2022 in paleomalacology explained

This list of fossil molluscs described in 2022 is a list of new taxa of fossil molluscs that were described during the year 2022, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2022.

Ammonites

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Acanthohoplites albedocostatus^[1]	Sp. nov	In press	Matsukawa	Early Cretaceous
Acantholytoceras costaevariavilis	Sp. nov	In press	Matsukawa	Early Cretaceous
Aulatornoceras beyrichi^[2]	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation		A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Buedesheimites	Gen. et sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation		A member of the family Tornoceratidae. Genus includes new species B. housei. Announced in 2022; the final article version was published in 2023.
Cadoceras (Catacadoceras) imlayi	Sp. nov	Valid	Kiselev	Middle Jurassic (Callovian)	Chinitna Formation
Cadoceras kniazevi	Sp. nov	Valid	Kiselev	Middle Jurassic (Callovian)
Cadoceras (Paracadoceras) pseudofrearsi	Sp. nov	Valid	Kiselev	Middle Jurassic (Callovian)
Cadomites zelenchukensis^[3]	Sp. nov	Valid	Mitta	Middle Jurassic (Bathonian)
Collignoniceras rhodanicum^[4]	Sp. nov	Valid	Amédro, Baudouin & Delanoy in Amédro et al.	Late Cretaceous (Turonian)
Crassotornoceras tenue	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation		A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Crioceratites tepehuaensis^[5]	Sp. nov	Valid	Esquivel-Macías et al.	Early Cretaceous
Cryptomorphiceras^[6]	Gen. et comb. et 2 sp. nov	Valid	Marchesi, Balini & Jenks	Late Triassic (Carnian)			A member of Trachyceratinae. The type species is C. compressum (Johnston, 1941); genus also includes new species C. crassum, and C. levicostatum. Announced in 2022; the final article version was published in 2023.
Dactylioceras kopiki^[7]	Sp. nov	Valid	Rogov in Rogov & Lutikov	Early Jurassic (Toarcian)		?	A member of the family Dactylioceratidae.
Diplesioceras fischeri^[8]	Sp. nov	Valid	Dietze, Schweigert & Chandler	Middle Jurassic (Bajocian)	Oolithe de Bayeux Formation
Egoianites jauberti^[9]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)			A member of the family Heteroceratidae.
Epitornoceras transmediterraneum^[10]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)			A member of the family Tornoceratidae.
Forresteria (Harleites) seidli^[11]	Sp. nov	Valid	Summesberger et al.	Late Cretaceous (Santonian)	Gosau Group (Streiteck Formation)
Heteroceras arnaudi	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)			A member of the family Heteroceratidae.
Heteroceras munierchalmasi	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)			A member of the family Heteroceratidae.
Heteroceras penagei	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Barremian)			A member of the family Heteroceratidae.
Hildoceras appenninicus^[12]	Sp. nov	Valid	Ridente	Early Jurassic (Toarcian)
Kepplerites (Gowericeras) copernici^[13]	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)
Kepplerites (Gowericeras) halleyi	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)
Kepplerites (Toricellites) besseli	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)
Kutatissites sudai	Sp. nov	In press	Matsukawa	Early Cretaceous
Lentitornoceras	Gen. et sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Red Ironstone Formation		A member of the family Tornoceratidae. The type species is L. materni.
Linguatornoceras acutilobatum	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation		A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Linguatornoceras sandense	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation		A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Lissoceras kubanense^[14]	Sp. nov	Valid	Mitta	Middle Jurassic (Bajocian)
Lissoceras paviai	Sp. nov	Valid	Mitta	Middle Jurassic (Bajocian)
Megatyloceras leteilensis^[15]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Aptian)			A member of the family Douvilleiceratidae.
Nejdia arabica^[16]	Sp. nov	Valid	Gelin	Early Jurassic (Toarcian)	Marrat Formation
Paracrochordiceras watanabei^[17]	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation		A member of the family Acrochordiceratidae.
Paradanubites ozashiense	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation		A member of Danubitoidea belonging to the family Danubitidae.
Parahedenstroemia? tatjanae^[18]	Sp. nov	Valid	Đaković, Krystyn & Sudar	Early Triassic		?	A member of the family Hedenstroemiidae.
Parapopanoceras involutum	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation		A member of Ceratitida belonging to the group Megaphyllitoidea and the family Parapopanoceratidae.
Paucitornoceras	Gen. et comb. nov	Valid	Korn & Bockwinkel	Devonian	Red Ironstone Formation		A member of the family Tornoceratidae. The type species is "Goniatites" paucistriatus d`Archiac & de Verneuil (1842); genus also includes "Tornoceras" rhysum Clarke (1898).
Ponticeras materni^[19]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Red Ironstone Formation		A member of Agoniatitida belonging to the group Gephuroceratina and the family Ponticeratidae.
Procheloniceras yajimai	Sp. nov	In press	Matsukawa	Early Cretaceous
Pseudocrioceras mazierei^[20]	Sp. nov	Valid	Frau & Delanoy	Early Cretaceous (Barremian/Aptian transition)
Pseudosaynella pacifica	Sp. nov	In press	Matsukawa	Early Cretaceous
Psilokhvalynites	Gen. et sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation		A member of Dinaritoidea belonging to the family Khvalynitidae. The type species is P. takaizumii.
Shasticrioceras nakazawai	Sp. nov	In press	Matsukawa	Early Cretaceous
Sigaloceras (Gulielmina) fabricius	Sp. nov	Valid	Mönnig & Dietl	Middle Jurassic (Callovian)
Tornoceras incisum	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation		A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Tornoceras ventrovaricatum	Sp. nov	Valid	Söte & Becker	Devonian (Frasnian)	Büdesheim Formation		A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023.
Witchellia companyi^[21]	Sp. nov	Valid	Sandoval	Middle Jurassic (Bajocian)			A member of the family Sonniniidae.
Witchellia striata	Sp. nov	Valid	Sandoval	Middle Jurassic (Bajocian)			A member of the family Sonniniidae.

