2022 in paleobotany explained

This paleobotany list records new fossil plant taxa that were to be described during the year 2022, as well as notes other significant paleobotany discoveries and events which occurred during 2022.

Algae

Charophytes

Name	Novelty	Status	Authors	Age	Unit	Location	Synonymized taxa	Notes	Images
Lamprothamnium elongatum^[1]	Sp. nov	In press	Feist & Floquet	Late Cretaceous				A charophyte.
Lamprothamnium ovoideum	Sp. nov	In press	Feist & Floquet	Late Cretaceous				A charophyte.
Pseudoharrisichara sedanoensis	Sp. nov	In press	Feist & Floquet	Late Cretaceous				A charophyte.

Charophyte research

• A study on the Paleocene charophyte flora from the South Gobi area in the Junggar Basin (China) and on the Paleogene fossil record of charophytes is published by Cao et al. (2022), who interpret their findings as evidence of the dispersal of charophyte lineages from Asia to Europe in the middle to late Eocene, possibly facilitated by waterbirds.^[2]

Chlorophytes

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ardeiporella^[3]	Gen. et comb. nov	Valid	Grgasović	Middle Triassic			A green alga belonging to the group Dasycladales. Genus includes "Oligoporella" karrerioidea Pia (1935).
Earltonella^[4]	Gen. et sp. nov		LoDuca in LoDuca et al.	Silurian (Llandovery)	Earlton Formation		A green alga belonging to the group Bryopsidales. Genus includes new species E. fredricksi.
Milanovicella? canadillana^[5]	Sp. nov	In press	Torromé & Schlagintweit	Late Cretaceous (Santonian–Campanian)			A green alga belonging to the group Dasycladales.
Neophysoporella	Gen. et comb. nov	Valid	Grgasović	Late Triassic, possibly also Middle Triassic			A green alga belonging to the group Dasycladales. Genus includes "Diplopora" lotharingica Benecke (1898), "Physoporella" jomdaensis Flügel & Mu (1982) and "Physoporella" zamparelliae Parente & Climaco (1999).
Protocodium^[6]	Gen. et sp. nov		Chai, Aria & Hua	Ediacaran	Dengying Formation		A green alga belonging to the family Codiaceae. Genus includes new species P. sinense.
Succodium luciae^[7]	Sp. nov	Valid	Vachard & Krainer	Permian-Triassic transition			A green alga belonging to the group Dasycladales.

Lycopodiopsida

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Lepacyclotes ordosensis^[8]	Sp. nov	Valid	Deng in Deng et al.	Middle Triassic (Ladinian)	Tongchuan Formation		A member of the family Isoetaceae.
Lycopodicaulis^[9]	Gen. et sp. nov	Valid	Herrera et al.	Early Cretaceous (late Barremian–early Aptian)	Huolinhe Formation		A member of the family Lycopodiaceae. Genus includes new species L. oellgaardii.
Multapicifolium^[10]	Gen. et sp. nov	Valid	Edwards, Li & Berry	Early Devonian			A member of Protolepidodendrales of uncertain phylogenetic placement. Genus includes new species M. sinense.
Nothostigma sepeensis^[11]	Sp. nov		Spiekermann, Jasper, Guerra-Sommer & Uhl in Spiekermann et al.	Permian (Cisuralian)			A member of Lycopodiopsida of uncertain affinities.
Omprelostrobus^[12]	Gen. et sp. nov		Liu et al.	Devonian (Famennian)	Wutong Formation		A member of Isoetales of uncertain affinities. Genus includes new species O. gigas.
Pleuromeia obovata^[13]	Sp. nov		Deng in Deng et al.	Middle Triassic (Ladinian)	Tongchuan Formation		A lycopsid.
Porongodendron^[14]	Gen. et sp. nov	Valid	Prestianni et al.	Carboniferous (Mississippian)			An isoetalean lycopsid. Genus includes new species P. minitensis.
Selaginella alata^[15]	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber		A species of Selaginella.
Selaginella cretacea^[16]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber		A species of Selaginella.

Lycopsid research

• Description of new fossil material of Guangdedendron micrum, providing new information on the morphology of this plant, is published by Gao et al. (2022).^[17]
• Xu, Liu & Wang (2022) describe new fossil material of Sublepidodendron grabaui from the Devonian (Famennian) Wutong Formation (China), providing new information on the morphology of the female reproductive organs of this plant.^[18]

Marchantiophyta

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Radula patrickmuelleri^[19]	Sp. nov	Valid	Feldberg, Schäfer-Verwimp & Renner in Feldberg et al.	Cretaceous (Albian-Cenomanian)	Burmese amber		A liverwort, a species of Radula.
Radula tanaiensis	Sp. nov	Valid	Feldberg, Schäfer-Verwimp & Renner in Feldberg et al.	Cretaceous (Albian-Cenomanian)	Burmese amber		A liverwort, a species of Radula.
Ricciopsis asturicus^[20]	Sp. nov	Valid	Santos et al.	Late Jurassic (Kimmeridgian)	Lastres Formation		A liverwort belonging to the family Ricciaceae.
Ricciopsis cortaderitaensis^[21]	Gen. et sp. nov	Valid	Savoretti et al.	Middle Triassic			A liverwort.
Ricciellites	Sp. nov	Valid	Savoretti et al.	Middle Triassic			A liverwort. Genus includes new species R. unsaltoensis.

