Many species are threatened and some from the Philippines (Lake Lanao) are already extinct. A survey carried out in 1992 only found three of the endemic Barbodes species,^[1] and only two (Barbodes lindog and B. tumba) were found in 2008.^[2] Several members of this genus were formerly included in Puntius.^[3]

Etymology

The name is derived from the Latin word barbus, meaning "barbel", and the Greek word oides, meaning "similar to".

Species

There are currently 48 recognized species in this genus, of which 15 are considered extinct and 2 considered possibly extinct:^[3]

• †Barbodes amarus Herre, 1924 (Pait)
• Barbodes aurotaeniatus (Tirant, 1885)
• Barbodes banksi (Herre, 1940)
• †Barbodes baoulan Herre, 1926 (Baolan)
• Barbodes binotatus (Valenciennes, 1842) (Spotted barb)
• Barbodes bunau (Rachmatika, 2005)
• Barbodes cataractae (Fowler, 1934)
• †Barbodes clemensi Herre, 1924
• Barbodes colemani (Fowler, 1937)
• †Barbodes disa Herre, 1932
• Barbodes dorsimaculatus (C. G. E. Ahl, 1923) (Blackline barb)
• Barbodes dunckeri (C. G. E. Ahl, 1929) (Bigspot barb)
• Barbodes everetti (Boulenger, 1894) (Clown barb)
• †Barbodes flavifuscus Herre, 1924 (Katapa-tapa)
• Barbodes hemictenus D. S. Jordan & R. E. Richardson, 1908
• †Barbodes herrei (Fowler, 1934)
• Barbodes ivis (Seale, 1910)
• Barbodes joaquinae (C. E. Wood, 1968)
• †Barbodes katolo Herre, 1924
• Barbodes kuchingensis (Herre, 1940)
• †Barbodes lanaoensis Herre, 1924 (Kandar)
• Barbodes lateristriga (Valenciennes, 1842) (Spanner barb)
• Barbodes lindog Herre, 1924 (Lindog) (possibly extinct)
• †Barbodes manalak Herre, 1924 (Manalak)
• Barbodes manguaoensis (A. L. Day, 1914)
• Barbodes microps (Günther, 1868)
• Barbodes montanoi (Sauvage, 1881)
• †Barbodes pachycheilus (Herre, 1924)
• †Barbodes palaemophagus (Herre, 1924)
• †Barbodes palata Herre, 1924
• Barbodes palavanensis (Boulenger, 1895)
• Barbodes polylepis J. X. Chen & D. J. Li, 1988
• Barbodes quinquemaculatus (Seale & B. A. Bean, 1907)
• †Barbodes resimus (Herre, 1924)
• Barbodes rhombeus (Kottelat, 2000)
• Barbodes sellifer (Kottelat & K. K. P. Lim, 2021)^[4]
• Barbodes semifasciolatus (Günther, 1868) (Chinese barb)
  • Barbodes semifasciolatus sachsii (Ahl, 1923) (Goldfinned barb)
• Barbodes sirang Herre, 1932 (Sirang) (possibly extinct)
• Barbodes snyderi Ōshima, 1919
• †Barbodes tras Herre, 1926
• †Barbodes truncatulus (Herre, 1924) (Bitungu)
• Barbodes tumba Herre, 1924
• Barbodes umalii (C. E. Wood, 1968)
• Barbodes wynaadensis (F. Day, 1873)
• Barbodes xouthos (Kottelat & H. H. Tan, 2011)^[5]
• Barbodes zakariaismaili (Kottelat & K. K. P. Lim, 2021)^[4]

Note on species list: Kottelat 2013 states that B. dorsimaculatus may not be referable to this genus and considers it to be species inquirenda. He also implies that Barbodes should be restricted to Southeast Asian and Philippine endemics and that the following species from eastern and southern Asia may not be referable to this genus: B. bovanicus, B. carnaticus, B. elongatus, B. polylepis and B. wynaadensis. Because these species fall outside of the geographic area of his paper, their position in Cyprinidae is not addressed.^[3]

Notes and References

1. Endangered Species Handbook: It's Too Late – Fish Extinctions. Retrieved 29 September 2012
2. Ismail . Gladys B. . Sampson . David B. . Noakes . David L. G. . The status of Lake Lanao endemic cyprinids (Puntius species) and their conservation . Environmental Biology of Fishes . 2013 . 97 . 4 . 425–434 . 10.1007/s10641-013-0163-1.
3. Kottelat . M. . The fishes of the inland waters of southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries . Raffles Bulletin of Zoology . Supplement . 2013 . 27 . 1–663 . July 8, 2021 . December 2, 2013 . https://web.archive.org/web/20131202225925/http://rmbr.nus.edu.sg/rbz/biblio/s27/rbz_S27.pdf . dead .
4. Kottelat, M. . Lim, K. K. P. . 2021 . Two new species of Barbodes from the Malay Peninsula and comments on ‘cryptic species’ in the B. binotatus group (Teleostei: Cyprinidae). . The Raffles Bulletin of Zoology . 69 . 522–540 . 10.26107/RBZ-2021-0069.
5. Kottelat . M. . Tan . H. H. . Systomus xouthos, a new cyprinid fish from borneo, and revalidation of Puntius pulcher (Teleostei: Cyprinidae) . Ichthyological Exploration of Freshwaters . 2011 . 22 . 3 . 209-214 . 0936-9902.