Armadillidium atticum explained

Armadillidium atticum is a European species of woodlouse endemic to Greece.[1] [2] It is a relatively small species and belongs to the so called "Armadillidium insulanum complex".[3]

Taxonomy

Armadilliidum atticum was originally described as a distinct species of Armadillidium based on specimens collected from the surroundings of Athens.[1] It belongs to the "Armadillidiae" section of the genus, which is characterized by rib-like side edges of the forehead that never form a "frontal brim" along with the frontal shield, a frontal triangle with lateral edges that never reach the areas of the ocelli, relatively steeply-dropping epimera on all tergites and anterior lobes of the first pair of epimera that are usually either not curved upwards or with a slight hint of a bend.[4] Additionally, the species is part of the "Armadillidium insulanum complex", a group of closely related, small sized pill bugs that occur in various Aegean Islands and the nearby Greek and Turkish coasts. Specifically, within this group, Armadillidium atticum is most closely related to Armadillidium insulanum sensu lato, from which it can safely been separated only by the more elongated first pleopod-exopodite of the male.[3]

Historically, three subspecies of Armadillidium atticum have been described. However, their validity is disputed with various sources accepting none, some or all of them.[3] [5] The subspecies, and their diagnostic characters, are the following:[4]

Description

Armadillidium atticum is a small to medium sized species, reaching maximum dimensions of only about 13.5 x 5.8 mm. Its tergites are covered with granulations and the coloration of the body consists of a pale greyish-brown hue with lighter epimera. The frontal shield surpasses the anterior edge of the head, has a straight to curved upper margin and its lateral corners are pronounced. The head itself has a groove in the middle, immediately behind the frontal shield. The lobes of the secondary antennae are trapezoidal and the distal article of the flagellum is slightly longer than the first. The hind margins of the first pair of epimera are rounded. The telson is triangular, as wide as long and has straight sides and a rounded tip.[1] [3] [4]

Concerning the sexual characters of the male, the first pereiopod has a weakly developed brush of short spines on the carpus and the seventh pereiopod a ventrally concave ischium, decorated with a longitudinal hair-field at the apical part. The first pleopod has an exopodite with a narrow, elongated posterior lobe and an endopodite with straight apex.[1] [3] [4]

Distribution

Armadillidium atticum is endemic to Central Greece and Cyclades. In particular, until now, the species has been found in the region of Attica on the mainland and the islands of Euboea, Salamis, Aegina and Kythnos in the Aegean Sea.[2] [3] [4]

Ecology

Armadillidium atticum is a relatively xerophilic species which can be found in various typical Mediterranean habitats, like maquis, phrygana and their intermediate morphs. Additionally, it is synanthropic and can frequently be found in croplands and in or around human settlements.[6]

Notes and References

  1. Die Landisopoden des Balkans. 3. Beitrag: Südbalkan. . Hans Strouhal . 1929. Zeitschrift für wissenschaftliche Zoologie . 133. 64 pp.
  2. World catalog of terrestrial isopods (Isopoda: Oniscidea) — revised and updated version . Helmut Schmalfuss . 2003 . Stuttgarter Beiträge zur Naturkunde, Serie A . 654 . 341 pp.
  3. The terrestrial isopods (Isopoda: Oniscidea) of Greece. 24th contribution: The genus Armadillidium (Armadillidiidae) on the Aegean islands . Helmut Schmalfuss . 2006 . Stuttgarter Beiträge zur Naturkunde, Serie A . 698 . 43 pp.
  4. Ordnung Isopoda (Landasseln).. Karl Schmölzer. 1965. Bestimmungsbücher zur Bodenfauna Europas. . 4-5. 468 pp.
  5. Armadillidium atticum. 257642.
  6. A study of terrestrial arthropod fauna among different habitat types on Mt. Parnitha (in Greek). Eva Pitta. 2009 . Bachelor's Thesis. 117 pp.