Ammonite research

• Mironenko & Naugolnykh (2022) describe a collection of cephalopod jaws from the Permian (Artinskian) Divjinskian Formation (Sverdlovsk Oblast, Russia), interpreted as likely fossil material of goniatitid ammonites belonging to the genus Uraloceras, and infer from the anatomy of the studied jaws that representatives of the genus Uraloceras were likely active predators.^[22]
• A study on the morphological change through ontogeny of beyrichitine ammonites from the Anisian of Nevada is published by Bischof, Schlüter & Lehmann (2022).^[23]
• A study aiming to determine whether the environmental changes around the Triassic–Jurassic boundary influenced development of septal thickness in ammonites, as well aiming to determine the relation between septal thickness and paleolatitudinal occurrences, and whether ontogenetic septal thickness-trajectories reflected paleogeographic origin or phylogenetic relationships of ammonites, is published by Weber et al. (2022).^[24]
• A study on the hydrodynamic properties of shells of Late Triassic and Early Jurassic ammonites is published by Hebdon et al. (2022).^[25]
• A study on the functional morphology of ammonites from the Middle Triassic to the Early Jurassic, reevaluating possible causes of an overall shift of ammonite shell forms during the studied periods, is published by Hebdon et al. (2022), who interpret their findings as indicating that the trade-offs between coasting efficiency, volume accretion per unit surface area accreted and diameter accretion per unit of surface area (three factors proposed as explanations of the shift of shell forms) were identical in the Triassic and in the Jurassic.^[26]
• Jaws of ammonites belonging to the Boreal family Polyptychitidae, with lower jaws closely resembling jaws of craspeditids, cardioceratids and hoplitoids, are described for the first time from the Valanginian of Russia by Mironenko & Mitta (2022).^[27]
• A study on changes of septal spacing between succeeding chambers of the shell of Gaudryceras tenuiliratum during its ontogeny is published by Kawakami, Uchiyama & Wani (2022).^[28]
• A study on the impact of the conch shape on ammonite locomotion, based on data obtained from tests of 3D-printed, biomimetic robots representing the oxycone morphology (based on Sphenodiscus lobatus and S. lenticularis), the serpenticone morphology (based on Dactylioceras commune), the sphaerocone morphology (based on Goniatites crenistria) and the morphospace center, is published by Peterman & Ritterbush (2022).^[29]

Other cephalopods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aphelaeceras azzelmattiense^[30]	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian–Viséan boundary interval)			A member of the family Trigonoceratidae.
Arctoteuthis angusta^[31]	Sp. nov	Valid	Efremenko	Early Cretaceous			A belemnite belonging to the family Cylindroteuthididae.
Binoleniceras^[32]	Gen. et sp. nov	Valid	Afhüppe & Becker	Devonian (Givetian)	Hagen-Balve Formation		A member of Discosorida belonging to the family Entimoceratidae. The type species is B. stichlingi.
Boreioteuthis mirifica	Sp. nov	Valid	Efremenko	Early Cretaceous			A belemnite belonging to the family Cylindroteuthididae.
Chisiloceras maocaopuense^[33]	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)			A member of the family Endoceratidae.
Domatoceras tuckeri^[34]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation		A member of the family Grypoceratidae.
Endolobus sturgeoni	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation		A member of the family Koninckioceratidae.
Enoploceras rieberi^[35]	Sp. nov	Valid	Pieroni	Middle Triassic (Anisian)	Besano Formation
Eximioceras^[36]	Gen. et sp. nov	Valid	Shchedukhin	Early Permian			A member of the family Rhiphaeoceratidae. The type species is E. venustum.
Holmiceras havelense^[37]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone
Kotoceras minosiphonatum	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)			A member of the family Endoceratidae.
Lispoceras orbis^[38]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Lituites baculus	Sp. nov	Valid	Aubrechtová & Korn	Middle Ordovician
Lituites bottkei	Sp. nov	Valid	Aubrechtová & Korn	Middle Ordovician
Lituites clavis	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone
Lituites dewitzi	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone
Lituites kruegeri	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone
Lituites nebeni	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone
Maccoyoceras concavum	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian–Viséan boundary interval)			A member of the family Trigonoceratidae.
Maccoyoceras habadraense	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Maccoyoceras pentagonum	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Maccoyoceras saharensis	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Metacoceras quadratum	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation		A member of the family Tainoceratidae.
Michelinoceras minutum	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian and Sandbian)
Michelinoceras multiseptum	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)
Pararhiphaeoceras parvum	Sp. nov	Valid	Shchedukhin	Early Permian			A member of the family Rhiphaeoceratidae.
Planetoceras destrictum	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Planetoceras transforme	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Pleuronautilus chisatoi^[39]	Sp. nov	Valid	Ehiro	Permian (Wordian)	Takakurayama Formation		A member of the family Tainoceratidae.
Proterovaginoceras rapidum	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Darriwilian)			A member of the family Endoceratidae.
Rhynchorthoceras kranepuhlense	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone
Rhynchorthoceras rugium	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone
Rineceras multituberculatum	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Rineceras rectangulatum	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Rineceras tenerum	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Solenochilus jackense	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation		A member of the family Solenochilidae.
Stroboceras ancilis	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Viséan)			A member of the family Trigonoceratidae.
Stroboceras mane	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Stroborineceras	Gen. et 2 sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae. The type species is S. insalahensis; genus also includes new species S. felis and possibly also "Nautilus (Trematodiscus)" meekianus Winchell (1862) and "Nautilus (Trematodiscus)" strigatus Winchell (1862).
Syllipsimopodi^[40]	Gen. et sp. nov	Disputed	Whalen & Landman	Carboniferous (Serpukhovian)	Bear Gulch Limestone		A member of Coleoidea of uncertain affinities. Originally assigned to the clade Vampyropoda; Klug et al. (2023) considered it to be a probable junior synonym of Gordoniconus beargulchensis.^[41] ^[42] The type species is S. bideni, named for Joe Biden.^[43]
Thrincoceras devolvere	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Tohokubelus^[44]	Gen. et sp. nov	Valid	Niko & Ehiro	Early Triassic (Olenekian)	Osawa Formation		A sinobelemnitid belemnite. Genus includes new species T. takaizumii.
Trilacinoceras filix	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone		A member of the family Lituitidae.
Trilacinoceras knoefleri	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone		A member of the family Lituitidae.
Trilobitoceras	Gen. et sp. et comb. nov	Valid	Korn & Bockwinkel	Carboniferous			A member of the family Trigonoceratidae. The type species is T. peculiaris; genus also includes "Nautilus (Trematodiscus)" planidorsalis Winchell (1862).
Vestinautilus angulatus	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Vestinautilus bicristatus	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Vestinautilus concinnus	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Vestinautilus inflexus	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Vestinautilus padus	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous			A member of the family Trigonoceratidae.
Vestinautilus papilio	Sp. nov	Valid	Korn & Bockwinkel	Carboniferous (Tournaisian)			A member of the family Trigonoceratidae.
Xainzanoceras densum	Sp. nov	Valid	Fang in Fang et al.	Ordovician (Katian)	Pagoda Formation		Possibly a member of the family Diestoceratidae.

