Arachniodes Explained
Arachniodes is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".[1]
The genus Arachniodes was first published by Carl Ludwig von Blume in 1828, with the single Indonesian species Arachniodes aspidioides.[2] The genus was not widely recognized until Mary Douglas Tindale transferred the two species (Byrsopteris amabilis and Byrsopteris aristata) into it in 1961.[3]
Species
The Pteridophyte Phylogeny Group classification of 2016 (PPG I) considers Arachniodes to have 60 species., the Checklist of Ferns and Lycophytes of the World accepted 78 species:
- Arachniodes ailaoshanensis Ching
- Arachniodes amabilis (Blume) Tindale
- Arachniodes amoena (Ching) Ching
- Arachniodes aristata (G.Forst.) Tindale
- Arachniodes assamica (Kuhn) Ohwi
- Arachniodes bella (C.Chr.) Ching
- Arachniodes blinii (H.Lév.) Nakaike
- Arachniodes carvifolia (Kunze) Ching
- Arachniodes caudata Ching
- Arachniodes cavaleriei (Christ) Ohwi
- Arachniodes chinensis (Rosenst.) Ching
- Arachniodes coniifolia (T.Moore) Ching
- Arachniodes cornucervi (D.Don) Fraser-Jenk.
- Arachniodes daklakensis Li Bing Zhang, N.T.Lu & X.M.Zhou
- Arachniodes davalliaeformis (Christ) Nakaike
- Arachniodes denticulata (Sw.) Ching
- Arachniodes fengii Ching
- Arachniodes festina (Hance) Ching
- Arachniodes formosa (Fée) Ching
- Arachniodes formosissima (Goldm.) Nakaike
- Arachniodes gigantea Ching
- Arachniodes globisora (Hayata) Ching
- Arachniodes grossa (Tardieu & C.Chr.) Ching
- Arachniodes hainanensis (Ching) Ching
- Arachniodes haniffii (Holttum) Ching
- Arachniodes hehaii Li Bing Zhang, N.T.Lu & X.F.Gao
- Arachniodes hekiana Kurata
- Arachniodes henryi (Christ) Ching
- Arachniodes hiugana Sa. Kurata
- Arachniodes hunanensis Ching
- Arachniodes insularis W.H.Wagner
- Arachniodes japonica (Kurata) Nakaike
- Arachniodes jinpingensis Y.T.Hsieh
- Arachniodes leucostegioides (C.Chr.) Ching
- Arachniodes longicaudata Li Bing Zhang, N.T.Lu & Liang Zhang
- Arachniodes longipinna Ching
- Arachniodes microlepioides (C.Chr.) comb. ined.
- Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi
- Arachniodes miyakei (H.Itô) Shimura ex Nakaike
- Arachniodes mutica (Franch. & Sav.) Ohwi
- Arachniodes neopodophylla (Ching) Nakaike
- Arachniodes nigrospinosa (Ching) Ching
- Arachniodes nipponica (Rosenst.) Ohwi
- Arachniodes oosorae H.Itô
- Arachniodes palmipes (Kunze) Fraser-Jenk.
- Arachniodes pseudoaristata (Tagawa) Ohwi
- Arachniodes pseudoassamica Ching
- Arachniodes pseudorepens Nakaike
- Arachniodes puncticulata (Alderw.) Ching
- Arachniodes quadripinnata (Hayata) Seriz.
- Arachniodes quangnamensis Li Bing Zhang, N.T.Lu & Liang Zhang
- Arachniodes repens Kurata
- Arachniodes rhomboidea (Schott) Ching
- Arachniodes rigidissima (Hook.) Ching
- Arachniodes sarasiniorum (Christ) Nakaike
- Arachniodes simplicior (Makino) Ohwi
- Arachniodes simulans (Ching) Ching
- Arachniodes sinomiqueliana (Ching) Ohwi
- Arachniodes sinorhomboidea Ching
- Arachniodes sledgei Fraser-Jenk.
- Arachniodes spectabilis (Ching) Ching
- Arachniodes squamulosa R.C.Moran & B. Øllg.
- Arachniodes standishii (T.Moore) Ohwi
- Arachniodes subamabilis Kurata
- Arachniodes superba Fraser-Jenk.
- Arachniodes tomitae Sa. Kurata
- Arachniodes tonkinensis (Ching) Ching
- Arachniodes tripinnata (Goldm.) Sledge
- Arachniodes tsiangiana (Ching) Nakaike
- Arachniodes webbiana (A.Braun) Schelpe
- Arachniodes wulingshanensis S.F.Wu
- Arachniodes yakusimensis (H.Itô) Nakaike
- Arachniodes ziyunshanensis Y.T.Xie
Some hybrids were also accepted:
- Arachniodes × azuminoensis Fujiw., Y.Matsuda, Yu. Abe & Otsuka
- Arachniodes × chibaensis Yashiro
- Arachniodes × clivorum Kurata
- Arachniodes × ikeminensis Seriz.
- Arachniodes × kenzo-satakei (Kurata) Kurata
- Arachniodes × kurosawae Shimura & Kurata
- Arachniodes × masakii Kurata
- Arachniodes × minamitanii Sa. Kurata
- Arachniodes × mirabilis Sa. Kurata
- Arachniodes × mitsuyoshiana Sa. Kurata
- Arachniodes × pseudohekiana Kurata
- Arachniodes × respiciens Sa. Kurata
- Arachniodes × sahashii Fraser-Jenk. & Kandel
- Arachniodes × sasamotoi Kurata
- Arachniodes × takayamensis Seriz.
Notes and References
- http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=102399 Arachniodes
- Blume, Carl (Karl) Ludwig von; Enumeratio Plantarum Javae fasc. 2: 241. 1828.
- Tindale, Mary Douglas; Contributions from the New South Wales National Herbarium 3(1): 90. 1961.
