Archisargidae Explained

Archisargidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It is part of the extinct superfamily Archisargoidea. Most members of the family are known from the Callovian-Oxfordian Daohugou biota of Inner Mongolia, China, and the equivalently aged Karabastau Formation of Kazakhstan. The family has been found to be paraphyletic with respect to Eremochaetidae in a cladistic analysis.^[1]

Taxonomy

• †subfamily Archisarginae Rohdendorf 1962
  • †Archirhagio Rohdendorf 1938^[2]
    • †Archirhagio gracilentus Wang et al. 2017^[3] Daohugou, China, Callovian
    • †Archirhagio obscurus Rohdendorf 1938 Karabastau Formation, Kazakhstan, Oxfordian
    • †Archirhagio striatus Zhang and Zhang 2003^[4] Daohugou, China, Callovian
    • †Archirhagio varius Zhang 2014^[5] Daohugou, China, Callovian
    • †Archirhagio zhangi Zhang et al. 2009 Tiaojishan Formation, China, Callovian
  • †Archisargus Rohdendorf 1938
    • †Archisargus maximus Mostovski 1997^[6] Karabastau Formation, Kazakhstan, Oxfordian
    • †Archisargus pulcher Rohdendorf 1938 Karabastau Formation, Kazakhstan, Oxfordian
    • †Archisargus spurivenius Zhang et al. 2007a^[7] Daohugou, China, Callovian
    • †Archisargus strigatus Zhang et al. 2007a Daohugou, China, Callovian
  • †Calosargus Mostovski 1997
    • †Calosargus (Calosargus) Mostovski 1997
      • †Calosargus (Calosargus) antiquus Zhang et al. 2007b^[8] Daohugou, China, Callovian
      • †Calosargus (Calosargus) bellus Zhang et al. 2007b Daohugou, China, Callovian
      • †Calosargus (Calosargus) daohugouensis Zhang et al. 2007b Daohugou, China, Callovian
      • †Calosargus (Calosargus) hani Zhang et al. 2007b Daohugou, China, Callovian
      • †Calosargus (Calosargus) niger Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian
      • †Calosargus (Calosargus) talbragarensis Oberprieler and Yeates 2012^[9] Talbragar Fossil Bed, Australia, Tithonian
      • †Calosargus (Calosargus) tatianae Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian
      • †Calosargus (Calosargus) tenuicellulatus Zhang et al. 2007b Daohugou, China, Callovian
      • †Calosargus (Calosargus) validus Zhang et al. 2007b Daohugou, China, Callovian
    • †Calosargus (Pterosargus) Mostovski 1997
      • †Calosargus (Pterosargus) sinicus Zhang 2010^[10] Daohugou, China, Callovian
      • †Calosargus (Pterosargus) thanasymus Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian
    • †Calosargus tatianae Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian
  • †Flagellisargus Zhang 2012^[11]
    • †Flagellisargus robustus Zhang 2012 Daohugou, China, Callovian
    • †Flagellisargus sinicus Zhang 2012 Daohugou, China, Callovian
    • †Flagellisargus venustus Zhang 2012 Daohugou, China, Callovian
  • †Mesosolva Hong 1983^[12]
    • †Mesosolva angustocellulata Mostovski 1996^[13] Karabastau Formation, Kazakhstan, Oxfordian
    • †Mesosolva balyshevae Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Mesosolva daohugouensis Zhang and Zhang 2003 Daohugou, China, Callovian
    • †Mesosolva dolosa Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Mesosolva hennigi Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Mesosolva huabeiensis (Hong 1983) Haifanggou Formation, China, Callovian
    • †Mesosolva imperfecta Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Mesosolva karataviensis (Mostovski 1996) Karabastau Formation, Kazakhstan, Oxfordian
    • †Mesosolva longivena Mostovski 1996 Shar Teeg, Mongolia, Tithonian
    • †Mesosolva parva Hong 1983 Haifanggou Formation, China, Callovian
    • †Mesosolva rohdendorfi Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Mesosolva sinensis Zhang et al. 2010^[14] Daohugou, China, Callovian
    • †Mesosolva zhangae Zhang 2012^[15] Daohugou, China, Callovian
  • †Novisargus Zhang 2014
    • †Novisargus rarus Zhang 2014 Daohugou, China, Callovian
  • †Origoasilus Zhang et al. 2011^[16]
    • †Origoasilus pingquanensis Zhang et al. 2011 Yixian Formation, China, Aptian
  • †Parvisargus Mostovski 1996^[17]
    • †Parvisargus malus Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Parvisargus peior Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
  • †Sharasargus Mostovski 1996
    • †Sharasargus fortis Zhang et al. 2008^[18] Daohugou, China, Callovian
    • †Sharasargus maculus Zhang 2014 Daohugou, China, Callovian
    • †Sharasargus oresbius (Ren 1998)^[19] Yixian Formation, China, Aptian
    • †Sharasargus ruptus Mostovski 1996 Shar Teeg, Mongolia, Tithonian
    • †Sharasargus spiniger Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
  • †Sinallomyia Zhang 2012
    • †Sinallomyia ruderalis (Ren 1998) Yixian Formation, China, Aptian
  • †Tabanisargus Zhang 2014
    • †Tabanisargus daohugouensis Zhang 2014 Daohugou, China, Callovian
• †subfamily Uranorhagioninae Zhang et al. 2010^[20] Grimaldi 2016 suggests that this grouping does not belong to the superfamily.
  • †Daohugosargus Zhang 2012
    • †Daohugosargus eximius (Zhang et al. 2008) Daohugou, China, Callovian
  • †Uranorhagio Zhang et al. 2010
    • †Uranorhagio asymmetricus (Zhang et al. 2010) Daohugou, China, Callovian
    • †Uranorhagio daohugouensis Zhang et al. 2010 Daohugou, China, Callovian
    • †Uranorhagio deviatus (Zhang et al. 2010) Daohugou, China, Callovian