Other cephalopod research

• A study on the phylogenetic relationships of Cambrian and Ordovician cephalopods is published by Pohle et al. (2022).^[45]
• Putative tentaculitoid Iwakiella ichiroi is reinterpreted as a sphaerorthoceratid orthocerid cephalopod by Niko (2022).^[46]
• A study on the identity of enigmatic black amorphous fossils often associated with belemnoid remains, based on the analysis of fossil material from the Upper Triassic (Carnian) Polzberg Lagerstätte (Austria), is published by Lukeneder & Lukeneder (2022), who interpret the enigmatic fossils as likely representing the preservation of a mineralized and secondarily carbonized cephalic-ocular-arm-cartilage complex of the belemnoid Phragmoteuthis bisinuata.^[47]
• A study on structures interpreted by Eduard Suess as beaks of Phragmoteuthis bisinuata is published by Doguzhaeva et al. (2022), who reinterpret the studied structures as cartilaginous remains of a prey, presumably juvenile fish; the authors also report a gladius of a previously unknown Carnian teuthid from the Cave del Predil (Italy), possibly similar to the Permian taxon Glochinomorpha stifeli, as well as an upper beak of a coleoid from the Anisian Buchenstein Formation (Italy), demonstrating typical coleoid upper beak morphology and composition, and interpreted by the authors as likely indicative of similar upper beak structure in P. bisinuata.^[48] Their interpretation of purported beaks of P. bisinuata as remains of a prey is subsequently contested by Lukeneder & Lukeneder (2022).^{[49] [50]}
• Belemnite species Liobelus acrei, otherwise known from the latest Jurassic and Early Cretaceous of the Boreal Realm, is reported from the Valanginian of the Vocontian Basin (France) by Mutterlose, Picollier & Dzyuba (2022), who interpret this finding as evidence of an isolated immigration of belemnites from the north in the early Valanginian, and establish two taxonomically different faunas of Tethyan belemnites.^[51]
• A study on the age of belemnites from the lower Santonian "Sponge horizon" of the Mozhzhelloovrazhnaya Formation in the Lower Volga region between Saratov and Volgograd (Russia), some of which are interpreted as redeposited from middle and upper Coniacian deposits on the basis of strontium, carbon and oxygen isotope data (with the first representatives of the genus Belemnitella interpreted as late Coniacian in age), is published by Zakharov et al. (2022).^[52]
• Revision of the type specimens of Eothinoceras americanum is published by Evans & Cichowolski (2022), who restrict the genus Eothinoceras to the type species, and propose a new scheme for classification of the genera belonging to the order Cyrtocerinida.^[53]
• A study on the distribution of nautiloids throughout the Cenozoic is published by Kiel, Goedert & Tsai (2022), who interpret their findings as indicating that from the Oligocene onward nautiloids became extinct in areas where pinnipeds appeared, while cetaceans (with possible exception of simocetids and agorophiids) were not found to significantly affect the demise of nautiloids.^[54]
• Fossil material of "Kummelonautilus" taiwanum is described from the Miocene Houdongkeng Formation (Taiwan) by Goedert, Kiel & Tsai (2022), who provisionally transfer this species to the genus Nautilus, and review the fossil record of the genus Nautilus in the Indo-Pacific region.^[55]
• A study on the external and internal morphology, affinities and ecology of Vampyronassa rhodanica, based on data on soft tissue of specimens from the La Voulte-sur-Rhône (France), is published by Rowe et al. (2022).^[56]
• First cephalopod statoliths from the Lower Cretaceous of Poland and United Kingdom are described by Pindakiewicz et al. (2022), who compare Mesozoic statoliths to those of extant cephalopods, and find the closest resemblance between Mesozoic forms and statoliths of extant idiosepiids.^[57]
• A study on the fossil material of Belosaepia from the Egemkapel Clay Member of the Tielt Formation and the Roubaix Clay Member of the Kortrijk Formation (Belgium), applying micro-CT imaging to the studied fossils, is published by Goolaerts et al. (2022), who identify growth lines in the studied fossils, and interpret their findings as indicating that the studied material belongs to representatives of a single species (B. tricarinata).^[58]

Bivalves

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Acila (Truncacila) interferencia^[59]	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation		A species of Acila.
Anniedhondtella^[60]	Gen. et sp. nov	Valid	Jagt, Cooper & Jagt-Yazykova	Late Cretaceous (Maastrichtian)	Maastricht Formation		A member of Trigoniida belonging to the family Rutitrigoniidae. Genus includes new species A. rieui.
Aphanaia sobopolensis^[61]	Sp. nov	Valid	Biakov & Kutygin	Early Permian
Arcopagia esteponensis^[62]	Sp. nov	Valid	Brunetti & Della Bella	Pliocene			A member of the family Tellinidae.
Arcopagia ligustica	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene			A member of the family Tellinidae.
Arcopella crestii	Sp. nov	Valid	Brunetti & Della Bella	Late Pliocene			A member of the family Tellinidae.
Arcopella pseudomista	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene			A member of the family Tellinidae.
Bathymodiolus labayugensis	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation		A species of Bathymodiolus.
Choniocardia olivieri^[63]	Nom. nov	Valid	Pacaud	Eocene			A member of the family Carditidae; a replacement name for Cardita (Miodon) bezanconi Cossmann (1887).
Conchocele pangasinanensis	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation		A member of the family Thyasiridae.
Crassatella (Bathytormus) ignava^[64]	Sp. nov	Valid	Berezovsky	Eocene	Malinovskaya Formation		A member of the family Crassatellidae.
Crassatella lassa^[65]	Sp. nov	Valid	Berezovsky	Late Eocene
Crassostrea sophiae	Nom. nov	Valid	Pacaud	Eocene			A species of Crassostrea; a replacement name for Ostrea elegans Deshayes (1832).
Cyclotellina hugoi	Nom. nov	Valid	Pacaud	Eocene			A member of the family Tellinidae; a replacement name for Tellina (Cyclotellina) bezanconi Cossmann (1906).
Gryphaea (Gryphaea) lavernockensis^[66]	Sp. nov	Valid	Hodges	Early Jurassic
Kasimlara ? antiqua^[67]	Sp. nov	Valid	Fang & Carter	Permian			A member of the family Kalenteridae belonging to the subfamily Healeyinae.
Liaoningoconcha^[68]	Nom. nov		Fang & Carter	Late Triassic	Yangcaogou Formation		A member of Cardiida belonging to the family Ferganoconchidae; a replacement name for Liaoningia Yu & Dong (1993). Ceccolini & Cianferoni (2021) proposed a replacement name Sinoliaoningia for the same genus in an earlier publication.^[69]
Megaxinus gorrospei	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation		A member of the family Lucinidae.
Meleagrinella golberti^[70]	Sp. nov	Valid	Lutikov & Arp	Early Jurassic (Toarcian)			A member of the family Oxytomidae. Published online in 2023, but the issue date is listed as December 2022.
Meleagrinella prima	Sp. nov	Valid	Lutikov in Lutikov & Arp	Early Jurassic (Toarcian)			A member of the family Oxytomidae. Published online in 2023, but the issue date is listed as December 2022.
Monginaia^[71]	Gen. et comb. nov	In press	Delvene et al.	Early Cretaceous (Barremian)	Camarillas Formation		A member of Unionida belonging to the new family Monginaiidae. Genus includes "Elliptio" galvensis Mongin (1966).
Myonia komiensiformis^[72]	Sp. nov	Valid	Biakov	Permian			A member of Pholadomyida belonging to the family Sanguinolitidae.
Myonia kutygini	Sp. nov	Valid	Biakov	Permian			A member of Pholadomyida belonging to the family Sanguinolitidae.
Oudardia pliorecens^[73]	Sp. nov	Valid	Brunetti & Della Bella	Plio-Pleistocene			A member of the family Tellinidae.
Palaeomutela (Palaeanodonta) berrutii^[74]	Sp. nov	Valid	Silantiev et al.	Permian (Kungurian)	Collio Formation		A member of Actinodontida belonging to the family Palaeomutelidae.
Palaeomutela (Palaeanodonta) guncinaensis	Sp. nov	Valid	Silantiev et al.	Permian (Kungurian)	Guncina Formation		A member of Actinodontida belonging to the family Palaeomutelidae.
Peronidia iberica	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene			A member of the family Tellinidae.
Peronidia intermedia	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene			A member of the family Tellinidae.
Pliocardia ballesterosi	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation		A species of Pliocardia.
Praebarrettia magnifica^[75]	Sp. nov		Mitchell	Late Cretaceous (Campanian)	Dry Hill Formation		A rudist bivalve.
Psammotreta santacatalinae	Sp. nov	Valid	Brunetti & Della Bella	Early Pliocene			A member of the family Tellinidae.
Sisonia	Gen. et sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation		A bivalve of uncertain affinities. The type species is S. frijellanae.
Squiresica^[76]	Gen. et comb. nov	Valid	Hybertsen, Goedert & Kiel	Oligocene	Lincoln Creek Formation	( )	A member of the family Vesicomyidae. The type species is "Archivesica" knapptonensis Amano & Kiel (2007); genus also includes "Archivesica" marincovichi Kiel & Amano (2010).
Verneuilnodon	Gen. et comb. nov	Valid	Fang & Carter	Permian			A member of the family Kalenteridae belonging to the subfamily Healeyinae; a new genus for "Mytilus (Modiola)" pallasi de Verneuil (1845).
Whitfieldiella catherinei	Sp. nov		Mitchell	Late Cretaceous (Campanian)	Dry Hill Formation		A rudist bivalve.
Whitfieldiella gracilis	Sp. nov		Mitchell	Late Cretaceous (Campanian)	Dry Hill Formation		A rudist bivalve.
Wolfgangella	Gen. et 2 sp. nov	Valid	Jagt, Cooper & Jagt-Yazykova	Late Cretaceous (Maastrichtian)	Maastricht Formation		A member of Trigoniida belonging to the family Rutitrigoniidae. Genus includes new species W. neilpearti and W. ignota.

