Cheirodes Explained

Cheirodes is a genus of darkling beetles in the Melanimonini tribe. It was formerly known as Anemia until 1973, when T. J. Spilman determined Anemia to be a synonym of Cheirodes Géné, 1839.^[1]

Subgenera

Cheirodes is split into several subgenera:

Cheirodes Géné, 1839
Anemiadena Ardoin, 1971^[2]
Histiaea Fairmaire, 1892
Pseudanemia Wollaston, 1864
Spinanemia Ardoin, 1971^[2]
Trichanemia Ardoin, 1971^[2]

Ammidanemia Reitter, 1904 is partly synonymous with Histiaea Fairmaire, 1892 and Pseudanemia Wollaston, 1864.^[2]

Species

Among the species within this genus are:

Subgenus Cheirodes Géné, 1839
- C. aethiopea (Ardoin, 1971)^[2] (Synonym: Anemia aethiopea Ardoin, 1971)
- C. chobauti (Reitter, 1898) (Synonyms: Anemia rotundicollis Reitter, 1884, Anemia reitteri Pic, 1899, Anemia chobauti (Reitter, 1898))
- C. durandi (Ardoin, 1971)^[2] (Synonym: Anemia durandi Ardoin, 1971)
- C. granulatus (Laporte, 1840) (Synonyms: Anemia crassa Wollaston, 1867, Anemia granulata Laporte, 1840)
- C. gruveli (Ardoin, 1971)^[2] (Synonym: Anemia gruveli Ardoin, 1971)
- C. humeralis (Ardoin, 1971)^[2] (Synonym: Anemia humeralis Ardoin, 1971)
- C. opacula (Fairmaire, 1882)
- C. punctata (Ardoin, 1971)^[2] (Synonym: Anemia punctata Ardoin, 1971)
- C. sardous Géné, 1839 (Synonym: Anemia sardoa (Géné, 1839))
  - C. sardoa denticulata (Wollaston, 1867) (Synonym: Anemia sardoa sudanica Gridelli, 1950)
- C. villiersi (Ardoin, 1971)^[2] (Synonym: Anemia villiersi Ardoin, 1971)
Subgenus Spinanemia Ardoin, 1971
- C. allardi (Ardoin, 1971)^[2] (Synonym: Anemia allardi Ardoin, 1971)
- C. amieti (Ardoin, 1971)^[2] (Synonym: Anemia amieti Ardoin, 1971)
- C. capensis (Ardoin, 1971)^[2] (Synonym: Anemia capensis Ardoin, 1971)
- C. cornutus (Pic, 1898) (Synonym: Anemia cornuta Pic, 1898)
  - C. cornutus panelii (Ardoin, 1971)^[2] (Synonym: Anemia cornutus panelii Ardoin, 1971)
- C. crispa (Quedenfeldt, 1885) (Synonym: Anemia crispa Quedenfeldt, 1885)
- C. curta (Ardoin, 1971)^[2] (Synonym: Anemia curta Ardoin, 1971)
- C. hottentota (Ardoin, 1971)^[2] (Synonym: Anemia hottentota Ardoin, 1971)
- C. kaszabi (Ardoin, 1971)^[2] (Synonym: Anemia kaszabi Ardoin, 1971)
- C. mystacina (Ardoin, 1971)^[2] (Synonym: Anemia mystacina Ardoin, 1971)
- C. ovatula (Fairmaire, 1891)
- C. pilosus (Tournier, 1868) (Synonyms: Anemia pilosa Tournier, 1868, Anemia fenyesi Reitter, 1897, Cheirodes fissidens Reitter, 1898)
- C. poteli Lillig & Ferrer, 2001^[3]
- C. roeri (Ardoin, 1976)^[4]
- C. royi (Ardoin, 1971)^[2] (Synonym: Anemia royi Ardoin, 1971)
  - C. royi tchadensis (Ardoin, 1971)^[2] (Synonym: Anemia royi tchadensis Ardoin, 1971)
- C. seriattipennis (Ardoin, 1971)^[2] (Synonym: Anemia seriattipennis Ardoin, 1971)
- C. thoracicus (Fairmaire, 1891) (Synonym: Anemia thoracicus Fairmaire, 1891)
- C. uhligi Ferrer, 2004^[5]
Subgenus Anemiadena Ardoin, 1971
- C. convexa (Gestro, 1881)
- C. giraudini (Ardoin, 1971)^[2] (Synonym: Anemia giraudini Ardoin, 1971)
Subgenus Trichanemia Ardoin, 1971
- C. schmitzi (Ardoin, 1971)^[2] (Synonym: Anemia schmitzi Ardoin, 1971)
Subgenus Histiaea Fairmaire, 1892
- C. angolensis (Ardoin, 1971)^[2] (Synonym: Anemia angolensis Ardoin, 1971)
- C. asperulus (Reitter, 1884) (Synonyms: Anemia denticulata Pic, 1923, Anemia aegyptiaca Pic, 1936, Anemia asperula Reitter, 1884, Anemia asperula var. seriesetosa Baudi, 1884)
- C. bidentulus (Fairmaire, 1892) (Synonym: Histiaea bidentula Ardoin, 1971)
- C. jaegeri Bremer, 2001
- C. kochii (Ardoin, 1976)^[4]
- C. sakalava (Alluaud, 1900)
- C. schultzei (Geibien, 1910) (Synonym: A. submetallica Peringuey, 1908)
Subgenus Pseudanemia Wollaston, 1864
- C. abyssinica (Ardoin, 1971)^[2] (Synonym: Anemia abyssinica Ardoin, 1971)
- C. brevicollis (Wollaston, 1864) (Synonyms: Anemia aphodioides Walker, 1871, Anemia rotundicollis Desbrochers, 1881, Anemia fausti Solsky, 1881, Anemia hauseri Reitter, 1894, Anemia striolata Fairmaire, 1894, Anemia pharao Reitter, 1897, Anemia curticollis Pic, 1923, Pseudanemia brevicollis Wollaston, 1864)
- C. drurei (Pic, 1923) (Synonym: Anemia drurei Pic, 1923)
- C. submetallicus (Raffray, 1873) (Synonyms: Anemia palaestina Pic, 1899, Anemia submetallica (Raffray, 1873), Anoemia submetallica Raffray, 1873)
C. africanus Ferrer, 2016^[6]
C. ahomanae Ferrer, 2016^[6]
C. anatolicus Ferrer, 2016^[6]
C. arabicus Ferrer, 2016^[6]
C. californicus (Horn, 1870) (Synonym: Anaemia californica Horn, 1870)
C. dentipes (Ballion, 1878) (Synonym: Microzoum dentipes Ballion, 1878)
C. guanchorum Ferrer, 2016^[6]
C. linnmani Ferrer, 2016^[6]
C. massauensis Ferrer, 2016^[6]
C. occulata (Wollaston, 1860)
C. otomanus Ferrer, 2016^[6]
C. ruthmuellerae Ferrer, 2016^[6]
C. sculpturata (Ritsema, 1875)