Marchantiophyta research

• New specimens of Radula heinrichsii, providing new information on the morphology of this liverwort, are described from the Cretaceous Burmese amber by Wang et al. (2022).^[22]

Ferns and fern allies

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Acrostichopteris alcainensis^[23]	Sp. nov	Valid	Skog & Sender	Early Cretaceous (Albian)	Escucha Formation		A member of the family Hymenophyllaceae.
Coniopteris antarctica^[24]	Sp. nov	In press	Trevisan et al.	Late Cretaceous		Antarctica
Diodonopteris virgulata^[25]	Sp. nov	In press	Zhou et al.	Early Permian			A botryopteid fern.
Discosoropteris^[26]	Gen. et 2 sp. nov	In press	Pšenička et al.	Carboniferous (Pennsylvanian)	Kladno Formation		A leptosporangiate fern. Genus includes new species D. chlupatum and D. zlatkokvacekii.
Dryopterites beishanensis^[27]	Sp. nov	In press	Ren & Sun in Ren et al.	Early Cretaceous	Chijinbao Formation		A fern Announced online in 2022 officially published in 2023
Gleichenia nagalingumiae^[28]	Sp. nov		Cantrill et al.	Miocene			A species of Gleichenia.
Hymenophyllites angustus	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber		A member of the family Hymenophyllaceae. Originally described as a species of Hymenophyllites, but subsequently moved to the genus Trichomanes sensu lato by Li et al. (2023).^[29]
Hymenophyllites kachinensis	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber		A member of the family Hymenophyllaceae.
Hymenophyllites setosus	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber		A member of the family Hymenophyllaceae.
Microlepia burmasia^[30]	Sp. nov	Valid	Long, Wang & Shi in Long et al.	Cretaceous	Burmese amber		A fern of uncertain affinities. Originally described as a dennstaedtiaceous fern belonging to the genus Microlepia, but this classification was contested by Zhang (2024).^[31] Published online in 2022, but the issue date of the article naming it is listed as March 2023.
Mikasapteris^[32]	Gen. et sp. nov	Valid	Nishida et al.	Late Cretaceous	Yezo Group		A probable stem polypod leptosporangiate fern. Genus includes new species M. rothwellii.
Paralophosoria^[33]	Gen. et sp. nov	Valid	Morales-Toledo, Mendoza-Ruiz & Cevallos-Ferriz	Middle Jurassic	Otlaltepec Formation		A member of the family Dicksoniaceae. Genus includes new species P. jurassica.
Phyllotheca douroensis^[34]	Sp. nov		Barbosa et al.	Carboniferous (Gzhelian)	Douro Carboniferous Basin		A member of Equisetales.
Scolecopteris zhoui^[35]	Sp. nov	In press	Zhang et al.	Early Permian	Taiyuan Formation		A member of Marattiales belonging to the family Psaroniaceae.
Wolfeniana^[36]	Nom. nov	Valid	Deshmukh	Devonian	Hampshire Group		A member of Stauropteridales; a replacement name for Gillespiea Erwin & Rothwell (1989).

Fern and fern ally research

• Pecopteris lativenosa is interpreted as a member of the late Paleozoic marattialean family Psaroniaceae by Li et al. (2022).^[37]

Gnetales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Bassitheca^[38]	Gen. et sp. nov	Valid	Manchester et al.	Late Jurassic	Morrison Formation		A gnetale. Genus includes the species B. hoodiorum.
Dichoephedra^[39]	Gen. et sp. nov	In press	Ren et al.	Early Cretaceous	Chijinbao Formation		A member of the family Ephedraceae. Genus includes new species D. beishanensis.

Bennettitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Dictyozamites barnardi^[40]	Sp. nov	Valid	Saadatnejad	Late Triassic (Rhaetian)	Kalariz Formation		A member of Bennettitales.
Dictyozamites fakhri	Sp. nov	Valid	Saadatnejad	Late Triassic (Rhaetian)	Kalariz Formation		A member of Bennettitales.
Kimuriella^[41]	Gen. et sp. nov	Valid	Pott & Takimoto	Late Jurassic (Oxfordian)	Tochikubo Formation		A member of Bennettitales. Genus includes new species K. densifolia.
Zamites pateri^[42]	Sp. nov	Valid	Kvaček	Late Cretaceous (Cenomanian)	Peruc–Korycany Formation

Ginkgophytes

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Glossophyllum lanceolatum^[43]	Sp. nov		Sun & Deng in Sun et al.	Late Triassic
Glossophyllum panii	Sp. nov		Sun & Deng in Sun et al.	Late Triassic
Pseudotorellia baganuriana^[44]	Sp. nov	In press	Nosova & Kostina	Early Cretaceous (Aptian–Albian)	Khuren Dukh Formation
Pseudotorellia zhoui^[45]	Sp. nov	In press	Dong et al.	Middle-Late Jurassic	Daohugou Beds
Umaltolepis zhoui	Sp. nov	In press	Dong et al.	Middle-Late Jurassic	Daohugou Beds

Ginkgophyte research

• Revision of Ginkgo abaniensis, based on data from leaves from the Jurassic Mura Formation (Russia), is published by Frolov & Mashchuk (2022), who emend the diagnosis of this species, and transfer Ginkgo abaniensis, Ginkgo glinkiensis and Ginkgo capillata to the genus Ginkgoites.^[46]

Conifers

Cheirolepidiaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Brachyoxylon yanqingense^[48]	Sp. nov		Cheng et al.	Late Jurassic	Tuchengzi Formation		A probable member of the family Cheirolepidiaceae.
Frenelopsis antunesii^[49]	Sp. nov	In press	Mendes & Kvaček	Early Cretaceous (Aptian–Albian)	Figueira da Foz Formation		A member of the family Cheirolepidiaceae.
Pseudofrenelopsis zlatkoi^[50]	Sp. nov		Kvaček & Mendes	Early Cretaceous (Aptian-Albian)	Figueira da Foz Formation

Cupressaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Cupressinanthus klebsii^[51]	Sp. nov	Valid	Sadowski, Schmidt & Kunzmann	Eocene		Europe (Baltic Sea region)	Cupressaceous pollen cone.
Patagotaxodia^[52]	Gen. et sp. nov	Valid	Andruchow-Colombo et al.	Late Cretaceous (Maastrichtian)	Lefipán Formation		A member of the family Cupressaceae. Genus includes new species P. lefipanensis.

Pinaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Keteleerioxylon changchunense^[53]	Sp. nov		Shi, Sun, Meng & Yu in Shi et al.	Early Cretaceous (Albian)	Yingcheng Formation		A Keteleeria-like wood morphogenus.
Nothotsuga mulaensis^[54]	Sp. nov		Li & Dong in Dong et al.	Miocene	Changtai Formation		A species of Nothotsuga.
Pinus prehwangshanensis^[55]	Sp. nov		Bazhenova, Wu & Jin in Bazhenova et al.	Late Pleistocene	Maoming Basin		A pine.
Pinus shengxianica^[56]	Sp. nov		Li, Hu & Xiao in Li et al.	Miocene	Shengxian Formation		A pine.

Podocarpaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Phyllocladoxylon antarcticum^[57]	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation		A podocarpaceous wood morphospecies Announced in 2022 Officially published in 2023
Podocarpoxylon resinosum	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation		A podocarpaceous wood morphospecies Announced in 2022 Officially published in 2023
Podocarpus mexicanoxylon^[58]	Sp. nov		Castañeda-Posadas	Miocene			A species of Podocarpus.

Other conifers

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aciphyllum^[61]	Gen. et sp. nov		Barbacka & Górecki in Barbacka et al.	Early Jurassic (Hettangian)	Zagaje Formation		A needle leaf similar to the leaves of Pinus. Genus includes new species A. triangulatum.
Ductoagathoxylon tsaaganensis^[62]	Sp. nov	In press	Cai, Zhang & Feng in Cai et al.	Late Permian
Sidashia^[63]	Gen. et sp. nov	In press	Forte, Kustatscher & Van Konijnenburg-van Cittert in Forte et al.	Middle Triassic (Anisian)			Genus includes new species S. tridentata.
Ullrichia^[64]	Gen. et comb. nov	Valid	Kerp et al.	Permian			The type species is "Lebachia" laxifolia (1939); genus also includes "L." intermedia (1939) and "L." mucronata (1939).

Conifer research

• Bodnar et al. (2022) reassess the anatomy and systematics of the permineralized conifer-like woods from the Triassic strata from Argentina, confirm the assignment of the logs related to the families Cupressaceae and Cheirolepidiaceae, as well as three taxa related to Araucariaceae (Agathoxylon cozzoi, Agathoxylon protoaraucana and Agathoxylon argentinum), and argue that the fossil woods previously assigned to the families Podocarpaceae and Taxaceae do not have enough preserved characters to support such assignment.^[65]
• A study on the pattern of conifer turnover across the Cretaceous-Paleogene boundary in the Raton and Denver basins (Colorado, United States) is published by Berry (2022).^[66]
• Mantzouka, Akkemik & Güngör (2022) describe fossil woods of Cupressinoxylon matromnense from the middle Miocene Eşelek volcanic deposits (Gökçeada, Turkey), preserved with feeding damage produced by members of the agromyzid genus Protophytobia, and supporting the existence of an eastern Mediterranean Miocene Climatic Optimum hotspot which additionally included Greek islands of Lemnos and Lesbos.^[67]

Flowering plants

Chloranthales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Canrightia foveolata^[68]	Sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation
Proencistemon	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation		Genus includes new species P. portugallicus.

Magnoliids

Laurales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Araliaephyllum silvapinedae^[69]	Sp. nov	In press	Rubalcava-Knoth & Cevallos-Ferriz	Cretaceous (Albian–Cenomanian)	La Cintura Formation
Argapaloxylon salvadorensis^[70]	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene			A member of the family Lauraceae.
Catula^[71] ^[72]	Gen. et sp. nov	Valid	Maccracken et al.	Late Cretaceous (Campanian)	Kaiparowits Formation		A member of the family Lauraceae. Genus includes new species C. gettyi.
Cryptocaryoxylon irregularis	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene	Hançili Formation	Turkey	Fossil wood of a member of the family Lauraceae.
Laurinoxylon scalariforme	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene			A member of the family Lauraceae.
Mezilaurinoxylon americana	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene			A member of the family Lauraceae.
Mezilaurinoxylon draconis	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene			A member of the family Lauraceae.
Mezilaurinoxylon miocenica	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene			A member of the family Lauraceae.

Piperales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aristospermum^[74]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Aptian–Albian)			A member of the family Aristolochiaceae. Genus includes new species A. huberi.
Siratospermum	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Late Cretaceous (Cenomanian)			A member of the family Aristolochiaceae. Genus includes new species S. mauldinense.

Monocots

Commelinid monocots

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sabalites ghughuaensis^[76]	Sp. nov	In press	Kumar, Hazra & Khan in Kumar et al.	Late Cretaceous-Paleocene (Maastrichtian-Danian)	Deccan Intertrappean Beds		A member of the family Arecaceae belonging to the subfamily Coryphoideae.
Sabalites umariaensis	Sp. nov	In press	Kumar, Hazra & Khan in Kumar et al.	Late Cretaceous-Paleocene (Maastrichtian-Danian)	Deccan Intertrappean Beds		A member of the family Arecaceae belonging to the subfamily Coryphoideae.

Monocot research

• Leaf fossils of costapalmate-palms belonging to the genus Sabalites are described from the ?Santonian–Campanian Belly River Group, Campanian Foremost Formation (Alberta, Canada) and Maastrichtian Frenchman Formation (Saskatchewan, Canada) by Greenwood, Conran & West (2022), who interpret the studied fossils as constraining climate reconstructions for the Late Cretaceous high mid-latitudes of North America (c. 55° N) to exclude significant freezing episodes; the authors also transfer the Late Cretaceous species "Geonomites" imperialis to the genus Phoenicites, and reassess Sabalites carolinensis as more likely to be Campanian than Coniacian–Santonian in age.^[77]
• A study on the impact of the absence of megaherbivores in the aftermath of the Cretaceous–Paleogene extinction event on the evolution of palms is published by Onstein, Kissling & Linder (2022).^[78]
• A study on the evolutionary history of palms belonging to the group Mauritiinae, as inferred from a phylogenetic analysis incorporating fossil data, is published by Bacon et al. (2022).^[79]