Bivalve research

• Late Triassic bivalves are identified for the first time in olistostromes from the Sêwa Formation in the Riganpeicuo area (Tibet, China) by Xiao et al. (2022).^[77]
• A study on the impact of the Triassic–Jurassic extinction event on the body-size distribution of bivalves, based on data from three study sites in the United Kingdom, is published by Opazo & Twitchett (2022).^[78]
• Wolkenstein (2022) presents evidence of abundance and diversity in the color patterns of Pleuronectites laevigatus from the Middle Triassic Muschelkalk of Central Europe.^[79]
• Asato, Nakayama & Imai (2022) study color patterns of freshwater bivalves from the Lower Cretaceous Kitadani Formation (Japan), providing evidence of the presence of similar shell color patterns in extant and Early Cretaceous bivalves.^[80]

Gastropods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Akburunella laminaris^[81]	Sp. nov	Valid	Guzhov	Miocene			Published online in 2023, but the issue date is listed as December 2022.
Akburunella nefanda archaica	Ssp. nov	Valid	Guzhov	Miocene			Published online in 2023, but the issue date is listed as December 2022.
Akburunella sinuosa	Sp. nov	Valid	Guzhov	Miocene			Published online in 2023, but the issue date is listed as December 2022.
Akburunella spinosa	Sp. nov	Valid	Guzhov	Miocene			Published online in 2023, but the issue date is listed as December 2022.
Ampezzopleura barremica^[82]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation
Angularia corallina^[83]	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation		A member of Caenogastropoda belonging to the family Purpuroideidae.
Angulomphalus senowbarii	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation		A member of Vetigastropoda belonging to the family Ptychomphalidae.
Anisocycla subcylindrica^[84]	Sp. nov	Valid	Pavia et al.	Pliocene			A member of the family Murchisonellidae.
Antiphora aurora	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation
Aplexa canculae^[85]	Sp. nov	Valid	Pacaud	Paleocene (Thanetian)			A species of Aplexa.
Aplexa cotiensis	Nom. nov	Valid	Pacaud	Eocene			A species of Aplexa; a replacement name for Physa (Aplecta) brevispirata Cossmann & Pissarro (1913).
Aplexa iota	Sp. nov	Valid	Pacaud	Paleogene			A species of Aplexa.
Aplexa oryza	Sp. nov	Valid	Pacaud	Paleogene			A species of Aplexa.
Aplexa risleius	Sp. nov	Valid	Pacaud	Paleocene (Thanetian)			A species of Aplexa.
Aplexa zhui	Nom. nov	Valid	Pacaud	Early Cretaceous (Albian)	Dalazi Formation		A species of Aplexa; a replacement name for Physa fusiformis Zhu (1980).
Arachnotropis^[86]	Gen. et comb. nov	Valid	Pérez, Ferrari & Ezcurra	Paleocene–Eocene		Antarctica	A member of the family Eucyclidae. The type species is "Calliotropis" microglyptophorus Darragh (1997); genus also includes "Calliotropis" antarchais Stilwell (2005).
Archaeocyclotus brevivillosus^[87]	Sp. nov	In press	Bichain et al.	Cretaceous	Burmese amber		A member of the family Cyclophoridae.
Archaeostylus^[88]	Gen. et sp. nov	Valid	Brook & Hayward	Pliocene (Waipipian)	Kaawa Formation		A member of the family Bothriembryontidae belonging to the subfamily Placostylinae. The type species is A. manukauensis.
Astralium ena^[89]	Sp. nov	Valid	Tomida, Inoue & Kase	Miocene	Ena Limestone		A species of Astralium.
Austrocypraea jimgracei^[90]	Sp. nov	Valid	Southgate & Roberts	Pliocene			A member of the family Cypraeidae.
Barycypraea iungo^[91]	Sp. nov	Valid	Seccombe & Aiken	Pleistocene			A species of Barycypraea.
Bathrotomaria teodorii^[92]	Sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)			A member of the family Pleurotomariidae.
Baylea tenera^[93]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation		A member of the family Eotomariidae.
Brachystomia succineiformis	Sp. nov	Valid	Guzhov	Miocene			A species of Brachystomia. Published online in 2023, but the issue date is listed as December 2022.
Ceritella convexa^[94]	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)			A member of the family Ceritellidae.
Choshipleura	Gen. et sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation		Genus includes new species C. striata.
Clathromangelia acuticostata^[95]	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Clathromangelia.
Clathromangelia mulderi	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Clathromangelia.
Clathromangelia oliverioi	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Clathromangelia.
"Clavatula" ariejansseni^[96]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Serravallian)	Týrtar Formation		A member of the family Clavatulidae.
"Clavatula" atatuerki	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Serravallian)	Týrtar Formation		A member of the family Clavatulidae.
"Clavatula" iberica^[97]	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)			A member of the family Clavatulidae.
"Clavatula" irisae	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation		A member of the family Clavatulidae.
"Clavatula" jarzynkae	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation		A member of the family Clavatulidae; a replacement name for Clavatula auingeri Finlay (1927).
"Clavatula" nana	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)			A member of the family Clavatulidae.
"Clavatula" pliogradata	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)			A member of the family Clavatulidae.
Clavatula sorini	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation	?	A species of Clavatula.
Clithon cooncreekense^[98]	Sp. nov	Valid	Garvie & Symonds	Late Cretaceous (Maastrichtian)	Coon Creek Formation		A species of Clithon.
Clithon duomillias	Sp. nov	Valid	Garvie & Symonds	Eocene	Cook Mountain Formation		A species of Clithon.
Clithon lisae	Sp. nov	Valid	Garvie & Symonds	Eocene	Cook Mountain Formation		A species of Clithon.
Clithon traceyi	Sp. nov	Valid	Garvie & Symonds	Eocene	Cook Mountain Formation		A species of Clithon.
Columbellaria rara^[99]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation		A member of the family Colombellinidae.
Conotomaria lecatae^[100]	Sp. nov	Valid	Chaix & Grenier	Late Cretaceous (Campanian)			A member of the family Pleurotomariidae.
Conus damianakisi^[101]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)			A species of Conus.
Conus davolii	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)		?	A species of Conus.
Conus dellabellai	Sp. nov	Valid	Pavia et al.	Pliocene			A species of Conus.
