Acaenitinae Explained

Acaenitinae is a subfamily of the parasitoid wasp family Ichneumonidae. Female Acaenitinae have a large triangular projecting genital plate.

Distribution

It is distributed on all continents except Antarctica, although only one specimen (from the genus Arotes) has ever been discovered in South America.

Taxonomy and phylogeny

The subfamily has traditionally been divided into two tribes (Acaenitini and Coleocentrini) and comprises 28 genera. The validity of tribes remains debated. Wahl and Gauld considered Coleocentrini paraphyletic and favored abandoning a tribal arrangement in 1998. In contrast, Klopfstein et al. and Bennett et al. found Acaenitini to be monophyletic in 2019. However, they also found that Coleocentrus, the type genus of Coleocentrini, was not recovered as sister to Acaenitini.

Genera

Acaenitini Förster, 1869

Acaenitus Latreille, 1809
Arotes Gravenhorst, 1829
Asperpunctatus Wang, 1989
Boloderma Morley, 1913
Dentifemura Sheng and Sun, 2010
Dimorphonyx Seyrig, 1932
Hieroceryx Tosquinet, 1896
Ishigakia Uchida, 1928
Jezarotes Uchida, 1928
Metachorischizus Uchida, 1928
Notaulites Seyrig, 1932
Paracollyria Cameron, 1906
Phaenolobus Förster, 1869
Phalgea Cameron, 1905
Phorotrophus Saussure, 1892
Prosacron Townes, 1971
Siphimedia Cameron, 1902
Spilopteron Townes, 1965
Yamatarotes Uchida, 1929
Yezoceryx Uchida, 1928

Coleocentrini Clément, 1938

Coleocentrus Gravenhorst, 1829
Cumatocinetus Sheng 2002,
Eremocinetus Viktorov, 1964
Hallocinetus Viktorov, 1962
Leptacoenites Strobl, 1902
Mesoclistus Förster, 1869
Procinetus Förster, 1869

Incertae sedis

Combivena Sheng & Sun, 2014