Basal eudicots

Proteales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Distefananthus^[80]	Gen. et sp. nov		Huegele & Wang	Early Cretaceous (Albian)	Dakota Formation		A platanaceous inflorescence. Genus includes new species D. hoisingtonensis.
Langeranthus	Gen et sp nov	Valid	Huegele & Manchester	Eocene Ypresian	Klondike Mountain Formation		A platanaceous flowering head. The type species is L. dillhoffiorum
Meliosma eosinica^[81]	Sp. nov		Moiseeva, Kodrul & Jin in Moiseeva et al.	Late Eocene	Huangniuling Formation		A species of Meliosma.
Nelumbo delinghaensis^[82]	Sp. nov		Luo & Jia in Luo et al.	Miocene	Upper Youshashan Formation		A species of Nelumbo.
Nelumbo fujianensis^[83]	Sp. nov	In press	Dong et al.	Miocene	Fotan Group		A species of Nelumbo.
Platimeliphyllum durhamensis^[84]	Comb nov	in press	(Wolfe)	Late Eocene	Puget Group "Upper Fultonian" Loc 9832	United States	A platanaceous leaf. Moved from "Fothergilla" durhamensis (1968).
Platimeliphyllum fushunensis	Comb nov	in press	(Chen)	Eocene	Fushun Formation	China	A platanaceous leaf. Moved from "Betula" fushunensis.
Sapindopsis chinensis^[85]	Sp. nov		Golovneva et al.	Early Cretaceous (Albian)	Dalazi Formation		A member of the family Platanaceae.
Sapindopsis orientalis	Sp. nov		Golovneva et al.	Early Cretaceous (Albian)	Frentsevka Formation		A member of the family Platanaceae.

Protealean research

• Redescription of the Okanagan Highlands genus Langeria with description of associated stipules and reproductive structures plus formal reassignment of the genus to Platanaceae by Huegele & Manchester is published.^[86]

Ranunculales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Berberis auriolensis^[87]	Sp. nov	Valid	Denk & Sami in Denk et al.	Pleistocene (Calabrian)			A species of Berberis.
Mahonia mangbangensis^[88]	Sp. nov		Tang et al.	Pliocene	Mangbang Formation		A species of Mahonia.
Palaeosinomenium hengduanensis^[89]	Sp. nov		Wu & Zhou in Wu et al.	Eocene	Shuanghe Formation		A member of the family Menispermaceae.

Superasterids

Aquifoliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ilex antiquorum^[90]	Nom. nov	Valid	Doweld	Late Cretaceous (Maastrichtian)			A holly; a replacement name for Ilex antiqua Knobloch & Mai (1986).
Ilex myricina	Nom. nov	Valid	Doweld	Miocene (Messinian)			A holly; a replacement name for Ilex myricoides Massalongo (1858).

Cornales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Blackwelloxylon^[92]	Nom. nov	Valid	Deshmukh	Pleistocene			A member of the family Cornaceae; a replacement name for Cornoxylon Blackwell (1982).
Exbeckettia^[93]	Gen. et comb. nov	Valid	Manchester & Collinson	Early Eocene	London Clay		A mastixioid fruit; a new genus for Beckettia mastixioides Reid & Chandler (1933).

Metteniusales

Superrosids

Fabales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Albizia yenbaiensis^[99]	Sp. nov	Announced	Nguyen, Su & J. Huang in Nguyen et al.	Miocene	Yen Bai Basin		An Albizia species. Announced in 2022 Officially published January 2023
Anadenantheroxylon kurupaum^[100]	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae.
Cedrelinga paleocatenaeformis	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A species of Cedrelinga.
Cercioxylon mediterraneum^[101]	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene	Hançili Formation	Turkey	Fossil wood of a member of the family Fabaceae.
Chloroleucoxylon	Gen. et sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae. Genus includes new species C. yukeriense.
Enterolobiumoxylon vassalloae	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae.
Leguminocarpum meghalayensis^[102]	Sp. nov	Announced	Bhatia, Srivastava & Mehrotra	Late Paleocene	Tura Formation		A fabaceous seed pod morphospecies. Announced in 2022 Officially published in 2023
Microlobiusxylon parafoetidus	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae.
Paleobowdichia^[103]	Gen. et comb. nov	Valid	Herendeen et al.	Latest Paleocene to late early Eocene	Lamar River Formation	( )	A member of Papilionoideae; a new genus for "Acacia" lamarensis Knowlton (1899).
Parapiptadenioxylon	Gen. et sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A Fabaceae genus. The type species is P. pararigida.
Parvileguminophyllum damalgiriensis	Sp. nov	Announced	Bhatia, Srivastava & Mehrotra	Late Paleocene	Tura Formation		A fabaceous leaf morphospecies. Announced in 2022 Officially published in 2023
Podocarpium tibeticum^[104]	Sp. nov	In press	Li, Huang & Su in Li et al.	Late Eocene	Lunpola Basin		A member of the family Fabaceae.
Pseudopiptadenioxylon	Gen. et sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae. Genus includes new species P. uniseriatum.
Tobya	Gen. et comb. nov	Valid	Herendeen et al.	Eocene	Cockfield Formation	( )	A member of Papilionoideae; a new genus for "Diplotropis" claibornensis Herendeen & Dilcher (1990).

Fabalean research

• New fossil material of members of the genus Bauhinia is described from the Eocene of the Puyang Basin (China) by Jia et al. (2022), who interpret their findings as the earliest reliable fossil records of Bauhinia in Asia.^[105]
• Moya et al. (2022) study the affinities of fossil legumes Entrerrioxylon victoriensis, Gossweilerodendroxylon palmariensis, Paraoxystigma concordiensis and Cylicodiscuxylon paragabunensis from the Cenozoic Paraná, Arroyo Feliciano and El Palmar formations (Argentina) with extant West African legumes, and discuss the possible migration routes by which these plants may have arrived in South America from Africa.^[106]

Fagales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Castanopsis zhoui^[107]	Sp. nov	In press	Wang et al.	Miocene	Fotan Group		A species of Castanopsis.
Comptonia hirsuta^[108]	Sp. nov		Xiao & Ji in Ji et al.	Miocene	Hannuoba Formation		A species of Comptonia.
Myricoxylon doganyurtensis	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene		Turkey	Fossil wood of a member of the family Myricaceae.
Nothofagoxylon ruei	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation		A nothofagaceous wood morphospecies Announced in 2022 Officially published in 2023
Pterocarya magnifructa^[109]	Sp. nov	Valid	Stults, Tiffney & Axsmith	Pliocene			A species of Pterocarya.
Quercus nanningensis^[110]	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation		An oak.
Quercus paleodisciformis	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation		An oak.
Quercus paleohui	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation		An oak.
Quercus yongningensis	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation		An oak.