Conus ictini	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Serravallian and Tortonian)			A species of Conus.
Conus lauriatragei	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Langhian to Tortonian)			A species of Conus.
Conus moissettei	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)			A species of Conus.
Conus villalvernensis	Sp. nov	Valid	Pavia et al.	Pliocene			A species of Conus.
Coptocheilus kachinensis^[102]	Sp. nov		Yu, Zhuo & Páll-Gergely	Cretaceous	Burmese amber		A species of Coptocheilus.
Cornirostra anistratenkorum umbilicaris	Ssp. nov	Valid	Guzhov	Miocene			Published online in 2023, but the issue date is listed as December 2022.
Crepidula bellardii	Sp. nov	Valid	Pavia et al.	Pliocene			A species of Crepidula.
Cretadiostoma	Gen. et 2 sp. nov		Yu, Zhuo & Páll-Gergely	Cretaceous	Burmese amber		A member of the family Pupinidae. Genus includes new species C. caperatum and C. umbilicarinatum.
Cryptoptyxis? fortiter	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation		A member of the family Maoraxidae.
Cyrillia clathrataeformis	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Cyrillia.
Cyrillia saldubensis	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Cyrillia.
Dentitriangularum^[103]	Gen. et sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)			A member of the family Cypraeidae. The type species is D. pacaudi.
Diaugasma mediterranea	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Diaugasma.
Dictyotomaria turrisbabel	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Caddo Creek Formation		A member of the family Phymatopleuridae.
Diempterus? multicostatus	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation		A member of the family Aporrhaidae.
Edrozeba	Gen. et 4 sp. et comb. nov	Valid	Guzhov	Miocene			A member of the family Hydrobiidae. The type species is E. caeca; genus also includes new species E. angulata, E. minuta and E. striata, as well as "Hydrobia" enikalensis Kolesnikov (1935). Published online in 2023, but the issue date is listed as December 2022.
Eirlysella	Gen. et sp. et comb. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation		A member of the family Luciellidae. The type species is E. buckhornensis; genus also includes "Trochus" hissingerianus de Koninck (1843), "Pleurotomaria" squamula Phillips (1836) and "Luciellina" ocultabanda Kues & Batten (2001).
Endoplocus acutus	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)			A member of the family Nerineidae.
Endoplocus inflatus	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)			A member of the family Nerineidae.
Ericusa ngayawang^[104]	Sp. nov	Valid	Yates	Miocene (Langhian)	Cadell Formation		A species of Ericusa.
Exelissa ursannella	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation		A member of Cerithioidea belonging to the family Cryptaulacidae.
Faureicerithium	Gen. et sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)			A member of the family Campanilidae. The type species is F. bidentata.
Favartia plioelata^[105]	Sp. nov	Valid	Brunetti & Della Bella	Pliocene			A species of Favartia.
Granulatocincta	Gen. et comb. et 3 sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene and Pliocene			A member of the family Clavatulidae. The type species is "Pleurotoma" granulato-cincta Münster in Goldfuss (1841); genus also includes new species G. callim, G. theoderichi and G. pelliscrocodili, as well as "Pleurotoma (Clavatula)" angelae Hoernes & Auinger (1891), "Clavatula" contorta Švagrovský (1958), "Pleurotoma (Clavatula)" nataliae Hoernes & Auinger (1891), "Pleurotoma" schreibersi Hörnes (1854), "Clavatula" calcarai Bellardi (1877), "Clavatula" curionii Bellardi (1877), "Clavatula" pellegrinii Simonelli (1896), "Pleurotoma" sotterii Michelotti (1847), "Clavatula" turbinata Bellardi (1877), "Clavatula" turgidula Bellardi (1877), "Clavatula" vigolensis Bellardi (1877), "Clavatula" rugata Bellardi (1877), "Murex" rusticus Brocchi (1814), "Pleurotoma" capgrandi Tournouër (1873), "Pleurotoma" obtruta Millet (1865), "Clavatula" delgadoi Vera-Peláez & Lozano-Francisco (2001) and possibly "Clavatula" boreointerrupta Kautsky (1925).
Hologyra callosa	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation		A member of the family Neritopsidae.
Hydrobia neofrauenfeldi	Sp. nov	Valid	Guzhov	Miocene			A species of Hydrobia. Published online in 2023, but the issue date is listed as December 2022.
Hydrobia orientalis^[106]	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation		A species of Hydrobia.
Iberellus colladoi^[107]	Sp. nov	Valid	Juárez-Ruiz & Altaba	Pliocene			A species of Iberellus.
Itieroptygmatis cylindrata	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)			A member of Nerineoidea belonging to the family Itieriidae.
Jenneria sandrae^[108]	Sp. nov	Valid	Daughenbaugh	Pleistocene (Chibanian)	Probably Bermont Formation		A species of Jenneria.
Leufroyia gradata	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Leufroyia.
Loriolotrema? nodosa	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation		A member of the family Brachytrematidae.
Lyria estotiensis^[109]	Sp. nov	Valid	Bail, Ledon & Pacaud	Oligocene (Chattian)			A species of Lyria.
Lyria lozoueti	Sp. nov	Valid	Bail, Ledon & Pacaud	Miocene (Aquitanian)			A species of Lyria.
Maoristylus pliocenicus	Sp. nov	Valid	Brook & Hayward	Pliocene (Waipipian)	Kaawa Formation		A member of the family Bothriembryontidae belonging to the subfamily Placostylinae.
Megaclavatula	Gen. et 2 sp. et comb. nov	Valid	Harzhauser, Landau & Janssen	Miocene		?	A member of the family Clavatulidae. The type species is M. grunerti; genus also includes new species M. pilleri, as well as "Pleurotoma (Clavatula)" amaliae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" antoniae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" evae Hoernes & Auinger (1891), "Pleurotoma" laevigatum Eichwald (1830), "Perrona" nemethi Kovács & Vicián (2021), "Pleurotoma" neudorfensis Schaffer (1898), "Pleurotoma tuberculosa" polonica Pusch (1837), "Pleurotoma (Clavatula) asperulata" var. subsculpta Schaffer (1912), "Pleurotoma" ernesti Toula (1901), "Pleurotoma" asperulata Lamarck (1822), Clavatula saubrigiana laurensii Peyrot (1931) and possibly "Pleurotoma (Clavatula)" szontaghi Strausz (1926), "Clavatula" taurofusulata Sacco (1890), "Clavatula" subdepressa Sacco (1890) and "Clavatula" tortonica Peyrot (1931).
Metacerithium boshuae	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation		A member of the family Metacerithiidae.