External links

Fauna Europaea: Genus Anemia
Book: I.. Löbl. A.. Smetana. 2008. Tenebrionoidea. Catalogue of Palaearctic Coleoptera. 5. Apollo Books. 9789004260900.

Notes and References

T.J.. Spilman. 1973. Nomenclatural problems in six genera of Tenebrionidae (Coleoptera). Proceedings of the Entomological Society of Washington. 75. 39–44.
P.. Ardoin. 1971. CONTRIBUTION A L'ETUDE DES ESPÈCES AFRICAINES ET MALGACHES DU GENRE ANEMIA LAPORTE [COL. TENEBRIONIDAE]]. Contribution of the study of the African and Madagascan species of the genus Anemia LAPORTE [Col. Tenebrionidae]. . Nouvelle Série. 7. 357–422. French.
Lillig. M.. Ferrer. J.. 2001. Eine neue Art der Gattung Cheirodes GENÉ, 1839 aus Zimbabwe (Coleoptera: Tenebrionidae). Acta Coleopterologica. 17. 31–34. German.
P.. Ardoin. 1976. Deux nouvelles espèces africanes du genre Anemia Laporte (Coleoptera: Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae. 22. 1–4. 1–4. French. subscription.
J.. Ferrer. 2004. Tenebrionidae (Coleoptera) de Namibia, avec descriptions de 12 espèces nouvelles. Zoosystematics and Evolution. 80. 2. 181–250. 10.1002/mmnz.20040800204. French.
J.. Ferrer. 2016. Contribución al conocimiento del género Cheirodes Géné 1839, con descripción de nuevas especies de Europa y África (Coleoptera, Tenebrionidae). Contributions to the knowledge of the genus Cheirodes Géné, 1839, with description of new species from Europe and África (Coleoptera, Tenebrionidae). Boletín de la Sociedad Entomológica Aragonesa. 59. 85–101. Spanish.