Malpighiales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Belenocarpa^[111]	Gen. et comb. nov	Valid	Hamersma et al.	Early Oligocene			A member of the family Euphorbiaceae; a new genus for "Jatropha" tertiara Berry.
Elatine odgaardii^[112]	Sp. nov	Valid	Bennike in Bennike et al.	Probably early Pleistocene			A species of Elatine. Announced in 2022; the final article version was published in 2023.
Mammeoxylon beylikduezuense^[113]	Sp. nov	In press	Akkemik et al.	Late Oligocene-Early Miocene	İstanbul Formation		A Mammea relative wood morphospecies
Mammeoxylon paramericana	Comb. nov	In press	(Nelson & Jud) Akkemik & D. Mantzouka	Miocene			A Mammea relative wood morphospecies Moved from Mammea paramericana (2017)
Parinari hilliana^[114]	Sp. nov	Valid	Grote in Grote, Duangkrayom & Jintasakul	Late Miocene	Tha Chang beds		A species of Parinari.
Parinari khoratensis	Sp. nov	Valid	Grote in Grote, Duangkrayom & Jintasakul	Late Miocene	Tha Chang beds		A species of Parinari.
Plukenetia minima^[115]	Sp. nov	In press	Poinar	Miocene	Dominican amber		A species of Plukenetia.

Malvales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Malvacipolloides deccanensis^[116]	Sp. nov		Manchester et al.	Late Cretaceous-Paleocene (Maastrichtian–Danian)	Deccan Intertrappean Beds		A member of the family Malvaceae.
Malvacipolloides intertrappea	Sp. nov		Manchester et al.	Late Cretaceous-Paleocene (Maastrichtian–Danian)	Deccan Intertrappean Beds		A member of the family Malvaceae.
Thespesia neopopulnea^[117]	Sp. nov	Valid	Hazra, Mahato & Khan in Hazra et al.	Pliocene	Rajdanda Formation		A species of Thespesia.

Malvalean research

• A study on the evolutionary history of Dipterocarpaceae, as indicated by biogeography of pollen fossils from Africa and India, molecular data and fossil amber records, is published by Bansal et al. (2022).^[118]

Myrtales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Hemitrapa zhangpuensis^[119]	Sp. nov		Dong et al.	Miocene			A member of the family Lythraceae belonging to the subfamily Trapoideae.
Myrtineoxylon hoffmannae	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation		A myrtaceous wood morphospecies. Announced in 2022 Officially published in 2023
Trapa natanifolia^[120]	Sp. nov		Han & Jia in Han et al.	Late Eocene	Bailuyuan Formation		A water caltrop.
Trapa qaidamensis^[121]	Sp. nov		Cai et al.	Miocene	Shangyoushashan Formation		A water caltrop.

Oxalidales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ceratopetalum suciensis^[122]	Sp nov	In press	Tang, Smith, & Atkinson	Late Cretaceous Campanian	Cedar District Formation		A Cunoniaceous species.
Connaroxylon^[123]	Gen. et sp. nov	Valid	Baas et al.	Cretaceous Maastrichtian-earliest Paleocene	Deccan Intertrappean Beds		A probable Connaraceous wood morphotaxon. The type species is C. dimorphum. First named in 2017 but failed ICBN requirements;^[124] subsequently validated in 2022.
Cunoniocarpa^[125]	Gen. et sp. nov	Valid	Matel et al.	Early Eocene	Huitrera Formation		A member of Cunoniaceae. The type species is C. stylosa.
Racemofructus	Gen. et sp. nov	Valid	Matel et al.	Early Eocene	Huitrera Formation		A member of Cunoniaceae. The type species is R. fasciculatus.
Weinmannioxylon trichospermoides	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation		A cunoniaceous wood morphospecies. Announced in 2022 Officially published in 2023

Oxalidalean research

Tand, Smith, and Atkinson describe the first North American instance of the previously Paleo-Antarctic Rainforest Lineage Cunoniaceae fruits from Sucia Island. Previously considered solely a Gondwanan family, the new species indicate a complex geographic history for the group.^[122]

Rosales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Eophylica^[126]	Gen. et sp. nov	Valid	Shi et al.	Cretaceous	Burmese amber		A rhamnaceous floral morphotaxon. The type species is E. priscastellata.
Ficoxylon fusiforme^[127]	Sp. nov	Valid	El-Noamani	Late Cretaceous	Taref Formation		A member of the family Moraceae.
Ficus fujianensis^[128]	Sp. nov	In press	Dong et al.	Miocene			A species of Ficus.
Ficus zhangpuensis	Sp. nov	In press	Dong et al.	Miocene			A species of Ficus.
Ventilago pliocenica^[129]	Sp. nov		Hazra et al.	Pliocene			A species of Ventilago.
Ventilago siwalika	Sp. nov		Hazra et al.	Miocene			A species of Ventilago.

Sapindales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Barkleya^[130]	Gen. et comb. nov	Valid	Manchester & Judd	Eocene Ypresian	Green River Formation		An anacardiaceous samara. The type species is B. schinoloxa. Moved from Anacardites schinoloxus (1929). Possibly the fruits of "Rhus" nigricans.
Canarium haominiae^[131]	Sp. nov	In press	Yin et al.	Miocene			A species of Canarium.
Canarium maomingense^[132]	Sp. nov		Xiang & Jin in Xiang et al.	Late Pleistocene	Maoming Basin		A species of Canarium.
Choerospondias mioaxillaris^[133]	Sp. nov		Xiao & Wu in Xiao et al.	Miocene	Shengxian Formation		A species of Choerospondias.
Choerospondias tiantaiensis	Sp. nov		Xiao & Wu in Xiao et al.	Miocene	Shengxian Formation		A species of Choerospondias.
Grimmipollis^[134]	Gen. et sp. nov		Huang, Morley & Hoorn in Huang et al.	Eocene	Yaw Formation		A member of the family Sapindaceae. Genus includes new species G. burmanica.
Koelreuteria kvacekii^[135]	Sp. nov	Valid	Chen, Del Rio & Su in Chen et al.	Eocene	Niubao Formation		A species of Koelreuteria.
Loxopteroides^[136]	Gen. et sp. nov	In press	Manchester & Judd	Eocene	Ione Formation	( )	A member of the family Anacardiaceae. Genus includes new species L. weeksae.
Vaudoisia^[137]	Gen. et comb. nov	Valid	Strullu-Derrien et al.	Eocene			A fruit of likely sapindalean affinity; a new genus for "Juglandicarya" gruetii Vaudois-Miéja (1976).