Metalycaeus fossilis^[110]	Sp. nov	In press	Yu, Páll-Gergely & Salvador	Miocene	Zhangpu amber		A species of Metalycaeus.
Microandonia	Gen. et sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A member of the family Raphitomidae. The type species is M. minutissima.
Microdrillia plioserratula^[111]	Sp. nov	Valid	Landau & Harzhauser	Pliocene			A species of Microdrillia.
Microschiza rhaetica	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation		A member of the family Purpurinidae.
Mitromorpha (Mitrolumna) velerinensis	Sp. nov	Valid	Landau & Harzhauser	Pliocene		?	A species of Mitromorpha.
Muricopsis landaui^[112]	Sp. nov	Valid	Merle in Merle, Garrigues & Pointier	Miocene			A species of Muricopsis.
Napaeus guanche^[113]	Sp. nov	In press	Miller et al.	Middle Pleistocene			A species of Napaeus.
Napaeus lipauges	Sp. nov	In press	Miller et al.	Middle Pleistocene			A species of Napaeus.
Narrimania saldubensis^[114]	Sp. nov	Valid	Landau & Mulder	Pliocene
Neoperrona	Gen. et comb. et sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)			A member of the family Clavatulidae. The type species is "Perrona" harzhauseri Kovács & Vicián (2021); genus also includes new species N. zoltanorum, as well as "Clavatula" taurinensis Bellardi (1877).
Neridomus mccallae	Sp. nov	Valid	Garvie & Symonds	Early Cretaceous (Albian)	Edwards Formation		A member of the family Neridomidae.
Neridomus minutissimus	Sp. nov	Valid	Garvie & Symonds	Early Cretaceous (Albian)	Walnut Formation		A member of the family Neridomidae.
Nerinea donosa	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)			A member of the family Nerineidae.
Neuburgensia	Gen. et comb. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian to Tithonian)			Possibly a member of Cerithioidea. The type species is "Gymnocerithium" concavum Janicke (1966); genus also includes "Cerithium" amabile Zittel (1873), "Gymnocerithium"? convexoconcavum Gründel, Keupp & Lang (2019), "Cerithium" crenato-cinctum Zittel (1873), "Cerithium" involvens Zittel (1873) and "Cerithium" perrotundum Cossmann (1913).
Nystia conoidalis^[115]	Sp. nov	Valid	Kadolsky & Morton	Oligocene (Rupelian)			A species of Nystia.
Odetta chirlii	Sp. nov	Valid	Pavia et al.	Pliocene			A species of Odetta.
Odostomia caucasica	Sp. nov	Valid	Guzhov	Miocene			A species of Odostomia. Published online in 2023, but the issue date is listed as December 2022.
Oleginina	Gen. et sp. et comb. nov	Valid	Harzhauser, Landau & Janssen in Landau, Harzhauser & Giannuzzi-Savelli	Miocene (Burdigalian to Tortonian)			A member of the family Clavatulidae. The type species is O. mandici; genus also includes "Pleurotoma (Clavatula)" agathae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" brigittae Hoernes & Auinger (1891), "Pleurotoma" doderleini Hörnes (1854), "Pleurotoma (Clavatula)" dorotheae Hoernes & Auinger (1891), "Pleurotoma" rumana Simionescu & Barbu (1940), Clavatula schreibersi szokolyensis Strausz (1960), "Pleurotoma" winterlingensis Quenstedt (1884), "Clavatula" aradasi Bellardi (1877), "Clavatula" baccifera Bellardi (1877), "Clavatula" consularis Bellardi (1877) and "Clavatula" stazzanensis Bellardi (1877). The original generic name was Olegia, but this name turned out to be preoccupied by Olegia Shaposhnikov (1979), necessitating the creation of a replacement name.
Oligoptyxis khorasanica^[116]	Sp. nov	Valid	Guzhov & Nazemi in Raisossadat et al.	Early Cretaceous (Aptian)			A member of the family Nerineidae.
Ondina curta	Sp. nov	Valid	Pavia et al.	Pliocene			A species of Ondina.
Ondina elongata	Sp. nov	Valid	Pavia et al.	Pliocene			A species of Ondina.
Ondina pseudovitrea	Sp. nov	Valid	Pavia et al.	Pliocene			A species of Ondina.
Ophieulima lobilloensis	Sp. nov	Valid	Landau & Mulder	Pliocene			A species of Ophieulima.
Otostoma cattoi	Sp. nov	Valid	Garvie & Symonds	Early Cretaceous (Albian)	Edwards Formation		A member of the family Otostomidae.
Ovalotrema	Gen. et comb. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)			A member of the family Brachytrematidae. The type species is "Turbo" cotteausius d'Orbigny (1853); genus might also include "Turbo" greppini De Loriol in De Loriol & Koby (1895).
Palaeoleuca edeni^[117]	Sp. nov	Valid	Pacaud	Paleocene (Thanetian) to Eocene (Ypresian)			A member of the family Ellobiidae.
Palaeoleuca gradata	Sp. nov	Valid	Pacaud	Paleocene (Thanetian) to Eocene (Ypresian)			A member of the family Ellobiidae.
Palaeoleuca hulvizensis	Nom. nov	Valid	Pacaud	Paleocene			A member of the family Ellobiidae; a replacement name for Auricula (Conovulus) pygmaea Morris (1854).
Paleothyca^[118]	Gen. et sp. nov	Valid	Forner, Domènech & Martinell	Early Cretaceous (Albian)			A member of the family Eulimidae. The type species is P. quinionarii.
Panneritina^[119]	Gen. et comb. et sp. nov	In press	Salvador & Yu	Jurassic and Cretaceous			A member of the new family Neritidae. Genus includes Chinese species formerly assigned to the genus Mesoneritina ("M." crassa Pan, 1982, "M." dakangensis Pan, 1978, "M." gansuensis Yu & Guo, 1982, "M." nanshanensis Pan, 1984, "M." opima Yu, 1974, "M." pustula Pan, 1980), as well as new species P. ambrae from the Kachin amber.
Papuliscala spinosa	Sp. nov	Valid	Landau & Mulder	Pliocene			A species of Papuliscala.
Paragoniozona ornata	Nom. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Lazy Bend Formation		A member of the family Phymatopleuridae; a replacement name for Pleurotomaria aspera Girty (1934).
Paragoniozona yanceyi	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation		A member of the family Phymatopleuridae.
Perrona grossi	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Florian Formation		A species of Perrona; a replacement name for Pleurotoma (Clavatula) auingeri Hilber (1879).
Perrona ilonae	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Sámsonháza Formation		A species of Perrona; a replacement name for Clavatula vindobonensis nodosa Csepreghy-Meznerics (1954).
Perrona koeberli	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation		A species of Perrona.
Perrona loetschi	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation		A species of Perrona.
Perrona wanzenboecki	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation		A species of Perrona; a replacement name for Pleurotoma concinna Handmann (1883).
Peruvispira kanchanaburiensis^[120]	Sp. nov	In press	Karapunar, Nützel & Ketwetsuriya	Permian	Ratburi Group		A member of the family Goniasmatidae.
