Arbutoideae Explained

The Arbutoideae are a subfamily in the plant family Ericaceae. Phylogenetic analysis supported all genera of the subfamily as monophyletic, except Arbutus. Moreover, it was suggested that the non-sister relationship between Mediterranean and North American species may be explained by a once widespread distribution in the Northern hemisphere before the Neogene.

The genera Arbutus, Arctostaphylos, Comarostaphylis form a particular type of mycorrhizal symbiosis with the fungus, Arbutoid mycorrhiza, which resembles ectomycorrhizas.

Genera List

Image Genus Living species
Arbutus
SpeciesCommon name Range
andrachneGreek strawberry treeMediterranean and Middle East
canariensisCanary madroneCanary Islands of Spain
pavariiLibya: Jebel Akhdar range in coastal Cyrenaica
unedoStrawberry treeEurope, north Africa, Middle East
arizonicaArizona madroneMexico, southwestern United States
bicolor.
madrensisMexico: Durango, Jalisco, Nayarit, Sinaloa
menziesiiPacific madroneCoastal areas of British Columbia, Washington, Oregon, California; Sierra Nevada
mollisMexico
occidentaliswestern Mexico
tessellata.northern and central Mexico from Chihuahua to Jalisco, Tlaxcala, and Veracruz
xalapensisTexas madroneCentral American, Mexico, southwestern United States
Arctostaphylos
  • Subgenus Micrococcus
    • Sect. Micrococcus
SpeciesCommon name Range
mendocinoensisPygmy manzanita
myrtifoliaIone manzanitaCalifornia (Amador, Calaveras Counties)
nissenanaNissenan manzanitaCalifornia (coastal and inland ranges north of San Francisco Bay)
nummulariaGlossyleaf manzanitaCalifornia (Mendocino County)
  • Subgenus Arctostaphylos, which has 3 sections:
    • Sect. Arctostaphylos
SpeciesCommon name Range
alpinaAlpine bearberry
bakeriBaker's manzanitaCalifornia (Sonoma County)
densifloraSonoma manzanitaCalifornia (Sonoma County)
edmundsiiLittle Sur manzanitaCalifornia (Monterey County)
franciscanaFranciscan manzanitaCalifornia (San Francisco County)
gabrielensisSan Gabriel manzanitaCalifornia (Los Angeles County)
glaucaBigberry manzanitaCalifornia and Baja California
hispidulaGasquet manzanitaCoastal mountain ranges of southern Oregon and northern California
hookeriHooker's manzanitaCalifornia
insularisIsland manzanitaCalifornia (Santa Cruz Island)
klamathensisKlamath manzanitaCalifornia (Klamath Mountains)
manzanitaCommon manzanita, whiteleaf manzanitaCalifornia (Coast Ranges and Sierra Nevada foothills)
mewukkaIndian manzanitaCalifornia (Sierra Nevada)
nevadensisPinemat manzanitaCalifornia
parryanaParry manzanitaCalifornia (southern)
patulaGreenleaf manzanitaWestern North America
pumilaSandmat manzanitaCalifornia (Monterey County)
pungensPointleaf manzanitaSouthwestern United States and to northern and central Mexico
rudisShagbark manzanitaCalifornia (southern central coast)
stanfordianaStanford's manzanitaCalifornia (Outer North Coast Ranges north of the San Francisco Bay Area)
uva-ursiBearberryEurope, Asia, North America
viscidaSticky manzanita, whiteleaf manzanitaCalifornia and Oregon
    • Sect. Foliobracteata
SpeciesCommon name Range
andersoniiSanta Cruz manzanitaSanta Cruz mountains (CA)
auriculataMount Diablo manzanitaMount Diablo (CA)
canescensHoary manzanitaCoastal ranges of SW OR and N CA
catalinaeSanta Catalina Island manzanitaSanta Catalina Island (CA)
columbianaHairy manzanitaWest coast from N. CA to S. BC
confertifloraSanta Rosa Island manzanitaSanta Rosa Island (CA)
cruzensisLa Cruz manzanitaMonterey and San Luis Obispo counties (CA)
glandulosaEastwood manzanitaCoastal slops from OR, CA, Baja California
glutinosaSchreiber's manzanitaSanta Cruz County (CA)
hooveriHoover's manzanitaSanta Lucia Mountains (CA)
imbricataSan Bruno Mountain manzanitaSan Bruno Mountain (CA)
lucianaSanta Lucia manzanitasouthern Santa Lucia Mountains (CA)
malloryiMallory's manzanitaInner North Coast Ranges west and northwest of the Sacramento Valley (CA)
montaraensisMontara manzanitaSan Bruno Mountain and Montara Mountain (CA)
montereyensisMonterey manzanitaMonterey County (CA)
morroensisMorro manzanitaMorro Bay (CA)
nortensisDel Norte manzanitaDel Norte County (CA), Curry County (OR), Josephine County (OR)
obispoensisSerpentine manzanitaSouthern Santa Lucia Mountains (CA)
osoensisOso manzanitaLos Osos Valley (San Luis Obispo, CA)
otayensisOtay manzanitaSan Diego County (CA)
pajaroensisPajaro manzanitaMonterey County, Santa Cruz County, San Benito County (CA)
pallidaPallid manzanita
pechoensisPecho manzanita
pilosulaLa Panzo manzanita
purissimaLa Purissima manzanita
refugioensisRefugio manzanita
regismontanaKings Mountain manzanita
silvicolaBonny Doon manzanita
tomentosaWoolyleaf manzanita
virgataBolinas manzanita
viridissimaWhitehair manzanita
wellsiiWells' manzanita
    • Sect. Pictobracteata
  • Unassigned
SpeciesCommon name Range
rainbowensisRainbow manzanita
gabilanensisGabilan manzanita
ohloneanaOhlone manzanita
Arctous
SpeciesCommon name Range
alpina (L.) Nied.
microphyllus C.Y.Wu
ruber (Rehder & E.H.Wilson) Nakai
Comarostaphylis
SpeciesCommon name Range
arbutoides
discolor
diversifolia
glaucescens
lanata
longifolia
mucronata
polifolia
sharpii
spinulosa
Ornithostaphylos
Xylococcus

External links