Other Eudicots

Other angiosperms

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Archaebuda^[139]	Gen. et sp. nov		Chen & Wang	Early Cretaceous (Barremian-Aptian)	Yixian Formation		A flower bud of an early angiosperm. Genus includes new species A. lingyuanensis.
Ascarinophyllum^[140]	Gen. et sp. nov	Announced 2022	Čepičková & Kvaček	Late Cretaceous (Cenomanian)	Peruc–Korycany Formation
Covidifructus^[141]	Gen. et sp. nov	Valid	Heřmanová et al.	Late Cretaceous (late Turonian-Santonian)	Klikov Formation		An angiosperm fruit of uncertain affinities, with similarities to the family Dilleniaceae. The type species is C. multicarpellatus.
Elasmostemon	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation		A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. paisii.
Endressistemon	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation		A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. cateficensis.
Fairlingtonia microgyna^[142]	Sp. nov		Du et al.	Early Cretaceous	Zhonggou Formation		A herbaceous eudicot.
Florigerminis^[143]	Gen. et sp. nov	In press	Cui et al.	Middle-Late Jurassic	Jiulongshan Formation		A possible flower bud. The type species is F. jurassica. First announced online 2021, Final article published in 2022.
Gansupeltata^[144]	Gen. et sp. nov	Valid	Wu et al.	Early Cretaceous (Aptian)	Chijinpu Formation		An early flowering plant. Genus includes new species G. beishanensis.
Herbifolia^[145]	Gen. et sp. nov	In press	Frolov & Enushchenko	Middle Jurassic (Aalenian)	Irkutsk Coal Basin		An angiosperm with leaf epidermal structure most similar to those of modern Asparagales and Liliales. Genus includes new species H. antiqua.
Honeytheca	Gen. et sp. nov	In press	Huegele & Manchester	Paleocene	Fort Union Formation		A flowering plant of uncertain affinities. Genus includes new species H. bighornensis.
Ibericarpus	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation		A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species I. cuneiformis.
Lingyuananthus^[146]	Gen. et sp. nov		Wang	Early Cretaceous (Barremian–Aptian)	Yixian Formation		An early angiosperm. Genus includes new species L. inexpectus.
Phylica piloburmensis	Sp. nov	Valid	Shi et al.	Cretaceous	Burmese amber		A flowering plant of uncertain affinities. Originally described as a species of Phylica. Oskolski et al. (2024) interpreted it as a flowering plant with an affinity to Rhamnaceae, possibly to an extinct basal lineage;^[147] on the other hand Beurel et al. (2024) interpreted it as more likely to have lauralean affinities, and made it the type species of the separate genus Nothophylica.^[148]
Santaniella^[149]	Gen. et 2 sp. nov		Gobo et al.	Early Cretaceous (Barremian-Aptian)	Crato Formation		Originally described as a member or a relative of the family Ranunculaceae, but subsequently considered to be a mesangiosperm of uncertain affinities, possibly a magnoliid.^[150] Genus includes new species S. lobata and S. acuta.
Todziaphyllum saportanum	Comb. nov	Announced	(Velenovský) Čepičková & Kvaček	Late Cretaceous (Cenomanian)	Peruc–Korycany Formation
Tolmania^[151]	Gen. et sp. nov	Valid	Edmonds, Stockey & Rothwell	Late Cretaceous (Maastrichtian)	St. Mary River Formation		An aquatic dicot. Genus includes new species T. aquatica.
Valvidistemon	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation		A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species V. globiferus.

General angiosperm research

• Surangea mohgaoensis, originally interpreted as fern megaspores, is reinterpreted as angiosperm fruits by Ramteke et al. (2022).^[152]
• Zhang et al. (2022) describe rich assemblages of spiny plant fossils from the Eocene (Bartonian) Niubao Formation (Tibet, China), preserving seven different spine morphologies, and interpret this finding as evidence of the presence of a diversity of spiny plants in Eocene central Tibet, as well as evidence of a rapid diversification of spiny plants in Eurasia around that time.^[153]
• A preliminary report on a new fossil angiosperm flora of the Lesvos Petrified Forest at Akrocheiras east of Sigri on Lesbos, Greece is given by Kafetzidou et al. Preliminary taxa identifications are given and commentary on the climactic implications are made.^[154]
• A study aiming to determine the relationship between past atmospheric CO₂ and temperature fluctuations and the shifts in diversification rates of Poaceae and Asteraceae is published by Palazzesi et al. (2022).^[155]