Peruvispira oklahomaensis	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation		A member of the family Goniasmatidae.
Phymatopleura conica	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation		A member of the family Phymatopleuridae.
Phymatopleura girtyi	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Wewoka Formation		A member of the family Phymatopleuridae.
Platyphysa nagensis	Nom. nov	Valid	Pacaud	Cretaceous			A replacement name for Physa prinsepii elongata Hislop (1860).
Pleurotomaria cloutetsensis	Sp. nov	Valid	Chaix in Chaix & Plicot	Late Cretaceous (Santonian)
Pleurotomella obesula	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Pleurotomella.
Pleurotomella turrita	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Pleurotomella.
Pomatiasia	Gen. et comb. nov	Valid	Guzhov	Miocene			A member of the family Hydrobiidae. Genus includes "Amnicola" cyclostomoides Sinzov (1880). Published online in 2023, but the issue date is listed as December 2022.
Propefavartia	Gen. et comb. nov	Valid	Merle in Merle, Garrigues & Pointier	Eocene			A member of the family Muricidae belonging to the subfamily Muricopsinae. The type species is "Murex" distans Deshayes (1835).
Pseudamnicola roblesi^[121]	Sp. nov	Valid	Albesa, López & Crespo	Early Miocene	Ribesalbes-Alcora Basin		A species of Pseudamnicola.
Pseudamnicola taodonggouensis	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation		A species of Pseudamnicola.
Pseudodaphnella iberica	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Pseudodaphnella.
Pseudodaphnella velerinensis	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Pseudodaphnella.
Pseudodostia lisbonensis	Sp. nov	Valid	Garvie & Symonds	Eocene	Lisbon Formation		A member of the family Neritidae.
Pseudomelania yamadai	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation
Pseudonerinea ? pseudomelaniformis	Sp. nov	Valid	Gründel et al.	Late Jurassic (Kimmeridgian)			A member of Nerineoidea belonging to the family Pseudonerineidae.
Purpuripullina	Gen. et comb. nov	Valid	Guzhov & Raisossadat in Raisossadat et al.	Cretaceous			The type species is "Natica" scalaris Conrad (1852).
Purpuroidea moosleitneri	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation		A member of Caenogastropoda belonging to the family Purpuroideidae.
Pusionella hofmanni	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation		A species of Pusionella.
Ringicula costata chokrakensis^[122]	Ssp. nov	Valid	Guzhov	Miocene			Published online in 2023, but the issue date is listed as December 2022.
Ringicula costata paratethica	Ssp. nov	Valid	Guzhov	Miocene			Published online in 2023, but the issue date is listed as December 2022.
Ringicula subglobosa	Sp. nov	Valid	Guzhov	Miocene			A species of Ringicula. Published online in 2023, but the issue date is listed as December 2022.
Saalensia lorioli	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation		A member of the family Brachytrematidae.
Shansiella (Oklahomaella) globilineata	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation		A member of the family Portlockiellidae.
Sigmesalia anatole^[123]	Sp. nov	Valid	Hoşgör & Pacaud	Paleocene (Danian)	Çaldağ Formation		A member of the family Turritellidae.
Sigmesalia erifensis	Sp. nov	Valid	Hoşgör & Pacaud	Paleocene (Danian)	Çaldağ Formation		A member of the family Turritellidae.
Skenea gofasi	Sp. nov	Valid	Landau & Mulder	Pliocene			A species of Skenea.
Specula estebbunensis	Sp. nov	Valid	Landau & Mulder	Pliocene			A species of Specula.
Spinatus ruizi^[124]	Sp. nov	Valid	Liverani & Wieneke	Miocene (Langhian)			A member of the family Strombidae.
Spiroscala quasipulchra	Nom. nov	Valid	Karapunar et al.	Permian			A member of the family Eotomariidae; a replacement name for Euconospira pulchra Batten (1958).
Spiroscala shwedagoniformis	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation		A member of the family Eotomariidae.
Striopusionella	Gen. et comb. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)			A member of the family Clavatulidae. The type species is "Clavatula" szekelyhidiae Kovács & Vicián (2021).
Stuoraxis kasei	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation		A member of the family Stuoraxidae.
Stuorella zapfei	Sp. nov	Valid	Nützel et al.	Late Triassic (Rhaetian)	Oberrhät Formation		A member of Vetigastropoda belonging to the family Stuorellidae.
Subpterynotus eocaenicus^[125]	Sp. nov		Merle, Lesport & Ledon	Eocene (Bartonian)			A species of Subpterynotus.
Subpterynotus mainotensis	Sp. nov		Merle, Lesport & Ledon	Miocene (Aquitanian)			A species of Subpterynotus.
Talassia plioalboranensis	Sp. nov	Valid	Landau & Mulder	Pliocene			A species of Talassia.
Teretia inflatissima	Sp. nov	Valid	Landau, Harzhauser & Giannuzzi-Savelli	Pliocene (Piacenzian)			A species of Teretia.
Thalassonerita hagai	Sp. nov	Valid	Kiel et al.	Miocene	Amlang Formation		A member of the family Phenacolepadidae.
Tomellana aueri	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation		A species of Tomellana.
Tomellana dulaii	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation		A species of Tomellana.
Tomellana onubensis	Sp. nov	Valid	Landau & Harzhauser	Early Pliocene			A species of Tomellana.
Tomellana postjouannetii	Sp. nov	Valid	Landau & Harzhauser	Pliocene (Piacenzian)			A species of Tomellana.
Tortaxis zhangpuensis	Sp. nov	In press	Yu, Páll-Gergely & Salvador	Miocene	Zhangpu amber
Tosapusia peregrina^[126]	Sp. nov	Valid	Bertaccini et al.	Pliocene			A species of Tosapusia.
Trigonostoma ariejansseni^[127]	Sp. nov	Valid	Roosen	Miocene			A species of Trigonostoma.
Tudorella lenneae^[128]	Sp. nov	Valid	Groh et al.	Pliocene			A species of Tudorella.
Tudorella rothei	Sp. nov	Valid	Groh et al.	Pliocene			A species of Tudorella.
Turbonilla yadongensis^[129]	Sp. nov		Li, Li & Garvie	Eocene (Ypresian)	Zhupure Formation		A species of Turbonilla.
Valvata complanusa	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation		A species of Valvata.
Varicoturbo^[130]	Gen. et sp. nov	Valid	Nützel, Schweigert & Karapunar	Late Triassic (Carnian)	San Cassiano Formation		A member of the family Seguenziidae. Genus includes new species V. microstriatus.
Worthenia (Yochelsonospira) kuesi	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Wetumka Formation		A species of Worthenia.
Xinjiangospira habita	Sp. nov	Valid	Zhu	Late Permian	Wutonggou Formation		A member of the family Hydrobiidae.