Notes and References

1. Grimaldi. David A.. Barden. Phillip. 2016-09-29. The Mesozoic Family Eremochaetidae (Diptera: Brachycera) in Burmese Amber and Relationships of Archisargoidea: Brachycera in Cretaceous Amber, Part VIII. American Museum Novitates. en. 3865. 1–29. 10.1206/3865.1. 89602433. 0003-0082.
2. B. B. Rohdendorf. 1938. Dvukrylye basekomye Mezozoya Kara-Tau. I. Brachycera i chast' Nematocera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7:29-67
3. Wang. Fengyan. Shih. Chungkun. Ren. Dong. Wang. Yongjie. January 2017. Quantitative assessments and taxonomic revision of the genus Archirhagio with a new species from Daohugou, China (Diptera: Archisargidae). Systematic Entomology. en. 42. 1. 230–239. 10.1111/syen.12204. 90656127. free.
4. J. Zhang and H. Zhang. 2003. Two new species of archisargids (Insecta: Diptera: Archisargidae) from the Upper Jurassic Daohugou Formation (Inner Mongolia, northeastern China). Paleontological Journal 37:409-412
5. Zhang. Junfeng. 2015-10-03. Archisargoid flies (Diptera, Brachycera, Archisargidae and Kovalevisargidae) from the Jurassic Daohugou biota of China, and the related biostratigraphical correlation and geological age. Journal of Systematic Palaeontology. en. 13. 10. 857–881. 10.1080/14772019.2014.960902. 84512383. 1477-2019.
6. M. B. Mostovski. 1997. On knowledge of fossil flies of the superfamily Archisargoidea (Diptera, Brachycera). Paleontological Journal 31(1):72-78
7. K. Y. Zhang, D. Yang, D. Ren and C. K. Shih. 2007. The earliest species of the extinct genus Archisargus from China (Diptera: Brachycera: Archisargidae). Annales Zoologici 57:827-832
8. Zhang. Kuiyan. Yang. Ding. Ren. Dong. Shih. Chungkun. 2007-11-23. The oldest Calosargus Mostovski, 1997 from the Middle Jurassic of China (Diptera: Brachycera: Archisargidae). Zootaxa. 1645. 1. 1–17. 10.11646/zootaxa.1645.1.1. 1175-5334.
9. Oberprieler. Stefanie K.. Yeates. David K.. July 2012. Calosargus talbragarensis new species: the first brachyceran fly from the Jurassic of Australia (Diptera, Archisargidae). Journal of Paleontology. en. 86. 4. 641–645. 10.1666/11-126R.1. 128737218. 0022-3360.
10. Zhang. Jun Feng. March 2010. Records of bizarre Jurassic brachycerans in the Daohugou biota, China (Diptera, Brachycera, Archisargidae and Rhagionemestriidae). Palaeontology. en. 53. 2. 307–317. 10.1111/j.1475-4983.2010.00934.x. free.
11. Zhang. Junfeng. September 2012. Distinct but rare archisargid flies from Jurassic of China (Diptera, Brachycera, Archisargidae) with discussion of the systematic position of Origoasilus pingquanensis Zhang et al., 2011. Journal of Paleontology. 86. 5. 878–885. 10.1666/12-028.1. 86217269. 0022-3360.
12. Y. C. Hong. 1983. Middle Jurassic Fossil Insects in North China 1-223
13. M. B. Mostovski. 1996. New Species of the Genus Mesosolva Hong, 1983 (Diptera, Archisargidae) from the Jurassic of Kazakhstan and Mongolia. The Continental Jurassic 60:329-332
14. Zhang. Kuiyan. Yang. Ding. Ren. Dong. Shih. Chungkun. March 2010. New archisargids from China (Insecta: Diptera). Entomological Science. en. 13. 1. 75–80. 10.1111/j.1479-8298.2009.00350.x. 84716263. free.
15. Junfeng. Zhang. 2012-11-06. Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China. ZooKeys. 238. 57–76. 10.3897/zookeys.238.3624. 1313-2970. 3496945. 23226708. free .
16. K. Y. Zhang, D. Yang, and D. Ren. 2011. A new brachyceran family Origoasilidae Fam. Nov. from the late Mesozoic of China (Insecta: Diptera). Acta Geologica Sinica 85(5):994-997
17. M. B. Mostovski. 1996. To the knowledge of Archisargoidea (Diptera, Brachycera). Families Eremochaetidae and Archisargidae. Russian Entomological Journal 5:117-124
18. Zhang. Kuiyan. Yang. Ding. Ren. Dong. June 2008. Middle Jurassic fossils of the genus Sharasargus from Inner Mongolia, China (Diptera: Archisargidae). Entomological Science. en. 11. 2. 269–272. 10.1111/j.1479-8298.2008.00271.x. 84139157.
19. D. Ren. 1998. Late Jurassic Brachycera from northeastern China (Insecta: Diptera). Acta Zootaxonomica Sinica 23:65-82
20. Zhang. Kuiyan. Yang. Ding. Ren. Dong. Shih. Chungkun. March 2010. An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera): FOSSIL FLIES FROM THE MIDDLE JURASSIC OF CHINA. Zoological Journal of the Linnean Society. en. 158. 3. 563–572. 10.1111/j.1096-3642.2009.00552.x.