Other plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aegianthus irkutensis^[156]	Sp. nov	In press	Nosova & Tekleva	Middle Jurassic	Prisayan Formation		Pollen cone with pollen of ginkgoalean or gnetophytalean affinity.
Aysenoxylon	Gen et sp nov	Announced	Pujana et al.	Oligocene	San José Formation		A wood morphospecies of uncertain affinity. The type species is A. patorarensis. Announced in 2022 Officially published in 2023
Bryokhutuliinia ignatovii^[157]	Sp. nov	Valid	Frolov, Kazanovsky & Enushchenko	Early Jurassic (Toarcian)	Middle Subformation of Prisayan Formation		A member of Bryopsida of uncertain affinities.
Combina^[158]	Gen. et sp. nov		Santos & Wang	Middle Triassic (Anisian)	Calcena Formation		A cone-like reproductive organ of a seed plant. Genus includes new species C. triassica.
Dioonitocarpidium rossicum^[159]	Sp. nov	Valid	Gomankov	Permian			A member of Cycadales.
Europoxylon garapensis^[160]	Sp. nov	In press	Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation		A gymnosperm.
Jarudia^[161]	Gen. et sp. nov		Shi et al.	Early Cretaceous	Huolinhe Formation		A seed-bearing structure of a corystosperm seed fern. Genus includes new species J. zhoui.
Komlopteris distinctiva	Sp. nov		Barbacka in Barbacka et al.	Early Jurassic (Hettangian)	Zagaje Formation		Cuticle of a seed fern.
Lesleya ceriacoi^[162]	Sp. nov	In press	Correia et al.	Carboniferous (Gzhelian)	Douro Carboniferous Basin		An early gymnosperm.
Palaeodichelyma kiritchkovae	Sp. nov	Valid	Frolov, Kazanovsky & Enushchenko	Early Jurassic (Pliensbachian)	Lower Subformation of Prisayan Formation		A member of Bryopsida of uncertain affinities.
Paragigantopteris^[163]	Gen. et sp. nov	In press	Ma et al.	Permian (Wuchiapingian)	Lungtan Formation		A gigantopterid. Genus includes new species P. qingloongensis.
Pauthecophyton hezhangensis^[164]	Sp. nov		Wang et al.	Devonian (Pragian-Emsian)	Danlin Formation		A euphyllophyte of uncertain affinities.
Piterophyton^[165]	Gen. et sp. nov	Valid	Naugolnykh	Ordovician			A rhyniophyte of uncertain affinities. The type species is P. caudatum.
Polycanaloxylon	Gen. et sp. nov	In press	Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation		A gymnosperm. Genus includes new species P. merlottii.
Psilophyton diakanthon^[166]	Sp. nov		Colston, Landaw & Tomescu	Devonian (Emsian)	Battery Point Formation		A member of the group Trimerophytopsida.
Renbernia^[167]	Gen. et sp. nov	In press	Friis, Crane & Pedersen	Early Cretaceous (Albian)	Potomac Group		A seed plant similar to Brenneria potomacensis. Genus includes new species R. zhoui.
Rhyniotaenium^[168]	Gen. et sp. nov	In press	Krings	Early Devonian	Rhynie chert		An alga, probably a green alga belonging to the family Mesotaeniaceae. Genus includes new species R. velatum.
Sinoglossa^[169]	Gen. et sp. nov	Valid	Zhang et al.	Middle Triassic	Linjia Formation		A member of Glossopteridales. The type species is S. sunii.
Taimyria^[170]	Gen. et sp. nov	Valid	Naugolnykh & Mogutcheva	Early Triassic (Induan)	Keshinskian/Keshinskaya Formation		A member of Peltaspermales belonging to the family Angaropeltaceae. Genus includes new species T. triassica.
Taungurungia^[171]	Gen. et sp. nov	Valid	McSweeney, Shimeta & Buckeridge	Devonian (Pragian–Emsian)	Norton Gully Sandstone Formation		A plant of uncertain affinities, similar to members of Zosterophyllopsida. Genus includes new species T. garrattii.
Teyoua^[172]	Gen. et sp. nov	In press	Huang, Liu & Xue	Devonian (probably Pragian)	Mangshan Group		A polysporangiate land plant. Genus includes new species T. antrorsa.
Traskia^[173]	Gen. et sp. nov	Valid	Rothwell et al.	Jurassic			A stem-cycad. Genus includes new species T. maahlae.
Vetiplanaxis obtusus^[174]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber		A moss belonging to the group Hypnodendrales.
Wilhowia^[175]	Gen. et sp. nov	Valid	Gensel	Devonian (Emsian)	Battery Point Formation		A basal euphyllophyte. Genus includes new species W. phocarum.
Xadzigacalix^[176]	Gen. et sp. nov	Valid	Klymiuk, Rothwell & Stockey	Early Cretaceous (Valanginian)			A gymnosperm of uncertain phylogenetic placement, possibly having affinities with gnetophytes or angiosperms. Genus includes new species X. quatsinoensis.
Xinhangia^[177]	Gen. et sp. nov		Yang & Wang	Devonian (Famennian)	Wutong Formation		A fern-like plant of uncertain affinities. Genus includes new species X. spina.

Other plant research

• A study on the xylem development in Leptocentroxyla, and on its implications for the knowledge of the evolution of pith, is published by Tomescu & McQueen (2022).^[178]
• Decombeix et al. (2022) report evidence of tylosis formation in permineralized wood of Dameria hueberi from the Tournaisian of Australia.^[179]
• The first comprehensive crown reconstruction of Medullosa stellata var. typica, based on data from a specimen from the Chemnitz petrified forest (Germany), is presented by Luthardt et al. (2022).^[180]
• Fossil material of Rhabdotaenia is reported from the Permian Umm Irna Formation (Jordan) by Blomenkemper et al. (2022), representing the northernmost occurrence of this Gondwanan leaf type reported to date.^[181]

Palynology

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Anapiculatisporites radiatus^[182]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore.
Apricasporites	Gen. et sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore. Genus includes new species A. cancellosus.
Camptotriletes inaequabilis	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore.
Camptotriletes suggrandis	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore.
Convolutispora inreligata	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore.
Endosporites circumsaeptus	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore.
Foveosporites magnus	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore.
Granulatisporites commutabilis	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation		A trilete spore.
Paxillitriletes permicus^[183]	Sp. nov	In press	Sui, McLoughlin & Feng in Sui et al.	Permian (Wuchiapingian–Changhsingian)	Xuanwei Formation		A lycophyte megaspore.
Sergipea multipapillata^[184]	Sp. nov		Hu et al.	Early Cretaceous	Bongor Basin		A gymnosperm pollen.
Volkheimerites^[185]	Gen. et sp. nov	In press	Narváez et al.	Paleocene (Danian)	Salamanca Formation		Pollen of a flowering plant. The type species is V. labyrinthus.
Yezopollis^[186]	Gen. et sp. nov	In press	Legrand, Yamada & Nishida	Late Cretaceous (Cenomanian–Turonian)	Mikasa Formation		A Normapolles-type flowering plant pollen. Genus includes new species Y. mikasaensis.