Gastropod research

• A study on transitions from water to land throughout the evolutionary history of gastropods is published by Vermeij & Watson-Zink (2022).^[131]
• A study on changes in gastropod diversity at the genus/subgenus level from the Norian to the Pliensbachian is published by Ferrari & Hautmann (2022), who report that gastropods lost 56% of genera/subgenera during the Triassic–Jurassic extinction event, which was much more than the average loss of marine life at the time, and attempt to determine potential causes of the end-Triassic extinctions of gastropods.^[132]
• A study aiming to determine the drivers of diversification for European freshwater gastropods over the past 100 million years is published by Neubauer et al. (2022).^[133]
• A study aiming to determine the factors which caused the diversification of European freshwater gastropods in the Late Cretaceous is published by Neubauer & Harzhauser (2022).^[134]
• Revision of the fossil material of Ancistrolepis from the Eocene (Ypresian) Llajas Formation (California, United States), representing the oldest occurrence of this genus reported to date, is published by Squires (2022).^[135]

Other molluscs

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Calloplax roederi^[136]	Sp. nov	Valid	Vendrasco, Powell & LaFollette	Miocene			A chiton.
Chiton solaris	Sp. nov	Valid	Vendrasco, Powell & LaFollette	Miocene			A chiton.
Dorispira avannga^[137]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A helcionellid.
Dorispira? lemdadensis^[138]	Sp. nov		Mghazli et al.	Cambrian
Dorispira septentrionalis	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A helcionellid.
Dorispira tavsenensis	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A helcionellid.
Dorispira tippik	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A helcionellid.
Erugoconus	Gen. et sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A member of Helcionelloida of uncertain affinities. The type species is E. acuminatus.
Hoarepora^[139]	Nom. nov	Valid	Mapes in Mazaev	Carboniferous (Pennsylvanian)			A rostroconch; a replacement name for Oxyprora Hoare, Mapes & Yancey (2002).
Ischnochiton loureiroi^[140]	Sp. nov	Valid	Dell'Angelo et al.	Pliocene	Mondego Basin		A chiton, a species of Ischnochiton.
Lepidochitona rochae	Sp. nov	Valid	Dell'Angelo et al.	Pliocene	Mondego Basin		A chiton, a species of Lepidochitona.
Scenella? siku	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A member of Helcionelloida. Originally tentatively assigned to the genus Scenella, but subsequently made the type species of the separate genus Hensoniconus.^[141]
Sermeqiconus	Gen. et comb. nov	Valid	Peel & Kouchinsky	Cambrian	Aftenstjernesø Formation		A helcionellid. The type species is "Figurina" polaris Peel (2021).
Tavseniconus	Gen. et sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A member of Helcionelloida of uncertain affinities. The type species is T. erectus.
Vendrascospira	Gen. et 2 sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation		A helcionellid. The type species is V. troelseni; genus also includes V. frykmani.
Yochelcionella gracilituba	Sp. nov		Mghazli et al.	Cambrian
Yochelcionella longituba	Sp. nov		Mghazli et al.	Cambrian

Other mollusc research

• Revision of the oldest known specimens of Jinonicella kolebabai from the Middle Ordovician strata of Ukraine and Belarus is published by Gubanov et al. (2022), who report the preservation of growth lines which were previously documented only in Silurian specimens.^[142]
• New information on the anatomy of Typhloesus wellsi is presented by Conway Morris & Caron (2022), who consider it plausible that Typhloesus was a pelagic mollusc with possible affinities with gastropods.^[143]

Notes and References

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3. Mitta . V. V. . The genus Cadomites (Ammonoidea: Stephanoceratidae) from the upper Bajocian-lower Bathonian (Middle Jurassic) of the Kuban River basin (Northern Caucasus, Russia) . 2022 . Paleontological Journal . 56 . 6 . 620–629 . 10.1134/S0031030122060089 . 2022PalJ...56..620M . 254249032 .
4. Amédro . F. . Baudouin . C. . Delanoy . G. . Devalque . C. . Robaszynski . F. . Une association d'ammonites rare dans la Zone à Romaniceras ornatissimum (Turonien moyen) du Massif d'Uchaux, Vaucluse, France . 2022 . Carnets Geol. . 22 . 9 . 333–393 . 10.2110/carnets.2022.2209 . 251192365 .
5. Esquivel-Macías . C. . León-Olvera . R. G. . Ramírez-Cardona . M. . Flores-Castro . K. . Reporte de Crioceratites (Gerth, 1925) con la descripción de C. tepehuaense n. sp. en el centro oriente de México y algunas consideraciones bioestratigráficas y paleobiogeográficas . 2022 . Tópicos de Investigación en Ciencias de la Tierra y Materiales . 9 . 9 . 1–12 . 10.29057/aactm.v9i9.9129 . 252743981 . free .
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