Research

• Review of the studies on the origin of the land flora is published by Bowman (2022).^[187]
• A study on the evolution of body plans of members of Viridiplantae, based on a review of the fossil record, molecular data and developmental biology, is published by Niklas & Tiffney (2022).^[188]
• A study on the biodiversity of land plants at the equator during their first major diversification in the Late Silurian–Early Devonian is published by Wellman et al. (2022).^[189]
• A study on the evolution of heterospory during the Devonian is published by Leslie & Bonacorsi (2022).^[190]
• Seven coniferous nurse logs that have been colonized by conifer and equisetalean roots are reported from four Permian intervals in the Ordos Basin (China) by Feng et al. (2022), indicating that conifer tree stems probably functioned as hosts to both conspecific and interspecific seedlings in the Cathaysian Flora.^[191]
• A study on the impact of the Intertropical Convergence Zone in the emerging South Atlantic region on Aptian plant communities from eight Brazilian sedimentary basins is published by Carvalho et al. (2022), who report evidence of an overall predominance of xerophytic plants, attesting to more dry conditions, and of a humidification trend towards the end of the late Aptian resulting in the predominance of hydrophytes, hygrophytes, tropical lowland flora and upland flora, indicative of prevalence of lowland and montane rainforests.^[192]
• A study on the distribution and relative abundances of major plant groups from the Albian Gates Formation (Alberta, Canada) is published by Kalyniuk et al. (2022).^[193]
• A study on the relationship between whole-genome duplication, seed traits and the selectivity of the survival of plants during the Cretaceous–Paleogene extinction event is published by Berry & Jaganathan (2022).^[194]
• New Oligocene flora is described from the Dong Ho Formation (Vietnam) by Huang et al. (2022), who interpret the studied fossils as evidence of long-term environmental, floristic and vegetational stability in this region since the Paleogene.^[195]
• Gentis et al. (2022) describe fossil wood specimens from the Miocene Natma Formation (Myanmar), representing an assemblage dominated by members of the families Fabaceae and Dipterocarpaceae, interpreted as coming from different types of low altitude forest ecosystems (tropical wet evergreen, tropical dry and deciduous, and tropical littoral), and interpreted as indicative of a monsoonal climate with an alternance of a dry season and a wet season.^[196]
• Abundant compression floras dominated by angiosperm leaves are described from two sites of probable Pliocene age in Brunei by Wilf et al. (2022), who interpret these floras as evidence of dipterocarp-dominated lowland rainforests in the Malay Archipelago before the Pleistocene.^[197]
• A study on the impact of the extinct Neotropical megafauna on the variability in plant functional traits and biome geography in Central and South America is published by Dantas & Pausas (2022).^[198]
• A study on plant material from rock overhangs from mid-late Holocene sites along the Kawarau-Cromwell-Roxburgh Gorges in Central Otago (New Zealand), much of which was likely transported as roosting material or consumed by moa birds, and on its implications for the knowledge of the mid-late Holocene regional vegetation of Central Otago and the knowledge of vegetation changes since mid-late Holocene, is published by Pole (2022).^[199]
• A study on the role of hydraulic failure in the evolution of early vascular plants is published by Bouda et al. (2022), suggesting that drought selection played a key role in the diversification of vascular arrangements beginning with the Devonian explosion.^[200]

Notes and References

1. Feist . M. . Floquet . M. . Charophytes from the Upper Cretaceous Castilian marine ramp and continental basins (central northern Spain): fossil assemblages and depositional environments . 2022 . Cretaceous Research . 140 . Article 105325 . 10.1016/j.cretres.2022.105325 . 2022CrRes.14005325F . 251681127 . free .
2. Cao . W. . Li . S. . Li . Q. . Stidham . T. A. . Wan . X. . Ni . X. . 2022 . Asian Paleocene charophyte records demonstrate Eocene dispersals from Asia to Europe . Journal of Paleontology . 96 . 3 . 706–714 . 10.1017/jpa.2021.118 . 2022JPal...96..706C . 246456520 .
3. Grgasović . T. . Taxonomy of the fossil calcareous algae: Revision of genera Physoporella Steinmann and Oligoporella Pia (Dasycladales) . 2022 . Carnets Geol. . 22 . 7 . 171–310 . 10.2110/carnets.2022.2207 . 251189240 .
4. LoDuca . S. T. . Meacher . M. . Pepper . P. . Brett . K. . Isotalo . P. A. . Earltonella fredricksi n. gen n. sp. and Thalassocystis striata (Chlorophyta, Bryopsidales) from the Silurian (Llandoverian) of the Timiskaming outlier, Ontario, Canada . 2022 . Journal of Paleontology . 97 . 2 . 516–532 . 10.1017/jpa.2022.86 . 252936182 .
5. Torromé . D. . Schlagintweit . F. . Milanovicella? canadillana sp. nov., an Upper Cretaceous supposedly calcitic Dasycladale (green algae) from the middle Santonian–lower Campanian of northeastern Spain . 2022 . Cretaceous Research . 141 . Article 105365 . 10.1016/j.cretres.2022.105365 . 252301204 .
6. Chai . S. . Aria . C. . Hua . H. . 2022 . A stem group Codium alga from the latest Ediacaran of South China provides taxonomic insight into the early diversification of the plant kingdom . BMC Biology . 20 . 1 . 199 . 10.1186/s12915-022-01394-0 . 36127662 . 9491005 . free .
7. Vachard . D. . Krainer . K. . Calcareous algae and foraminifers across the Permian-Triassic boundary interval (uppermost Bellerophon Formation and basal Werfen Formation) in the Dolomites (South Tyrol – Trentino, Italy) . 2022 . Palaeontographica Abteilung A . 324 . 1–6 . 1–173 . 10.1127/pala/2022/0128 . 2022PalAA.324....1V . 250292126 .
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